Click the number at the start of a key lead to highlight both that lead and its corresponding lead. Click again to show only the two highlighted leads. Click a third time to return to the full key with the selected leads still highlighted.
Key to Cyperus, Key ZD: Key to Cyperus, subgenus Cyperus -- Spikelets quadrangular to subterete; scales 1-8, persistent rachillas deciduous, articulate at the base
1 Culms, rays, and leaves bluish-glaucous, papillose; leaves harshly scabrid on margins and midrib; spikes with numerous well-developed branches or secondary rays; spikelets dark reddish brown at maturity
1 Culms, rays, and leaves green, not papillose; leaves glabrous to scabridulous; spikes solitary or with a few reduced basal branches; spikelets various.
2 All but the uppermost spikelets strongly (-weakly in some forms of C. hystricinus) reflexed; scales (3.8-) 4-4.9 (-5.4) mm long, terminal involute and prolonged distally, midrib often bent downward; achenes linear to linear-oblong, (2.2-) 2.5-3 (-2.3) mm long; [section Umbellati, subsection Longistylis, C. retrofractus complex]
3 Spikes mostly at least 2× as long as wide, slightly (or not) narrowed to base, spikelets reflexed to spreading; culms, leaves, and bracts glabrous except for marginal prickles on leaves and bracts; rachilla wing claspingachene for its entire length (concealing it in abaxial view); floral scales golden-brown when mature
4 Rays scabrid throughout their lengths; leaves and bracts puberulent; longest ray equal to or longer than the longest bract; culms roundly trigonous to subterete; terminal scalespinose distally; scales 1-2 (-3); spikes tight, burlike, turbinate
6 Spikes dense, the rachis obscured when pressed; achenes lacking a stipitatebase, apiculate at the apex; [native of Mexico and Central America, introduced northward along the Gulf and Atlantic coasts]
6 Spikes relatively loose, the rachis usually visible when pressed; achenes cuneate to a stipitatebase, broadly rounded to acute at the apex; [native, s. TX, uncertain reports]
7 Spikelets very closely spaced, obscuring the rachis in life and when pressed; [section Umbellati, subsection Brevistylis]
8 Spikes broadly and appearing (because of longer spikelets) relatively loosely ellipsoid to ovoid-cylindric, 2.5-7 cm long × 1.5-6 cm wide; spikelets (11-) 12-25 (-28) mm long; achenes 2.2-3 mm long (if < 2.5 mm long, noticeably falcate).
11 Terminal scale of spikelet reduced, sterile, involute, with a minute but distinct mucro to 0.3 mm long, usually uncinate (hooked); fertile scales (if more than one) not overlapping on the same side of the rachilla, usually red-brown or orangeish laterally at maturity; leaf blades 1-3 mm wide; [Cyperus retroflexus complex]
12 Fertile floral scales 1.9-3.0 (-3.3) mm long; rachilla wing usually membranaceous throughout; rachilla lacking lateral nerves; longest spikelets 2.8-5.8 (-8.0) mm long; terminal sterile floral scale of spikelet often much reduced, less than 2/3 the length of fertile floral scales; longest peduncle less than 2.7 (-3.9) cm long; plants diminutive, culms 3-35 (-45) cm tall
12 Fertile floral scales (2.8-) 3.0-3.9 mm long; rachilla wing usually chartaceous beyond clasped achene angle, border membranaceous; rachilla with two lateral nerves, one along each side of the median; longest spikelets 4.9-9.0 (-11.3) mm long; terminal sterile floral scale usually not greatly reduced, 2/3 or more the length of fertile floral scales; longest peduncle (0.5-) 2.4-6.8 cm long; plants more robust, culms usually >25 cm tall
11 Terminal scale of spikelet not differentiated; scalesacute, greenish-hyaline to brownish or straw-colored laterally; scales overlapping; leaf blades 1.5-5 mm wide.
13 Spikes relatively loosely ovoid, 8-20 mm wide, with (5-) 10-50 (-70) spikelets, lacking basal branches; spikelets (2.2-) 3.5-10 mm long, angular in ×-section, greenish-hyaline to pale yellow-green; bracts horizontal
13 Spikes densely oblong-ellipsoid to ovoid, 6-8 (-10) mm wide, with (13-) 30-200 (-274) spikelets, occasionally with short basal branches; spikelets 2.2-4 (-4.5) mm, subterete, straw-colored to brown at maturity; bracts horizontal or ascending to erect; [Cyperus retrorsus complex]
14 Achenes oblong, usually abruptly constricted at the ends; spikes ovoid to oblong, to 2× as long as wide (rarely longer in C. retrorsus); scalesappressed, usually reddish-brown at maturity; bracts ascending to erect (sometimes horizontal in C. nashii).
15 Spikes > 1.5× as long as wide, each with (42-) 50-200 (-274) spikelets; achenes 0.4-0.5 (-0.6) mm wide; stems moderately cespitose; [widespread and weedy]
15 Spikes < 1.5 × as long as wide, each with (13-) 30-92 spikelets; achenes 0.5-0.7 mm wide; stems single to slightly cespitose; [Florida scrub, northward and westward in coastal dunes]
16 Rays 8-27; anthers 0.8-1.0 mm long; achenes 1.6-1.7mm long × 0.6-0.7 mm wide
7Spikelet attachments spaced apart at least proximally, rachis not obscured when pressed and usually also visible in life.
17 Fertile floral scales 1-3 (-5), not overlapping on the same side of the spikelet (if more than one); terminal scale reduced, sterile and involute, mucro usually uncinate; achenes (1.7-) 1.9-2.6 mm long; [section Umbellati, Cyperus retroflexus complex]
18 Longest spikelets 10-21 mm long, strongly contorted; distalscales with scabridmidrib and mucro 0.6-1.9 mm long; longest floral scale of spikelet (3.5-) 3.7-4.8 mm long; anthers 0.5-1.3 mm long; achenes > 3× as long as wide; [TX]
18 Longest spikelets usually < 10 mm long, at most somewhat flexuous with curved tips; distalscales with glabrousmidrib and mucro 0.1-0.3 (-0.5) mm long; longest floral scale of spikelet (2.1-) 2.5-3.5 (-4.0) mm long; anthers 0.3-0.5 (-0.6) mm long; achenes 2-3 (-3.3)× as long as wide; [more widespread]
17 Fertile floral scales 3-8, reaching the next scale or overlapping on the same side of the spikelet; terminal scale not involute and sterile, mucro straight; achenes various.
20 Spikelets ±terete; scales 4.5-5.3 mm long, persistent, laterally reddish brown or tinted with reddish brown if paler and stramineous; achenes usually at least half as long as floral scale; achenes linear-oblong, 2.6-3 mm long, 5-6× as long as wide, > 0.5 mm wide; anthers 1-1.5 mm long; spikes usually broadly and convexly ellipsoid or ovoid, usually unbranched (or rarely with 1-2 reduced proximal branches); [section Umbellati, subsection Brevistylis]
20 Spikelets flattened; scales 3.2-4.5 (-6) mm long, laterally golden yellow to stramineous; achenes less than half as long as floral scale; achenes oblong, (1.5-) 1.8-2.4 mm long, 3-4× as long as wide, < 0.4 mm wide; anthers 0.3-0.5 mm long; spikes usually parallel-sided, ovoid-cylindric, frequently branched; [section Strigosi]
24 Floral scales pale olivaceous to yellowish, not mucronate (rarely with a mucro to 0.1 mm long); spikes cylindric, loose, 6-10 spikelets per 5 mm span of the rachis; culms 0.5-1 mm in diameter; leaves 0.8-2.3 (-3) mm wide; [more widespread]; [section Tetragoni]
24 Floral scales straw-colored to reddish-brown, acute or mucronate, mucro up to 0.2 mm long; spikes loosely to densely ovoid to oblong, spikelets 8-21 per 5 mm span of the rachis; culms 1-2.6 mm in diameter; leaves (1-) 3-6 (-10) mm wide; [TX].
25 Floral scales not mucronate; spikes mostly narrowly oblong (usually 2× or more as long as wide); rachis, spikelets, bracteoles, peduncles, and prophylls not reddish brown striate or maculate (at 10× magnification); [section Tetragoni]
25 Floral scalesmucronate, mucro at least 0.2 mm long; spikes densely ovoid to loosely ellipsoid-oblong (to 1.5× as long as wide); rachis, spikelets, bracteoles, peduncles, and prophylls usually conspicuously reddish brown striate or maculate (at 10× magnification); [section Strigosi].
26 Spikes denser, spikelets 11-21 per 5 mm span of the rachis (at least in the upper portion), often the upper spikelets ascending; scale color bilaterally variable, reddish or reddish-brown to whitish; spikelets estipitate or stipe to 0.2 mm long; spikes ellipsoid to oblong