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4 Plants perennial, often with woody base; leaves and stems glabrescent; corollas white, purple, or yellow, with single, double, or triple whorls; fruits spineless, pericarp tuberculate | |
5 Corolla limbs with 5 acuminate lobes longer than the 5 lobules; calyces canescent-puberulent (especially along veins); abaxial leaf surface canescent-puberulent (especially along veins), trichomes short-appressed or curved |