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Key to Carex, Key C: Subkey in Carex

Cyperaceae

Carex

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1 Perigynia glabrous, papillate or not.
  2 Lateral spikes usually pedunculate; lowermost inflorescence bracts sometimes with sheath; peduncles with prophyll at base.
    3 Terminal spike gynecandrous, pistillate flowers as many as or more numerous than staminate; lateral spikes short, not much longer than wide; fresh perigynia white-pulverulent
    3 Terminal spike usually staminate or, sometimes, gynecandrous, staminate flowers then more numerous than pistillate; lateral spikes oblong, distinctly longer than wide; fresh perigynia green or slightly glaucous.
      4 Pistillate scales (at least the lower) long-awned
      4 Pistillate scales obtuse to acuminate or cuspidate.
        5 Perigynia smooth; style persistent on the achene
        5 Perigynia often papillose over most of the surface; style deciduous
  2 Lateral spikes sessile; bracts sheathless; peduncles without (or rarely with) a prophyll
          6 Perigynia papillose (visible at 20× magnification).
             7 Terminal spike staminate, androgynous, or gynecandrous (if gynecandrous, the staminate flowers more numerous than the pistillate); lateral spikes at least 2× as long as wide
             7 Terminal spike staminate or gynecandrous (if gynecandrous, the pistillate flowers more numerous than the staminate); lateral spikes not much longer than wide
          6 Perigynia smooth.
               8 Terminal spike gynecandrous; lateral spikes gynecandrous or pistillate.
                       12 Margins of perigynia flat, at least in the upper ½, flat portion (measured at the tip of the achene and base of beak) > (0.1-) 0.2 mm wider
                          13 Achenes rounded at apex (style dehiscing at the surface of the achene); style conspicuously enlarged at the base
                          13 Achenes with short apiculus formed by the persistent base of the style; style not conspicuously enlarged at base
                       12 Margins of perigynia rounded, or with flat portion < 0.1 mm wide.
                            14 Margins of perigynia rounded or with a very narrow rounded edge; achenes nearly filling the perigynium bodies
                            14 Margins of perigynia sharply edged or narrowly winged; achenes distinctly smaller than the perigynium bodies.
                              15 Inflorescences in fruit 1-1.5× as long as wide
                              15 Inflorescences in fruit 1.5-2 (or more)× as long as wide.
                                16 Lowermost perigynia in each spike spreading
                                16 Lowermost perigynia in each spike ascending or erect.
                                  17 Perigynium serrulate on the margins of the upper body and lower beak
                                  17 Perigynium entire on the margins of the upper body and the lower beak
               8 Terminal spike androgynous (rarely entirely staminate or entirely pistillate); lateral spikes androgynous, staminate, or pistillate.
                 9 Sheath fronts of lower cauline leaves transversely rugose.
                   10 Perigynia mostly > 2× as long as wide, widest near the base
                   10 Perigynia mostly < 2× as long as wide, widest near middle.
                     11 Inflorescence usually branched, at least at the base, usually with > 15 spikes; pistillate scales usually yellow or brown, sometimes with hyaline margins, 3-veined
                     11 Inflorescence unbranched or with 1 or 2 short branches at the base, with < 15 spikes; pistillate scales greenish hyaline, 1-veined
                 9 Sheath fronts of lower cauline leaves smooth (or very weakly and indistinctly transversely rugose).
                                    18 Fronts of leaf sheaths dotted red, brown, or yellow.
                                       19 Perigynia widest near the base; culms usually > 1 mm wide
                                       19 Perigynia widest near the middle; culms usually < 1 mm wide.
                                         20 Plants densely cespitose, with short rhizomes; pistillate scales acute to acuminate
                                         20 Plants loosely cespitose, sometimes with long rhizomes; pistillate scales (at least the upper) obtuse
                                    18 Fronts of leaf sheaths not dotted red, brown, or yellow.
                                           21 Perigynium beak < 0.25 mm long, the margins entire
                                           21 Perigynium beak > 0.25 mm long, the margins often serrulate.
                                             22 Upper leaves of culms with front of sheaths green-veined, not differentiated from the rest of the sheath
                                             22 Upper leaves of culms with front of sheaths with at least a narrow hyaline or whitish-hyaline band extending at least ½ the length of the sheath.
                                               23 Perigynia with flat, winglike margins > 0.1 mm wide; plants long-rhizomatous, not cespitose, sometimes forming large colonies
                                               23 Perigynia without a flat margin, or with a flat margin < 0.1 mm wide; plants short-rhizomatous or inconspicuously rhizomatous, cespitose or not, sometimes forming large colonies.
                                                 24 Plants colonial from long rhizomes or stoloniferous (proliferating from last year’s decumbent sterile culms)
                                                    25 Plants long stoloniferous, proliferating from last year’s decumbent sterile culms
                                                    25 Plants rhizomatous from creeping rhizomes 1.8-3.5 mm in diameter
                                                 24 Plants cespitose.
                                                      26 Spikes not consistently androgynous, the terminal either entirely staminate or pistillate, the lateral spikes irregularly pistillate, or staminate, or mixed
                                                      26 Spikes consistently androgynous, occasionally some of the lateral spikes entirely pistillate.
                                                        27 Perigynium widest near the base, tapering from base to beak
                                                        27 Perigynium widest above the base, often abruptly beaked

Key to Carex, Key D: Subkey in Carex

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1 Pistillate spikes all or in part borne on the elongate, aboveground stem.
  2 Bracts of the lowermost non-basal spike with well-developed sheath > 4 mm long.
    3 Beak of perigynium with distinct teeth > 0.6 mm long
    3 Beak of perigynium entire, notched, or with indistinct teeth < 0.6 mm long.
      4 Bracts of the lowermost non-basal spike bladeless, or with a blade < 2 mm long.
        5 Perigynia 2.0-2.9 mm long; leaf blades 0.9-3.3 mm wide
        5 Perigynia 4-5 mm long; leaf blades 4-8 mm wide
      4 Bracts of the lowermost non-basal spike with blade > 3 mm long (and often much longer).
          6 Achene tip with persistent, enlarged, circular style base
          6 Achene tip with at most a short apiculus.
             7 Leaves pubescent or pilose
             7 Leaves usually glabrous, to scabrous on the veins.
               8 Bases of plants brown
               8 Bases of plants distinctly red or purple.
                 9 Lowermost pistillate scales awned; leaves somewhat septate-nodulose; plants usually long-rhizomatous and forming large clonal colonies
                 9 Lowermost pistillate scales obtuse to acuminate; leaves not septate-nodulose; plants cespitose, short-rhizomatous
  2 Bracts of the lowermost non-basal spike sheathless or with sheath < 4 mm long.
                   10 Perigynia > 10 mm long.
                     11 Pistillate spikes globose, about as long as wide; staminate spikes usually 1
                     11 Pistillate spikes cylindric, much longer than wide; staminate spikes 1-8
                   10 Perigynia < 10 mm long.
                       12 Perigynium beak with 2 teeth > 0.6 mm long
                       12 Perigynium beak entire or with teeth < 0.6 mm long.
                          13 Terminal spike gynecandrous or pistillate
                            14 Perigynia papillate
                            14 Perigynia glabrous or pubescent.
                              15 Sheaths and blades both glabrous
                              15 Sheaths and/or blades pubescent
                          13 Terminal spike staminate (or rarely androgynous).
                                16 Leaf sheaths (and usually the blades as well) pubescent.
                                  17 Pistillate scales sometimes pubescent; pistillate spikes with 40-200 perigynia
                                  17 Pistillate scales glabrous; pistillate spikes with < 40 (-50) perigynia.
                                    18 Perigynia usually < 3.2 mm long, the apex rounded and beakless, or abruptly beaked
                                    18 Perigynia > 3.5 mm long, the tip tapering or abruptly beaked.
                                       19 Longer peduncles of pistillate spikes > 1 cm long; perigynia > 3× as long as wide, tapering gradually to the base
                                       19 Longer peduncles of pistillate spikes 0-1 cm long; perigynia < 3× as long as wide, abruptly contracted to a short stipe at the base.
                                         20 Perigynia distinctly 20-30-veined; beak < 0.5 mm long
                                         20 Perigynia veinless except for 2 marginal veins; beak > 0.7 mm long
                                16 Leaf sheaths and blades glabrous to scabrous.
                                           21 Achene tip with persistent, enlarged, circular style base
                                           21 Achene tip with at most a short apiculus.
                                             22 Leaf blades scabrous on the upper surface; beak of perigynium recurved
                                             22 Leaf blades glabrous on the upper surface, often with rough margins or tip; beak of perigynium straight.
                                               23 Fronts of sheaths of lower leaves ladder-fibrillose; leaves and sheaths septate-nodulose (sometimes obscurely so)
                                               23 Fronts of leaf sheaths not ladder-fibrillose, sometimes breaking into longitudinal fibers; leaves and sheaths not septate-nodulose.
                                                 24 Perigynia strongly 12-30-veined.
                                                    25 Pistillate scales with 3-7 (-10) veins
                                                    25 Pistillate scales with 1-3 veins.
                                                      26 Leaf blades, at least toward the tip, M-shaped in cross-section when young, the upper surface usually with 2 marginal veins more prominent than the midvein; staminate spikes 1-4
                                                      26 Leaf blades V-shaped in cross-section when young, the upper surface lacking 2 marginal veins more prominent than the midrib; staminate spike 1
                                                 24 Perigynia 0-12-veined.
                                                        27 Proximal pistillate spike pendulous, perigynia usually > 30 per spike; terminal staminate spikes 1-3; [exotic]
                                                        27 Proximal pistillate spike erect, perigynia < 25 per spike; terminal staminate spikes 1; [native].
                                                          28 Plants with at least some pistillate spikes basal; culms much shorter than the leaves
                                                          28 Plants with most pistillate spikes on obvious elongated stems; culms shorter than or longer than the leaves.
                                                            29 Upper leaves with blades > 2 cm long, longer than the sheaths
                                                            29 Upper leaves bladeless or with blades < 1 cm long and also shorter than the sheaths

Key to Carex, Key E: Subkey in Carex

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1 Apex of perigynium beak with 2 teeth at least (0.4-) 0.5 mm long.
  2 Perigynia with 2 distinct marginal veins, otherwise veinless or only very faintly veined; leaves not septate-nodulose
  2 Perigynia with 5+ distinct veins; at least proximal leaves septate-nodulose, rarely not.
    3 Perigynia with serrulate wing on margins; spikes usually 20+, sessile, pistillate or androgynous and similar in appearance, forming a dense, ovoid or oblong head, rarely with proximal spike separated
    3 Perigynia without wing on margins; spikes not more than 10, usually at least proximal shortly pedunculate, sometimes subsessile, distal and proximal spikes usually dissimilar in appearance, usually not crowded into a dense head.
      4 Pistillate scales obtuse to acuminate, awnless or at most with a rough apiculus.
        5 Staminate spike usually 1; perigynia 15-20-veined, each 10-20 mm long
        5 Staminate spikes usually 1-3+; perigynia 6-15 (-22)-veined, each 4-10 (-12.5) mm.
          6 Perigynia 6-14 (-15)-veined
          6 Perigynia 14-25-veined
             7 Perigynia narrowly ovate to subglobose, 2-2.5× as long as wide
             7 Perigynia elliptic to ovate, 3-4.5× as long as wide
      4 Pistillate scales, at least some, with a scabrous awn.
               8 Perigynia (9-) 10+ mm long.
                 9 Pistillate spikes cylindric, much longer than wide; staminate spikes (1-) 2-6
                 9 Pistillate spikes globose, ovoid, or cylindric, about as long as wide; staminate spikes usually 1
               8 Perigynia < 9 mm long.
                   10 Upper pistillate scales with awn > ½ as long as the body
                   10 Upper pistillate scales acute to short-awned, awn < ½ as long as the body.
                     11 Beak of perigynia with teeth 1.1-3 mm long
                     11 Beak of perigynia with teeth often < 1 mm long.
                       12 Staminate spikes 1 (-4); perigynium beak > 1.7 mm long
                       12 Staminate spikes 1-6; perigynium beak often < 1.7 mm long.
                          13 Perigynia veined only at base; widest leaves < 4 mm wide
                          13 Perigynium veined to tip of body and often into beak; widest leaves mostly > 4 mm wide.
                            14 Pistillate scales with margins serrulate distally; awn, when present, rough.
                              15 Perigynia 3.1-4.8 × 0.9-1.5 (-1.8) mm; staminate spikes 1-2
                              15 Perigynia 2.5-8 × 1.2-3.5 mm; staminate spikes usually 3-7
                            14 Pistillate scales with margins entire; awn, when present, usually smooth.
                                16 Perigynia 9-15-veined
                                16 Perigynia 14-25-veined.
                                  17 Perigynia narrowly ovate to subglobose, 2-2.5× as long as wide
                                  17 Perigynia elliptic to ovate, 3-4.5× as long as wide
1 Apex of perigynium beak entire, emarginate, or with 2 teeth < 0.5 mm long.
                                    18 Perigynia minutely papillose, at least distally (at 20× magnification).
                                       19 Some roots with yellow-brown felty covering
                                       19 Roots brown or black, without yellow-brown felty covering, rarely with white felty covering.
                                         20 Leaf blades and/or sheaths pubescent, at least at junction of blade and sheath
                                         20 Leaf blades and sheaths glabrous.
                                           21 Terminal spike gynecandrous
                                           21 Terminal spike staminate or androgynous.
                                             22 Pistillate scales obtuse to acuminate or with awn < 1 mm long; staminate scales not awned
                                             22 Lower pistillate scales with awn 1.2-3.5 mm long; staminate scales awned
                                    18 Perigynia not papillose, essentially smooth.
                                               23 Terminal spike gynecandrous.
                                                 24 Perigynia with distinct beak 0.5-4 mm long.
                                                    25 Larger spikes usually with > 50 perigynia; achenes 2-3 mm long
                                                    25 Larger spikes with < 40 perigynia; achenes 1-2 mm long
                                                 24 Perigynia beakless or with beak < 0.5 mm long.
                                                      26 Adaxial side of leaves with 2 marginal veins more prominent than midvein; young leaves M-shaped in cross section.
                                                        27 Perigynia ovoid, usually > 1.5× as long as wide; apex tapered to beak; sheaths of proximal leaves not septate-nodulose
                                                        27 Perigynia subcircular, < 1.2× as long as wide; apex rounded, abruptly beaked; sheaths of proximal leaves septate-nodulose
                                                      26 Adaxial side of leaves without 2 marginal veins more prominent than midvein; young leaves V-shaped or rounded in cross section.
                                                          28 Leaf blades and/or sheaths pubescent, at least at junction of blade and sheath
                                                          28 Leaf blades and sheaths glabrous.
                                                            29 Perigynia erect or ascending
                                                            29 Perigynia ascending, spreading at about right angles, or reflexed when mature
                                               23 Terminal spike staminate or androgynous.
                                                              30 Leaves and/or sheaths pubescent, at least at junction of blade and sheath.
                                                                 31 Larger leaves 8-23 mm wide, only the sheaths pubescent
                                                                 31 Leaves < 8 mm wide, blades and usually also the sheaths pubescent.
                                                                   32 Perigynium beak 0.5-3 mm long, often > 1 mm long, about 1/2 the length of the body; proximal pistillate scales awned
                                                                   32 Perigynium beak absent or not more than 0.5 (-0.7) mm long, not > 1/4 the length of the body; proximal pistillate scales acute, acuminate or cuspidate.
                                                              30 Leaves and sheaths usually glabrous, rarely papillose.
                                                                     33 Style persistent on achene in fruit; larger leaves and sheaths usually at least sparsely septate-nodulose, rarely not.
                                                                       34 Perigynia (9-) 10+ mm long; staminate spike usually 1
                                                                       34 Perigynia < 10 mm long; staminate spikes 1-5 (-7).
                                                                         35 Pistillate scales with margins serrulate distally, apex usually rough-awned.
                                                                           36 Distal pistillate scales with apex acute to short-awned, awn < 1/2 as long as the body
                                                                           36 Distal pistillate scales with apex long-awned, awn > 1/2 as long as the body
                                                                         35 Pistillate scales with margins entire, apex awnless or with short, smooth awn.
                                                                              37 Perigynia not inflated, thick walled, brownish, dull
                                                                              37 Perigynia slightly to strongly inflated, thin walled, yellowish to purplish, shiny
                                                                     33 Style deciduous; larger leaves and sheaths sometimes septate-nodulose, more often not.
                                                                                38 Leaf blades, at least widest, M-shaped in cross section when young, adaxial surface with 2 marginal veins more prominent than midvein.
                                                                                  39 Widest leaf blades 8-23 mm wide, septate-nodulose
                                                                                  39 Widest leaf blades < 6 mm wide, not septate-nodulose.
                                                                                    40 Pistillate scales awned, longest awn > 0.5 mm long
                                                                                    40 Pistillate scales obtuse to acuminate or short-awned, awn < 0.2 mm long
                                                                                38 Leaf blades V-shaped in cross section when young, adaxial surface without 2 marginal veins more prominent than midvein.
                                                                                      41 Proximal perigynia in each spike spreading at right angles or reflexed at maturity; leaf blades and sheaths sparsely septate-nodulose
                                                                                      41 Proximal perigynia in each spike ascending or spreading-ascending; leaf blades and sheaths not septate-nodulose.
                                                                                        42 Plant base brown or blackish, without trace of red or purple
                                                                                        42 Plant base red or purple tinged, sometimes only sparsely so.
                                                                                           43 Perigynia distinctly veined on faces; pistillate scales brown or black