1 Pistillate spikes all from the
base of the plant
1 Pistillate spikes all or in part borne on the elongate, aboveground stem.
2 Bracts of the lowermost non-
basal spike with well-developed
sheath > 4 mm long.
3 Beak of
perigynium with distinct teeth > 0.6 mm long
3 Beak of
perigynium entire , notched, or with indistinct teeth < 0.6 mm long.
4 Bracts of the lowermost non-
basal spike bladeless, or with a
blade < 2 mm long.
5 Perigynia 2.0-2.9 mm long; leaf blades 0.9-3.3 mm wide
5 Perigynia 4-5 mm long; leaf blades 4-8 mm wide
4 Bracts of the lowermost non-
basal spike with
blade > 3 mm long (and often much longer).
6 Achene tip with at most a short
apiculus .
7 Leaves usually
glabrous , to
scabrous on the veins.
8 Bases of plants distinctly red or purple.
9 Lowermost
pistillate scales awned; leaves somewhat
septate -
nodulose ; plants usually long-
rhizomatous and forming large
clonal colonies
2 Bracts of the lowermost non-
basal spike sheathless or with
sheath < 4 mm long.
10 Perigynia > 10 mm long.
11 Pistillate spikes
cylindric , much longer than wide;
staminate spikes 1-8
10 Perigynia < 10 mm long.
12 Perigynium beak with 2 teeth > 0.6 mm long
12 Perigynium beak entire or with teeth < 0.6 mm long.
15 Sheaths and blades both
glabrous
15 Sheaths and/or blades
pubescent
13 Terminal
spike staminate (or rarely
androgynous ).
16 Leaf sheaths (and usually the blades as well)
pubescent .
18 Perigynia usually < 3.2 mm long, the
apex rounded and beakless, or abruptly beaked
18 Perigynia > 3.5 mm long, the tip tapering or abruptly beaked.
19 Longer peduncles of
pistillate spikes > 1 cm long;
perigynia > 3× as long as wide, tapering gradually to the
base
19 Longer peduncles of
pistillate spikes 0-1 cm long;
perigynia < 3× as long as wide, abruptly
contracted to a short
stipe at the
base .
20 Perigynia distinctly 20-30-veined;
beak < 0.5 mm long
20 Perigynia veinless except for 2
marginal veins;
beak > 0.7 mm long
16 Leaf sheaths and blades
glabrous to
scabrous .
21 Achene tip with at most a short
apiculus .
22 Leaf blades
scabrous on the upper surface;
beak of
perigynium recurved
22 Leaf blades
glabrous on the upper surface, often with rough margins or tip;
beak of
perigynium straight.
23 Fronts of sheaths of lower leaves ladder-
fibrillose ; leaves and sheaths
septate -
nodulose (sometimes obscurely so)
23 Fronts of leaf sheaths not ladder-
fibrillose , sometimes breaking into
longitudinal fibers; leaves and sheaths not
septate -
nodulose .
24 Perigynia strongly 12-30-veined.
25 Pistillate scales with 3-7 (-10) veins
25 Pistillate scales with 1-3 veins.
26 Leaf blades, at least toward the tip, M-shaped in cross-section when young, the upper surface usually with 2
marginal veins more prominent than the
midvein ;
staminate spikes 1-4
26 Leaf blades V-shaped in cross-section when young, the upper surface lacking 2
marginal veins more prominent than the
midrib ;
staminate spike 1
24 Perigynia 0-12-veined.
27 Proximal pistillate spike erect,
perigynia < 25 per
spike ; terminal
staminate spikes 1; [native].
28 Plants with at least some
pistillate spikes
basal ; culms much shorter than the leaves
28 Plants with most
pistillate spikes on obvious elongated stems; culms shorter than or longer than the leaves.
29 Upper leaves with blades > 2 cm long, longer than the sheaths
29 Upper leaves bladeless or with blades < 1 cm long and also shorter than the sheaths
1 Apex of
perigynium beak with 2 teeth at least (0.4-) 0.5 mm long.
2 Perigynia with 2 distinct
marginal veins, otherwise veinless or only very faintly veined; leaves not
septate -
nodulose
2 Perigynia with 5+ distinct veins; at least
proximal leaves
septate -
nodulose , rarely not.
3 Perigynia with
serrulate wing on margins; spikes usually 20+,
sessile ,
pistillate or
androgynous and similar in appearance, forming a dense,
ovoid or
oblong head, rarely with
proximal spike separated
3 Perigynia without wing on margins; spikes not more than 10, usually at least
proximal shortly
pedunculate , sometimes subsessile,
distal and
proximal spikes usually dissimilar in appearance, usually not
crowded into a dense head.
4 Pistillate scales obtuse to
acuminate , awnless or at most with a rough
apiculus .
5 Staminate spikes usually 1-3+;
perigynia 6-15 (-22)-veined, each 4-10 (-12.5) mm.
6 Perigynia 6-14 (-15)-veined
7 Perigynia narrowly
ovate to
subglobose , 2-2.5× as long as wide
7 Perigynia elliptic to
ovate , 3-4.5× as long as wide
4 Pistillate scales , at least some, with a
scabrous awn .
8 Perigynia (9-) 10+ mm long.
9 Pistillate spikes
cylindric , much longer than wide;
staminate spikes (1-) 2-6
9 Pistillate spikes
globose ,
ovoid , or
cylindric , about as long as wide;
staminate spikes usually 1
10 Upper
pistillate scales with
awn > ½ as long as the body
10 Upper
pistillate scales acute to short-awned,
awn < ½ as long as the body.
11 Beak of
perigynia with teeth 1.1-3 mm long
11 Beak of
perigynia with teeth often < 1 mm long.
12 Staminate spikes 1 (-4);
perigynium beak > 1.7 mm long
12 Staminate spikes 1-6;
perigynium beak often < 1.7 mm long.
13 Perigynia veined only at
base ; widest leaves < 4 mm wide
13 Perigynium veined to tip of body and often into
beak ; widest leaves mostly > 4 mm wide.
14 Pistillate scales with margins
serrulate distally;
awn , when present, rough.
15 Perigynia 3.1-4.8 × 0.9-1.5 (-1.8) mm;
staminate spikes 1-2
15 Perigynia 2.5-8 × 1.2-3.5 mm;
staminate spikes usually 3-7
14 Pistillate scales with margins
entire ;
awn , when present, usually smooth.
16 Perigynia 14-25-veined.
17 Perigynia narrowly
ovate to
subglobose , 2-2.5× as long as wide
17 Perigynia elliptic to
ovate , 3-4.5× as long as wide
1 Apex of
perigynium beak entire ,
emarginate , or with 2 teeth < 0.5 mm long.
18 Perigynia minutely
papillose , at least distally (at 20× magnification).
19 Some roots with yellow-brown felty covering
19 Roots brown or black, without yellow-brown felty covering, rarely with white felty covering.
20 Leaf blades and/or sheaths
pubescent , at least at junction of
blade and
sheath
20 Leaf blades and sheaths
glabrous .
22 Lower
pistillate scales with
awn 1.2-3.5 mm long;
staminate scales awned
18 Perigynia not
papillose , essentially smooth.
24 Perigynia with distinct
beak 0.5-4 mm long.
25 Larger spikes usually with > 50
perigynia ; achenes 2-3 mm long
25 Larger spikes with < 40
perigynia ; achenes 1-2 mm long
24 Perigynia beakless or with
beak < 0.5 mm long.
26 Adaxial side of leaves with 2
marginal veins more prominent than
midvein ; young leaves M-shaped in cross section.
27 Perigynia ovoid , usually > 1.5× as long as wide;
apex tapered to
beak ; sheaths of
proximal leaves not
septate -
nodulose
27 Perigynia subcircular, < 1.2× as long as wide;
apex rounded, abruptly beaked; sheaths of
proximal leaves
septate -
nodulose
26 Adaxial side of leaves without 2
marginal veins more prominent than
midvein ; young leaves V-shaped or rounded in cross section.
28 Leaf blades and/or sheaths
pubescent , at least at junction of
blade and
sheath
28 Leaf blades and sheaths
glabrous .
29 Perigynia ascending , spreading at about right angles, or
reflexed when mature
30 Leaves and/or sheaths
pubescent , at least at junction of
blade and
sheath .
31 Larger leaves 8-23 mm wide, only the sheaths
pubescent
31 Leaves < 8 mm wide, blades and usually also the sheaths
pubescent .
32 Perigynium beak 0.5-3 mm long, often > 1 mm long, about 1/2 the length of the body;
proximal pistillate scales awned
32 Perigynium beak absent or not more than 0.5 (-0.7) mm long, not > 1/4 the length of the body;
proximal pistillate scales acute ,
acuminate or
cuspidate .
30 Leaves and sheaths usually
glabrous , rarely
papillose .
33 Style persistent on
achene in fruit; larger leaves and sheaths usually at least sparsely
septate -
nodulose , rarely not.
34 Perigynia (9-) 10+ mm long;
staminate spike usually 1
34 Perigynia < 10 mm long;
staminate spikes 1-5 (-7).
35 Pistillate scales with margins
serrulate distally,
apex usually rough-awned.
36 Distal pistillate scales with
apex acute to short-awned,
awn < 1/2 as long as the body
36 Distal pistillate scales with
apex long-awned,
awn > 1/2 as long as the body
35 Pistillate scales with margins
entire ,
apex awnless or with short, smooth
awn .
37 Perigynia not
inflated , thick walled, brownish, dull
37 Perigynia slightly to strongly
inflated , thin walled, yellowish to purplish, shiny
33 Style deciduous ; larger leaves and sheaths sometimes
septate -
nodulose , more often not.
38 Leaf blades, at least widest, M-shaped in cross section when young,
adaxial surface with 2
marginal veins more prominent than
midvein .
39 Widest leaf blades 8-23 mm wide,
septate -
nodulose
39 Widest leaf blades < 6 mm wide, not
septate -
nodulose .
40 Pistillate scales awned, longest
awn > 0.5 mm long
40 Pistillate scales obtuse to
acuminate or short-awned,
awn < 0.2 mm long
38 Leaf blades V-shaped in cross section when young,
adaxial surface without 2
marginal veins more prominent than
midvein .
41 Proximal perigynia in each
spike spreading at right angles or
reflexed at maturity; leaf blades and sheaths sparsely
septate -
nodulose
41 Proximal perigynia in each
spike ascending or spreading-
ascending ; leaf blades and sheaths not
septate -
nodulose .
42 Plant
base brown or blackish, without trace of red or purple
42 Plant
base red or purple tinged, sometimes only sparsely so.
43 Perigynia distinctly veined on faces;
pistillate scales brown or black
43 Perigynia veinless or with veins only proximally;
pistillate scales white-
hyaline or red-brown
1 Apex of
perigynium beak terminated by 2 teeth, mostly > 0.5 mm long.
2 Perigynia > 4× as long as wide (8-15 × 1-3 mm).
3 Apical teeth of
perigynium beak erect; achenes
oblong -
obovoid or
obovoid ; at least
proximal pistillate scales with 3-5 (-7)-veined center
3 Apical teeth of
perigynium beak reflexed ; achenes elongate;
pistillate scales with 1-veined center
2 Perigynia < 4× as long as wide.
4 Perigynia > (9-) 10 mm long.
5 Sheaths, at least the
proximal sheath fronts, densely
tomentose at mouth;
apical teeth of
perigynium beak often > 1 mm long
5 Sheaths
glabrous ;
apical teeth of
perigynium beak not more than 1 mm.
6 Perigynia 7-11-, 5-12-, or 12-25-veined
6 Perigynia 12-34-veined.
7 Basal and
proximal leaf sheaths yellowish to brown, without trace of red or purple
4 Perigynia < 10 mm long.
8 At least
proximal pistillate scales with long, rough
awn .
9 Perigynia with 2 strong
marginal veins, otherwise veinless or veined only proximally; leaves not
septate -
nodulose .
9 Perigynia with 5+ strong veins extending length of bodies; leaves
septate -
nodulose .
10 Perigynium beak with
apical teeth (0.4-) 0.6-3 mm long, often > 1 mm long
10 Perigynium beak with
apical teeth < 1 mm long.
11 Perigynium body
obovoid , widest distally;
proximal bract > 3× as long as the
inflorescence
11 Perigynium body
ovoid or
lanceoloid or
ellipsoid , widest at middle or proximally;
proximal bract usually < 2× as long as the
inflorescence .
12 Perigynium beak 0.9-1.7 mm long; mature
perigynia dull
12 Perigynium beak 0.2-6 mm long; mature
perigynia somewhat
glossy
13 Culms with some red or purple at
base .
14 At least
proximal perigynia in each
spike spreading or
reflexed ; leaves sparsely
septate -
nodulose
13 Culms brown or black at
base , without trace of red or purple.
15 Pistillate scales with 5-7-veined center
15 Pistillate scales with 1-3-veined center.
16 Leaves not
septate -
nodulose
16 Leaves at least sparingly
septate -
nodulose
1 Apex of
perigynium beak entire ,
emarginate , or with teeth mostly < 0.5 mm long.
17 Leaf blades variously hairy.
18 Plant brown or black at
base
18 Plant with at least some red or purple at
base
19 Bracts without blades.
20 Leaves much wider, often over 10 mm wide
19 Bracts, at least the
proximal , with distinct
blade .
21 Young leaves V-shaped or rounded in cross section,
adaxial surface without 2
marginal veins more prominent than
midvein or other veins.
22 Perigynium beak with 2 distinct terminal teeth, usually > 0.2 mm long.
23 At least
proximal pistillate scales awned,
awn at least 1/2 as long as body; leaves conspicuously
septate -
nodulose .
24 Base of
culm strongly red or purple; leaves 2-4 mm wide
24 Base of
culm brown, without or with only trace of red or purple; leaves 4-15 mm wide.
25 Distal pistillate scales acute ,
acuminate , or short-awned,
awn less than 1/2 length of body
25 Distal pistillate scales with
awn at least about as long as body
23 Pistillate scales obtuse to
acuminate , sometimes very shortly awned; leaves not or only weakly
septate -
nodulose .
26 Plants with long-creeping rhizomes; leaves usually not
septate -
nodulose
26 Plants with short-
ascending rhizomes; leaves at least sparingly
septate -
nodulose .
22 Perigynium beak entire ,
emarginate , or with 2
apical teeth < 0.2 mm long.
28 Perigynia rounded at
base , sides proximally distinctly
convex .
29 Perigynia with veins slightly prominent
29 Perigynia with veins slightly
impressed
28 Perigynia tapering at
base , sides straight or only slightly
convex proximally.
30 Perigynia rounded at
apex , beakless or with
beak < 0.3 mm long.
31 Widest leaf blades < 5 mm wide; plants usually not
cespitose
31 Widest leaf blades usually > 5 mm wide; plants usually
cespitose
30 Perigynia tapering to
beak at least > (0.3-) 0.5 mm long.
32 Proximal perigynia in each
spike spreading at right angles or
reflexed at maturity; leaves and sheaths sparsely
septate -
nodulose
32 Proximal perigynia in each
spike ascending or spreading-
ascending ; leaves and sheaths not
septate -
nodulose .
33 Plants usually
colonial ;
perigynia and leaves often
papillose
33 Plants
cespitose ;
perigynia and leaves not
papillose
21 Leaf blades M-shaped in cross section when young,
adaxial surface with 2
marginal veins more prominent than
midvein and other
marginal veins, sometimes apparent only on
proximal leaves and on
proximal part of
blade .
34 Perigynium beakless or with
beak < 0.5 mm long,
entire or with 2
apical teeth < 0.1 mm long.
35 Widest leaf blades (usually
basal ) > 10 mm wide.
36 Distal pistillate spikes erect,
perigynia < 30 per
spike ; [native].
37 Perigynia trigonous with rounded angles in cross section
37 Perigynia sharply
trigonous in cross section
35 Widest leaf blades < 10 mm wide.
38 Perigynia not prominently veined, veins somewhat
impressed
38 Perigynia at least slightly prominently veined or veinless except for 2
marginal veins.
39 Perigynia rounded at
base , sides proximally
convex
39 Perigynia tapering at
base , sides proximally straight,
concave or slightly
convex .
40 Perigynia transversely
wrinkled ; leaf sheaths sparsely
septate -
nodulose
40 Perigynia not transversely
wrinkled ; leaf sheaths not
septate -
nodulose .
41 Perigynia sharply
trigonous in cross section
41 Perigynia terete or with rounded angles in cross section.
42 Perigynia with 2
marginal veins and not more than 10 veins on faces
42 Perigynia with more than 12 veins.
43 Plants usually
colonial , not
cespitose ;
perigynia papillose at least faintly.
34 Perigynium with distinct
beak > 0.5 mm, usually with
apical teeth > 0.1 mm long.
44 Perigynia with 2 distinct
marginal veins, otherwise veinless or with very weak veins proximally.
45 Perigynium beak entire ,
emarginate , or with
apical teeth < 0.1 mm long
45 Perigynium beak with
apical teeth 0.2-1 mm.
46 Perigynium beak usually < 1 mm long.
46 Perigynium beak sometimes > 2 mm long
44 Perigynia with more than 2 veins extending most of length of bodies.
47 Pistillate scales with narrow, indistinct bodies, scarcely enlarged at bases
47 Pistillate scales with wide, flat bodies, sometimes terminating in awns.
49 Perigynium with not more than 5 veins extending length of body on
abaxial face, veinless or weakly veined on
adaxial face
49 Perigynium with > 8 equally prominent veins on both faces.
50 Perigynia bluntly angled in cross section
50 Perigynia sharply angled in cross section
48 Perigynium beak with 2 distinct teeth at
apex , usually > 0.2 mm long.
51 Pistillate spikes, at least the
proximal ,
lax ,
proximal and middle
perigynia separated by internodes at least 1/4 their length; leaves and sheaths not
septate -
nodulose
51 Pistillate spikes dense,
proximal and middle
perigynia separated by internodes less than 1/10 their length;
proximal leaves and sheaths sparingly
septate -
nodulose .
52 Pistillate scales , at least
proximal , awned,
awn at least 1/4 of
scale body length.
53 Distal pistillate scales acuminate or with
awn less than 1/2 body length;
staminate spikes usually 2+
53 Distal pistillate scales awned,
awn usually more than 1/2 body length;
staminate spikes 1, sometimes with 1 much smaller
spike laterally
52 Pistillate scales obtuse to
acute or
apiculate ,
apiculus not more than 1/10 of body length.
54 Culms purple-red at
base ; achenes with
style persistent
54 Culms brown at
base , without trace of red or purple; achenes with
style deciduous