1 Culm nodes , at least the lower,
bearded (often
retrorsely ).
2 Nodes with a
viscid band below the
beard ;
internodes and
sheaths velvety-
pubescent ;
vernal culm blades 9-20 cm long, 9-20 mm wide
3 Second
glume and
sterile lemma acute to short-
acuminate , some in a given specimen 0.3 mm or more longer than fertile
lemma .
4 Ligule an eciliate membrane;
peduncle scabrous ; first
glume 0.3-0.6 (-0.8) mm long,
reniform to suborbicular
4 Ligule a
ciliate membrane or entirely
ciliate ;
peduncle smooth; first
glume 0.7-1.6 mm long, broadly
ovate to
lanceolate .
5 Ligule a
ciliate membrane; larger leaves 10-20 cm long;
spikelets pubescent , (2.6-) 2.8-3.4 (-3.6) mm long; first
glume (0.9-) 1.0-1.6 mm long
5 Ligule entirely
ciliate ; larger leaves 9-14 cm long;
spikelets glabrous , 2.1-2.6 mm long; first
glume 0.7-1.2 mm long
3 Second
glume and
sterile lemma blunt to subacute, shorter than, equaling, or barely exceeding fertile
lemma , never by as much as 0.3 mm.
6 Ligule ciliate , 0.2-1.1 (-2.1) mm long; leaf
blades 5-14 cm by 5-17 mm; second
glume 0-0.3 mm shorter than fertile
lemma .
7 Culms 30-75 cm long; lower
sheaths glabrous to sparsely
papillose -
pilose ;
panicle pulvini glabrous to sparsely
pilose ;
spikelets 1.8-2.6 mm long; first
glume narrowly
triangular (
triangular -
ovate ); second
glume 0-0.1 mm shorter than fertile
lemma
7 Culms 60-150 cm long; lower
sheaths papillose -
pilose ;
panicle pulvini pubescent ;
spikelets 2.2-2.8 (-3.0) mm long; first
glume broadly
ovate ; second
glume 0.05-0.3 mm shorter than fertile
lemma
6 Ligule a
ciliate or eciliate membrane (0-) 0.3-0.9 mm long; leaf
blades 5-28 cm by 5-40 mm; second
glume 0-0.1 mm shorter than fertile
lemma .
8 Lower
sheaths papillose -
hispid with spreading
hairs ;
ligule a membrane 0.4-0.9 mm long; leaf
blades 10-28 cm long
8 Lower
sheaths glabrous ,
puberulent , or softly
villous basally;
ligule either a membrane 0-0.3 mm long or a
ciliate membrane 0.4-0.7 mm long; leaf
blades 5-18 cm long.
9 Ligule a
ciliate membrane 0.4-0.7 mm long; leaves 15-40 mm wide;
spikelets 1.6-2.0 mm wide
9 Ligule a membrane 0-0.3 mm long; leaves 5-25 mm wide;
spikelets 1.1-1.3 mm wide
10 Cauline blades 5-18 cm long, 6-14 mm wide, averaging about 10 × as long as wide, the
margins nearly parallel;
spikelets 2.9-3.7 mm long; first
glumes 1.1-2.6 mm long, 2/5 or more as long as the
spikelets
10 Cauline blades 7-12 cm long, (6-) 12-25 mm wide,
ovate -
lanceolate , averaging about 6 × as long as wide, the
margins curved most of their length;
spikelets 2.0-3.2 mm long; first
glumes 0.5-1.0 mm long, about 1/4 as long as
spikelets .
11 Culms erect;
blades more nearly symmetrical;
spikelets 2.0-3.2 mm long;
sterile lemma apex usually blunt to rounded
11 Culms basally
decumbent ;
blades strongly asymmetrical-
falcate ;
spikelets 3.0-3.2 mm long;
sterile lemma apex usually pointed
1 Ligule with a dense ring of short
hairs 0.5-1 mm long, and behind it a ring of longer
hairs 1-5 (-8) mm long called the pseudoligule (see introduction for discussion of the so-called double
ligule ).
2 Culm bases hard, thickened,
corm -like; plants producing
basal branches instead of
rosettes , the branches ultimately producing the largest leaves, 4-13 cm long and 4-11 mm wide;
vernal spikelets 2.3-2.7 mm long, 1.2-1.5 mm wide,
autumnal spikelets 2.6-2.9 mm long, 1.0-1.1 mm wide;
spikelets basally
attenuate , the
glumes attached 0.2-0.4 mm below expanding
base of upper
glume and lower
lemma ; [currently known only from sand ridges in peninsular FL]
2 Plants without this combination of characters.
3 Culms > 100 cm long;
node pubescence densely
retrorse ; larger
blades 12-20 cm long;
spikelets acute ;
vernal spikelets 1.9-2.3 mm long, slightly < 2× as long as wide;
autumnal spikelets 2.2-2.5 mm long, > 2× as long as wide
3 Culms < 100 cm long (rarely longer);
node pubescence various; larger
blades 3-12 cm long;
spikelets rounded to blunt, < 2× as long as wide.
4 Plants velvety-
pubescent throughout;
nodes retrorsely bearded ;
internode pubescence retrorse to spreading;
blades usually
ciliate above
base as well as basally; mature
vernal panicles 3-7 cm long, 2-5 cm wide, about 2/3 as wide as long
4 Plants densely
pubescent to
glabrate , but lacking velvety
pubescence ;
nodes with spreading to
ascending pubescence (sometimes
retrorse in
D. villosissimum );
internode pubescence ascending ,
appressed , or spreading;
blades ciliate only at
base (sometimes appearing
ciliate laterally in
D. ovale var.
ovale ); mature
vernal panicles 4-10 cm long, nearly or equally as wide.
5 Lower
internodes with
retrorse or spreading
hairs to 3+ mm long;
blade adaxial surface
pilose with
hairs to 4+ mm long, these
appressed and angled away from the leaf
axis ;
spikelets 2.1-2.5 mm long
6 Spikelets 2.1-2.6 mm long; lower
culm blades usually sparsely
appressed -
pubescent on both surfaces, eciliate or
ciliate at the
base only
6 Spikelets 2.5-3.1 mm long; lower
culm blades usually
glabrous adaxially except for long
hairs at or near the
margin (appearing
ciliate ),
appressed -
pubescent abaxially
1 Ligule a single structure, without a pseudoligule.
7 Culm bases hard, thickened,
corm -like; plants producing
basal branches instead of
rosettes , the branches ultimately producing the largest leaves, 4-13 cm long and 4-11 mm wide;
vernal spikelets 2.3-2.7 mm long, 1.2-1.5 mm wide,
autumnal spikelets 2.6-2.9 mm long, 1.0-1.1 mm wide;
spikelets basally
attenuate , the
glumes attached 0.2-0.4 mm below expanding
base of upper
glume and lower
lemma ; [currently known only from dry sand ridges in peninsular FL]
7 Plants without this combination of characters.
8 Ligule 2-5 mm long,
ciliate
9 Ligule a stramineous to light brown membrane, with or without terminal
cilia ;
peduncle antrorsely scabrous but not hairy.
10 Panicle rachis smooth,
pellucid -
punctate ; first
glume 0.3-0.6 (-0.8 ) mm long, as broad as or broader than long,
truncate to
obtuse ; larger leaves 10-25 cm long, 8-15 mm wide;
ligule 0.5-1.3 mm long; lowest elongate
culm internode > 2 mm in diameter; lowest
nodes usually
glabrous or
pubescent
10 Panicle rachis scabrous or smooth, not
pellucid -
punctate ; first
glume 0.5-1.1 mm long, longer than wide, rounded to
acute ; larger leaves 3.5-12 cm long, 3-9 mm wide;
ligule 0.1-0.6 mm long; lowest elongate
culm internode < 2 mm in diameter; lowest
nodes retrorsely bearded or
glabrous .
11 Lowest
nodes usually
retrorsely bearded ;
ligules (0.1-) 0.3-0.6 mm long; largest
vernal blades 7-12 cm long, (4.5-) 6-9 mm wide;
panicle peduncle scabrous ;
spikelets ovate -
lanceolate ,
acute , 2.0-2.4 mm long; first
glume lanceolate , blunt to
acute ; fertile
lemma smooth
11 Lowest
nodes usually
glabrous ;
ligules 0.1-0.2 (-0.3) mm long; largest
vernal blades 3.5-7 cm long, 3-6 mm wide;
panicle peduncle smooth;
spikelets elliptic , blunt to
acute , 1.6-2.2 mm long; first
glume ovate to
rotund , rounded to
acute ; fertile
lemma papillose
9 Ligule entirely of white
hairs ;
peduncle variously hairy or
glabrous , but not
antrorsely scabrous .
12 Culms without a
viscid band below the
retrorsely , spreading, or
antrorsely bearded nodes ;
internodes and
sheaths not velvety-
pubescent ,
blades velvety-
pubescent in some.
13 Basal as well as
cauline blades linear , 5-15 cm long,; 1-4 mm wide, 30× or more as long as wide;
spikelets 2.8-3.8 (-4.5) mm long
13 Blades < 15× as long as wide (if > 15×, then
spikelets 1.9-2.5 mm long and
sheaths retrorsely pilose in
D. laxiflorum ).
14 Blades (15-) 20× or more as long as wide, increasing in length upwards on
culm ;
blade adaxial surface (weakly-) strongly
scabrous ;
sheaths glabrous ; second
glume and
sterile lemma exceeding fertile
lemma
14 Blades < 15× as long as wide, similar or more often decreasing in size upwards on
culm (if increasing, then
sheaths strongly
retrorsely pilose in
D. laxiflorum );
blade adaxial surface smooth to moderately
scabrous ; second
glume and
sterile lemma shorter than to exceeding fertile
lemma .
15 Nodes retrorsely bearded ; larger
vernal leaves 7-15 mm wide.
16 Sheaths retrorsely pilose with
hairs 2-3 mm long;
basal leaves usually numerous,
ascending , similar in size and shape to the
culm leaves;
culms branching only at the
base in
autumnal phase
16 Sheaths glabrous or
pilose (if
pilose , then
hairs not both
retrorse and 2-3 mm long);
basal leaves
rosette -forming, usually much shorter than
culm leaves;
culms branching above
base in
autumnal phase.
17 Plants often > 1 m in length; lowest
internodes moderately to densely
villous ;
vernal cauline leaves 6-15;
spikelets 1.8-2.2 mm long,
obovoid to broadly
ellipsoid , rounded to blunt,
pubescent
15 Nodes spreading to
antrorsely bearded ; larger
vernal leaves 3-10 mm wide.
18 Culm internodes , at least the lower,
strigose ,
pilose , or
villous ;
culm nodes bearded with
ascending or spreading
hairs ;
blade surfaces
glabrous or variously hairy.
19 Lower
nodes bearded with erect-
ascending , soft, and long
hairs ; mid-
culm blades usually 20× or more as long as wide.
20 Spikelets (3.0-) 3.2-3.8 mm long,
fusiform , pointed at summit,
attenuate at
base , with both
glumes attached 0.3-0.5 mm below
sterile lemma , the
autumnal spikelets 3.5-3.8 mm long; larger
vernal blades 3-6 mm wide, the lower and mid-
culm blades of similar width;
autumnal blades involute
20 Spikelets 1.8-3.1 mm long,
obovate , blunt, and the
base not
attenuate except in
D. arenicoloides with
autumnal spikelets 2.3-3.1 mm long; larger
vernal blades 2-8 mm wide, the lower usually wider and often shorter than mid-
culm blades ;
autumnal blades involute or flat.
21 Longest
vernal blades to 16 cm, widest
vernal blades 4-8 mm, usually longitudinally
wrinkled ;
vernal and
autumnal spikelets 2.3-3.1 mm long;
autumnal blades flat, the larger to 9 cm × 2-4 mm
21 Longest
vernal blades to 6 (
D. aciculare ) or 12 (
D. arenicoloides ) cm long, 2-5 mm wide, not noticeably
wrinkled ;
vernal spikelets 1.5-2.8 mm long,
autumnal spikelets either 1.5-2.3 (
D. aciculare ) or 2.3-3.1 (
D. arenicoloides );
autumnal blades involute , the larger to 6 cm × 1.5 mm.
22 Longest
vernal blades to 6 (-8) cm, widest
vernal blades 2-5 mm;
vernal and
autumnal spikelets 1.7-2.3 mm long, blunt, not
attenuate , the
glumes attached <0.2 mm below
sterile lemma ; first
glumes 0.6-0.9 mm long; larger
autumnal blades to 3 cm by 1 mm
22 Longest
vernal blades to 12 cm, widest
vernal blades 3-4 (-5) mm;
vernal spikelets 2.1-2.8 mm long;
autumnal spikelets 2.3-3.1 mm long, pointed,
attenuate , the
glumes attached 0.3-0.5 mm below
sterile lemma ; first
glumes 0.7-1.5 mm long; larger
autumnal blades to 6 cm by 1.5 mm
19 Lower and often mid-
culm nodes bearded with spreading, stiffish, and short-to-long
hairs ; mid-
culm blades usually 15× or less as long as wide.
23 Blades stiff, often longitudinally
ribbed , at least the lower
villous or strongly
pilose on the
abaxial surface, and
pilose to
hirsute on the
adaxial surface, at least proximally.
24 Spikelets 2.3-3.0 mm long;
ligule < 1 mm long; mid-
culm blades 4-8 mm wide
24 Spikelets 1.7-2.0 (-2.3) mm long;
ligule 1-2.5 mm long; mid-
culm leaves 2-5 mm wide
23 Blades not noticeably stiff nor longitudinally
ribbed ,
pubescent or
strigose underneath,
glabrous above or with a few long
hairs near the
base .
25 Spikelet bases strongly
attenuate ,
apices blunt to beaked; first
glume (2/5-) ½-2/3 (-4/5) as long as
spikelet , attached 0.3-0.5 (-0.7) mm belong expansion of second
glume and
sterile lemma ; larger
vernal blades 3-6 (-7) mm wide
25 Spikelet bases not noticeably
attenuate ,
apices rounded to
obtuse ; first
glume < ½ as long as
spikelet , attached < 0.3 mm below expansion of second
glume and
sterile lemma ; larger
vernal blades 5-12 mm wide.
26 Culms 50-140 cm long; lowest
internodes usually purplish; leaf
sheaths often
viscid -spotted; leaf
blades glabrous abaxially,
glabrous and
scabrous adaxially;
spikelets 1.8-2.2 mm long
26 Culms 15-60 cm long; lowest
internodes usually olive- or bluish-green; leaf
sheaths not
viscid -spotted; leaf
blades appressed -
pubescent abaxially,
glabrous and smooth to
appressed pubescent adaxially;
spikelets 2.1-3.1 mm long.
27 Spikelets 2.1-2.6 mm long; lower
culm blades usually sparsely
appressed -
pubescent on both surfaces, eciliate or
ciliate at the
base only
27 Spikelets 2.5-3.1 mm long; lower
culm blades usually
glabrous adaxially except for long
hairs at or near the
margin (appearing
ciliate ),
appressed -
pubescent abaxially
2 Leaf
blades , at least the lower,
cordate or subcordate at the
base , 4-25 mm wide.
3 Culm blades 4-12 mm wide;
spikelets obpyriform when viewed dorsally, strongly plano-
convex when viewed laterally, usually markedly reddish-purple basally; fertile
lemma papillose
3 Culm blades 5-25 mm wide;
spikelets elliptic to
elliptic -
obovoid when viewed dorsally or laterally, greenish to faintly purple-tinged basally; fertile
lemma not
papillose .
4 Ligule ciliate , 0.2-0.8 mm long;
culm blades 5-14 mm wide;
spikelets 1.8-2.6 mm long; first
glumes 0.5-1 mm long
4 Ligule an eciliate membrane 0-0.3 mm long;
culm blades 5-25 cm wide;
spikelets 2.0-3.7 mm long; first
glumes 0.5-2.6 mm long.
5 Internodes crisp-
puberulent ; larger
culm blades 4-8 (-11) cm long, 5-10 (-12) mm wide, broadest near the
base ;
spikelets 2.1-2.7 mm long; first
glumes 0.7-0.9 mm long
5 Internodes glabrous to sparsely
pubescent , larger
culm blades 5-18 cm long, 5-25 mm wide, broadest near the middle or broadly
linear ;
spikelets 2.0-3.7 mm long; first
glumes 0.5-2.6 mm long.
6 Cauline blades 5-18 cm long, 6-14 mm wide, averaging about 10 × as long as wide, the
margins nearly parallel;
spikelets 2.9-3.7 mm long; first
glumes 11-26 mm long, 2/5 or more as long as the
spikelets
6 Cauline blades 7-12 cm long, (6-) 12-25 mm wide,
ovate -
lanceolate , averaging about 6 × as long as wide, the
margins curved most of their length;
spikelets 2.0-3.2 mm long; first
glumes 5-10 mm long, about 1/4 as long as
spikelets .
7 Culms erect;
blades more nearly symmetrical;
spikelets 2.0-3.2 mm long;
sterile lemma apex usually blunt to rounded
7 Culms basally
decumbent ;
blades strongly asymmetrical-
falcate ;
spikelets 3.0-3.2 mm long;
sterile lemma apex usually pointed
2 Leaf
blades tapering to the
base , 2-12 mm wide.
8 Ligule a stramineous to light brown membrane, with or without terminal
cilia .
9 Ligule a
ciliate membrane;
spikelets (2.6) 2.8-3.4 (3.6) mm long; first
glume (0.9) 1.1-1.6 mm long
9 Ligule an eciliate membrane;
spikelets 1.6-2.8 (3.0) mm long; first
glume 0.3-1.1 mm long.
10 Panicle rachis smooth,
pellucid -
punctate ; first
glume 0.3-0.6 (-0.8 ) mm long, as broad as or broader than long,
truncate to
obtuse ; larger leaves 10-25 cm long, 8-15 mm wide;
ligule 0.5-1.3 mm long; lowest elongate
culm internode > 2 mm in diameter; lowest
nodes usually
glabrous or
pubescent
10 Panicle rachis scabrous or smooth, not
pellucid -
punctate ; first
glume 0.5-1.1 mm long, longer than wide, rounded to
acute ; larger leaves 3.5-12 cm long, 3-9 mm wide;
ligule 0.1-0.6 mm long; lowest elongate
culm internode < 2 mm in diameter; lowest
nodes retrorsely bearded or
glabrous .
11 Lowest
nodes usually
retrorsely bearded ;
ligules (0.1-) 0.3-0.6 mm long; largest
vernal blades 7-12 cm long, (4.5-) 6-9 mm wide;
panicle peduncle scabrous ;
spikelets ovate -
lanceolate ,
acute , 2.0-2.4 mm long; first
glume lanceolate , blunt to
acute ; fertile
lemma smooth
11 Lowest
nodes usually
glabrous ;
ligules 0.1-0.2 (-0.3) mm long; largest
vernal blades 3.5-7 cm long, 3-6 mm wide;
panicle peduncle smooth;
spikelets elliptic , blunt to
acute , 1.6-2.2 mm long; first
glume ovate to
rotund , rounded to
acute ; fertile
lemma papillose
8 Ligule of short white
hairs or absent.
12 Leaves basally disposed, usually matted or cushion-forming, larger than the mid and upper
culm leaves;
blade margins uniformly
papillose -
ciliate ;
culms branching only at the
base , 0.5-3.5 dm tall;
internodes glabrous or sparsely
pubescent
12 Basal leaves usually
rosette -forming, not matted or cushion-forming, usually much smaller than
culm leaves;
blade margins glabrous , or
ciliate only below the middle (or sometimes
papillose -
ciliate throughout in
D. portoricense ssp.
patulum , which has densely
puberulent internodes );
culms branching at the
nodes in age, 1.5-7.5 dm tall.
13 Blades progressively longer upwards on
culm , the larger 9-20 cm long and 3-8 mm wide, (15-) 20 × or more as long as wide, firm but not stiff, not longitudinally
grooved or
wrinkled ;
blade adaxial surface (weakly-) strongly
scabrous ;
sheaths glabrous ; second
glume and
sterile lemma exceeding fertile
lemma
13 Blades similar or more often decreasing in size upwards on
culm , the larger 3-16 cm long and 2-25 mm wide; if 15 × or more as long as wide, then
blades stiff and longitudinally
grooved or
wrinkled (sect.
Angustifolia );
blade adaxial surface smooth to moderately
scabrous ;
sheaths glabrous or
pilose ; second
glume and
sterile lemma shorter than to exceeding fertile
lemma .
14 Blades of mid-
culm leaves typically long and stiff,
acuminate ,
linear or narrowly
lanceolate , usually > 10× as long as wide, only 2-6 mm wide when < 8 cm long.
15 Vegetative parts
glabrous (
spikelets pubescent , lowest
internodes and
sheaths sometimes sparsely
pubescent ,
blades and
sheaths sometimes
ciliate ); mature
panicles less than ¼ as wide as long, the branches erect-
ascending , the
spikelets often subsecund;
autumnal blades 4-10 cm long,
involute , < 2 mm wide;
spikelets 1.8-2.2 mm long.
16 Larger
vernal blades 5-8 cm long and 3-5 mm wide;
vernal panicles usually less than ¼ as wide as long,
spikelets often subsecund along the strongly
ascending to erect branches;
spikelets 1.8-2.2 mm long, second
glume and
sterile lemma pubescent , summits blunt, about equaling fertile
lemma ; [of coastal wetlands and Piedmont
prairies from NC and AR south to the Caribbean and Central America]
16 Larger
vernal blades 5-9 cm long and 2-3 mm wide;
vernal panicles usually ¼ to ½ as wide as long, the branches
ascending ;
spikelets 2.3-3.0 mm long, second
glume and
sterile lemma sparsely
pubescent to
glabrous , summits pointed to blunt, exceeding fertile
lemma ; [of c. and s. FL]
15 Vegetative parts
pubescent , at least in the lower portion of plant; mature
panicles usually more than half as wide as long, the branches spreading-
ascending , the
spikelets not noticeably subsecund;
autumnal blades 1-6 cm long,
involute and < 2 mm wide in
D. aciculare and
D. arenicoloides , or to 9 cm long, flat and 2-4 mm wide in
D. angustifolium (
autumnal blade dimensions not yet known for
D. wilcoxianum );
spikelets 1.5-3.2 mm long.
17 Culms 15-35 cm long; all
culm leaf
blades long-
hirsute on both surfaces, their
sheaths densely
papillose -
hirsute ; leaf
blades of the primary branches often exceeding the immature
vernal panicles
17 Culms 15-75 cm long;
culm leaf
blades glabrous to sparsely
pubescent , or the lower softly
villous abaxially in
D. arenicoloides ; lower
sheaths usually
villous , mid-
culm and upper
sheaths glabrous (though often
ciliate basally); leaf
blades of primary branches not exceeding immature
vernal panicles .
18 Spikelets (3.0-) 3.2-3.8 mm long,
fusiform , pointed at summit,
attenuate at
base , with both
glumes attached 0.3-0.5 mm below
sterile lemma , the
autumnal spikelets 3.5-3.8 mm long; larger
vernal leaf
blades 3-6 mm wide, the lower and mid-
culm leaf
blades of similar width;
autumnal leaf
blades involute
18 Spikelets 1.8-3.1 mm long,
obovate , blunt, and the
base not
attenuate (except in
D. arenicoloides with
autumnal spikelets 2.3-3.1 mm long); larger
vernal leaf
blades 2-8 mm wide, the lower usually wider and often shorter than mid-
culm leaf
blades ;
autumnal leaf
blades involute or flat.
19 Longest
vernal blades to 16 cm, widest
vernal leaf
blades 4-8 mm, usually longitudinally
wrinkled ;
vernal and
autumnal spikelets 2.3-3.1 mm long;
autumnal leaf
blades flat, the larger to 9 cm by 2-4 mm
19 Longest
vernal leaf
blades to 6 cm (
D. aciculare ) or 12 cm (
D. arenicoloides ) long, 2-5 mm wide, not noticeably
wrinkled ;
vernal spikelets 1.5-2.8 mm long,
autumnal spikelets either 1.5-2.3 (
D. aciculare ) or 2.3-3.1 (
D. arenicoloides );
autumnal leaf
blades involute , the larger to 6 cm by 1.5 mm.
20 Longest
vernal leaf
blades to 12 cm, widest
vernal leaf
blades 3-4 (-5) mm;
vernal spikelets 2.1-2.8 mm long;
autumnal spikelets 2.3-3.1 mm long, pointed,
attenuate , the
glumes attached 0.3-0.5 mm below
sterile lemma ; first
glumes 0.7-1.5 mm long; larger
autumnal leaf
blades to 6 cm by 1.5 mm
20 Longest
vernal leaf
blades to 6 (-8) cm, widest
vernal leaf
blades 2-5 mm;
vernal and
autumnal spikelets 1.7-2.3 mm long, blunt, not
attenuate , the
glumes attached < 0.2 mm below
sterile lemma ; first
glumes 0.6-0.9 mm long; larger
autumnal leaf
blades to 3 cm by 1 mm
14 Blades of mid-
culm leaves
lanceolate , thin or firm but not stiff, usually < 10× as long as wide, usually 7 mm or more wide when as much as 8 cm long.
21 Spikelet bases strongly
attenuate ,
apices blunt to beaked; first
glume (2/5-) ½-2/3 (-4/5) as long as
spikelet , attached 0.3-0.5 (-0.7) mm below expansion of second
glume and
sterile lemma
21 Spikelet bases not noticeably
attenuate ,
apices rounded to
acute ; first
glume < ½ as long as
spikelet , attached < 0.3 mm below expansion of second
glume and
sterile lemma .
22 Spikelets 2.9-3.8 mm long, broadly
elliptic , rounded at the summit, with broad and thick
nerves
22 Spikelets 2.1-2.9 mm long,
elliptic or
obovate , rounded or pointed at the summit, the
nerves often raised, but not broad and thick.
24 Culms 50-150 cm long; at least lowest
sheaths papillose -
pilose .
25 Lowest
internodes usually purple-
villous ;
nodes retrorsely (-spreading)
bearded ;
spikelets 1.8-2.2 mm long, moderately densely
pubescent ; first
glume broadly
ovate , subacute, 0.4-0.7 mm long, usually < 1.5 × as long as wide; second
glume about as long as
sterile and fertile
lemmas ; fertile
lemma 1.7-1.8 mm long
25 Lowest
internodes glabrous or
pubescent or spreading
villous , usually not purplish;
nodes glabrous , sparsely
pubescent to sparsely
bearded ;
spikelets 2.2-2.8 (-3.0) mm long, sparsely
pubescent ; first
glume 0.7-1.1 mm long, usually > 1.5 × as long as wide; second
glume shorter than
sterile and fertile
lemmas ; fertile
lemma 2.1-2.2 mm long
24 Culms 18-100 cm long;
sheaths glabrous or the lowest softly
pubescent but not
papillose -
pilose .
26 Spikelets strongly plano-
convex when viewed laterally,
obpyriform when viewed dorsally, (2.1-) 2.2-2.6 mm long; fertile
lemma and
palea papillose or smooth; first
glume usually broadened above
base , rounded or broadly tapered distally;
glume bases often reddish-purple.
27 Culm leaves 2-5 mm wide; fertile
lemma and
palea smooth, shiny
27 Culm leaves 4-12 mm wide; fertile
lemma and
palea papillose -roughened
23 Culm internodes crisp-
puberulent (sparsely so in
D. webberianum) ;
sheaths puberulent or
glabrous .
28 Spikelets elliptic , sub-
acute to pointed, greenish or faintly purple-tinged basally
28 Spikelets strongly plano-
convex when viewed laterally,
obpyriform when viewed dorsally, broadly rounded, usually markedly reddish-purple basally.
29 Fertile
lemma and
palea papillose ;
spikelets 2.2-2.6 mm long; lower
culm blades 6-12 mm wide,
glabrous
29 Fertile
lemma and
palea smooth (minutely
reticulate but not
papillose );
spikelets (1.8) 1.9-2.2 (-2.5) mm long; lower
culm blades 2-8 mm wide,
glabrous ,
glabrate , or
puberulent (especially abaxially)
2 Ligule (1.5-) 2-5 mm long;
sheaths glabrous to variously
pubescent ;
internodes glabrous or
pubescent ;
nodes glabrous , or
bearded with
ascending , spreading, or tangled
hairs ; leaves 3-10 cm long, 3-10 mm wide;
spikelets 0.8-1.9 mm long
2 Ligule 1-2 mm long;
sheaths sparsely to moderately spreading short-
pilose ;
internodes glabrous ;
nodes retrorsely bearded ; leaves 1-4 cm long, 2-5 mm wide;
spikelets 1.2-1.4 mm long
3 Basal leaves
rosette -forming, usually much smaller than the
culm leaves, not matted or cushion-forming;
culms branching at the mid and upper
nodes in age.
4 Blades of mid-
culm leaves typically long and
acuminate ,
linear or narrowly
lanceolate , usually 10-20× as long as wide, only 2-5 mm wide when < 8 cm long.
5 Vernal leaf
blades 9-20 cm long, the larger usually 5-8 mm wide, becoming longer upwards on the
culm , (weakly-) strongly
scabrous on the
adaxial surface;
spikelets sparsely
pubescent to
glabrous
5 Vernal leaf
blades to 11(14) cm long, those on the upper
culm smaller or similar in size to those below, the
adaxial surface not
scabrous except sometimes on the
margins (or
scabrous in D. neuranthum with leaves no more than 5 mm wide and
spikelets moderately
glandular papillose -
pubescent ).
6 Spikelets (
glandular -)
papillose -
pubescent ;
blades 1-3 (-5) mm wide; first
glume 0.8-1.0 mm long;
culms to 4 dm tall
6 Spikelets glabrous ;
blades 3-8 mm wide; first
glume 0.3-1.1 mm long;
culms to 10 dm tall.
7 Leaves 3-8 mm wide;
panicle (8-) 20-40 mm wide; first
glume 0.6-1.1 mm long, blunt to
acute
7 Leaves 3-5.5 mm wide;
panicle 2-5 mm wide; first
glume 0.3-0.4 mm long,
truncate to
obtuse
4 Blades of mid-
culm leaves
lanceolate , mostly 10× or less as long as wide, usually 7 mm or more wide when as much as 8 cm long.
8 Spikelets elliptic ,
oblong , or
obovate ; lower
culm blades 3-12 (-15) mm wide, thin, tapered to the
base ; plants often freely branching in age, becoming top-heavy with a mass of
fascicled , reduced leafy branches and
inflorescences
8 Spikelets broadly
elliptic to suborbicular; lower
culm blades 6-30 mm wide, broad and
cordate to subcordate at the
base ; plants sparingly branched in age, not becoming top-heavy with
fascicled , reduced leafy branches and
inflorescences .
9 Leaf
blades not noticeably thick;
ligules ciliate , to 0.8 mm long;
spikelets 1.8-2.6 mm long,
elliptic ,
obtuse to subacute; first
glumes 0.5-1.0 mm long
9 Leaf
blades thickish;
ligule obsolete or with
cilia < 0.8 mm long;
spikelets 0.9-1.9 mm long,
obovoid to
spherical , rounded; first
glumes 0.2-0.8 mm long.
10 Spikelets 0.9-1.2 mm long; longer
blades 6-8 cm long, erect to erect-
ascending
10 Spikelets 1.2-1.9 mm long; longer
blades 8-20 cm long,
ascending or the uppermost erect.
11 Mid-
culm blades to 25 cm long, 14-30 mm wide, the uppermost 10-15+ cm long;
vernal panicles to 20 cm long, often less than half as wide as long;
spikelets 1.3-1.6 (-1.7)
11 Mid-
culm blades to 10 cm long, 5-11 (-14) mm wide, the uppermost 3-9 cm long;
vernal panicles to 14 cm long, usually more than half as wide as long;
spikelets (1.4-) 1.5-1.8 mm long
3 Basal leaves similar to or larger than the mid and upper
culm leaves, often matted or cushion-forming;
culms branching at the
base (also at mid and upper
nodes in
D. chamaelonche vars. and
D. dichotomum var.
glabrifolium ).
12 Longer
blades > 6 cm; if only 6 cm, then
sheaths retrorsely long-
pilose (
D. laxiflorum ).
13 Spikelets 1.2-1.5 mm long,
glabrous
13 Spikelets 1.7-2.3 (-2.8) mm long,
pubescent .
14 Longer
blades 6-18 cm long by 7-12 mm wide;
sheaths conspicuously
retrorsely long-
pilose ;
nodes bearded with
retrorse or spreading
hairs ;
spikelets (1.4-) 1.9-2.3 (-2.5) mm long
14 Longer
blades 10-35 cm long by 2-6 mm wide;
sheaths glabrous to variously
pilose , but not conspicuously
retrorsely long-
pilose ;
nodes variously
pubescent to
glabrate ;
spikelets 1.7-2.3 (-2.8) mm long.
15 Fresh plants light green to blue-green; leaf
sheaths usually
pilose ;
ligule of largest
culm leaves 3-6× as wide as long;
culm blades to 35 cm long, usually
pilose on the upper surface
15 Fresh plants darker green; leaf
sheaths usually
glabrous ;
ligule of largest
culm leaves usually > 6× as wide as long;
culm blades to 15 cm long, usually
glabrate to
glabrous on the upper surface
16 Blades 1-3 mm wide,
glabrous , eciliate or basally
ciliate ;
spikelets 0.9-1.4 mm long.
17 Spikelets pubescent , 1.2-1.4 mm long;
blades involute , often
falcate , 2.5-6 cm long; [
endemic to c. and s. FL scrub]
17 Spikelets glabrous , 0.9-1.2 mm long;
blades flat, not
falcate , 1.5-4 (-5) cm long; [wet sites, VA s. to FL, w. to e. LA]
16 Blades 3-8 mm wide;
spikelets 1.1-2.1 mm long (if < 1.5 mm, then
blades either
pubescent on one or both surfaces or
ciliate to the
apex ).
18 Spikelets pubescent , 1.5-2.1 mm long;
blade surfaces
glabrous
19 Blades glabrous , or sparsely
pilose only near the
adaxial base ;
spikelets 1.4-2.1 mm long
19 Blades pilose , at least abaxially;
spikelets 1.1-1.6 mm long