1 Leaves either 2-3-foliolate or
palmately 4-11-foliolate (all the
leaflets attached at a common point).
2 Leaves 2-foliolate; fruit a
capsule , opening by a
circumscissile lid
2 Leaves either 3-foliolate or
palmately or pedately 4-11-foliolate.
3 Inflorescence an
involucrate head subtended by
phyllaries , the heads solitary or many and variously arrayed in secondary
inflorescences , the
ovary inferior , the
corolla connate and
tubular at least basally, the
calyx absent, the
stamens 5, the fruit a
cypsela
3 Inflorescence , flower, and fruit structure various, but not with the combination of features as above.
4 Inflorescence various, usually not an
umbel (sometimes an
umbel in
Oxalis in OXALIDACEAE);
ovary superior ; fruit an aggregate,
legume ,
berry , or 2-valved
capsule .
5 Leaflets either
entire or barely and very shallowly
crenulate or notched at the tip (but otherwise
entire ).
6 Inflorescence a
spadix , surrounded by a
spathe ; fruit a
berry ; [
Monocots {illogically keyed here because of the likelihood of being mistaken for a
dicot }]
7 Flowers bilaterally symmetrical; fruit a
legume ; [plant of uplands]
7 Flowers radially symmetrical; fruit a 2-valved or 5-valved
capsule ; [plant of uplands or wetlands]
8 Leaflets not notched at the tip; flowers white; [plants of
saturated saturated or ponded wetlands]
8 Leaflets notched at the tip; flowers pink, white, or yellow; [plants of uplands or
temporarily flooded wetlands]
9 Petals 5 or more;
stamens 10 or more; fruit either a
legume or an aggregate of
achenes or
follicles
10 Stamens many, fused into a
staminal tube;
carpels 10-20, in a ring;
pubescence stellate (sometimes mixed with simple
hairs )
10 Stamens 10-many, separate, or fused but not all into a
staminal tube;
carpel either 1 (FABACEAE), or 3-7 in a ring (RANUNCULACEAE), or many and
spirally arranged on a
conical receptacle (RANUNCULACEAE or ROSACEAE)
11 Leaflets serrulate ; flowers bilaterally symmetrical; fruit a
legume ;
corolla variously colored, including white
11 Leaflets serrate ; flowers radially symmetrical; fruit an aggregate of
achenes or of
follicles ;
corolla white or yellowish or greenish.
12 Fruit an aggregate of
follicles
12 Fruit an aggregate of
achenes (borne on a fleshy, expanded
receptacle in
Fragaria and some
Potentilla )
13 Flowers lacking a
hypanthium
13 Flowers with a
hypanthium
1 Leaves 1-
pinnately compound (all
leaflets attached to a central
rachis ) or more complexly compound (with several orders of branching, some
leaflets at least attached to second-order branches from the
rachis ).
14 Leaves 1-
pinnately compound (all
leaflets attached to a central
rachis ).
15 Inflorescence an
involucrate head subtended by
phyllaries , the heads solitary or many and variously arrayed in secondary
inflorescences , the
ovary inferior , the
corolla connate and
tubular at least basally, the
calyx absent, the
stamens 5, the fruit a
cypsela
15 Inflorescence , flower, and fruit structure various, but not with the combination of features as above.
16 Flowers bilaterally symmetrical; fruit a
legume
16 Flowers radially symmetrical; fruit a
silique /
silicle , or a
schizocarp of
mericarps , or an
achene .
17 Petals 0 or 5 (if 0, the
sepals petaloid );
stamens 2, 4, 5, or many.
18 Stamens 2, 4, or many; fruit an
achene
14 Leaves more complexly compound (with 2 or more orders of branching, some
leaflets at least attached to second-order branches from the
rachis ).
19 Inflorescence an
involucrate head subtended by
phyllaries , the heads solitary or many and variously arrayed in secondary
inflorescences , the
ovary inferior , the
corolla connate and
tubular at least basally, the
calyx absent, the
stamens 5, the fruit a
cypsela
19 Inflorescence , flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head subtended by
bracts , but then with other features differing, such as
stamens 4, or green
calyx present, or
petals separate, or fruit a
schizocarp of
mericarps , etc.).
20 Leaf
segments or ultimate
lobes linear or
lanceolate , > 2× as long as wide, < 4 mm wide.
21 Inflorescence various, but not an
umbel ;
ovary superior ; fruit an aggregate of
follicles or an elongate
capsule .
22 Carpels 2, fused; fruit an elongate
capsule ; flowers bilaterally symmetrical
22 Carpels 5-10 or many, separate; fruit an aggregate; flower radially symmetrical
20 Leaf
segments or ultimate
lobes ovate or
elliptic , < 3× as long as wide, > 5 mm wide.
24 Fruit a 5-seeded
drupe
23 Inflorescence various, but not an
umbel ; fruit an aggregate of
follicles or
achenes , an elongate
capsule , or a
naked seed resembling a
drupe .
25 Leaflets with < 10 ultimate ‘points’ (
lobe or
tooth terminations), these rounded to broadly
acute , often large in comparison to the
leaflet and appearing as “sublobes”;
pistil 1 or 4-many.
26 Corolla bilaterally symmetrical; fruit an elongate
capsule ; [cultivated
exotic , rarely
persistent near gardens]
26 Corolla radially symmetrical; fruit an aggregate of
follicles or
achenes , or a
naked seed resembling a
drupe ; [native plants of moist to dry forests and rock outcrops].
27 Leaflets 5-8 cm long, obviously longer than broad;
pistil 1; fruit a
naked blue seed resembling a
drupe ; flowers mainly 3
-merous
27 Leaflets 1-6 cm long, about as long as broad if > 4 cm long;
pistils 4-many; fruit an aggregate of
follicles or
achenes ; flowers mainly 4-5
-merous
25 Leaflets with >11 ultimate ‘points’ (
lobe or
tooth terminations), these
acuminate to
acute ;
pistils 1-8.
28 Pubescence of the stem and lower leaf surface
glandular ; flowers
unisexual , on the same plant (
monoecious );
stamens 10;
pistils 2, partly fused; fruit an aggregate of
follicles
28 Pubescence of the stem and lower leaf surface non-
glandular (or absent); flowers either
bisexual (the plants
hermaphroditic ), or
unisexual and the male and female flowers on separate plants (the plants
dioecious );
stamens 15 or more;
pistils 1-8, separate; fruit an aggregate of
follicles , a
follicle , or a red or white
berry .
29 Flowers
unisexual (plants
dioecious );
carpels 3-4 per
pistillate flower;
inflorescence a
panicle of
racemes , with numerous branches; fruit an aggregate of
follicles
1 Leaves either 3-foliolate or
palmately 4-11-foliolate (all the
leaflets attached at a common point, or the
leaflets slightly
pedate in
Helleborus foetidus in RANUNCULACEAE).
2 Inflorescence an
involucrate head subtended by
phyllaries , the heads solitary or many and variously arrayed in secondary
inflorescences , the
ovary inferior , the
corolla connate and
tubular at least basally, the
calyx absent, the
stamens 5, the fruit a
cypsela
2 Inflorescence , flower, and fruit structure various, but not with the combination of features as above.
3 Leaflets obviously and sharply
serrate ;
pistils 1 or 5-many; fruit an
achene , or an aggregate of
achenes ,
drupelets , or
follicles .
4 Leaflets 7-11, slightly
pedate in their arrangement, evergreen
5 Pistil 1; fruit an
achene
5 Pistils 5-many; fruit an aggregate of
achenes ,
drupelets , or
follicles
3 Leaflets entire , finely
denticulate , or very obscurely
toothed (or irregularly
serrate or
lobed in
Cardamine in BRASSICACEAE);
pistil 1; fruit simple, a
legume ,
capsule ,
silique , or
berry .
6 Corolla bilaterally symmetrical;
petals 5; fruit a
legume ;
carpel 1
6 Corolla radially symmetrical;
petals 4 or 5; fruit either an elongate
capsule or a
berry ;
carpels 1, 2, or 5.
7 Leaflets radially arranged at the summit of the
petiole , not differentiated in size or placement into a terminal
leaflet and 2
lateral leaflets ;
leaflets prominently notched at the
apex ;
petals 5, yellow;
inflorescence axillary ,
cymose or
umbelliform ;
carpels 5
7 Leaflets differentiated in size and placement into a terminal
leaflet and 2 or more
lateral leaflets ;
leaflets not regularly notched at the
apex (a few may be slightly
retuse );
petals 4, white, pink, or yellow;
inflorescence terminal and
racemose ;
carpels 1 or 2.
8 Carpels 2; fruit a red
berry ;
petals connate , purplish-blue
8 Carpel 1; fruit a
capsule ;
petals separate, white, pink, or yellow.
9 Stem leaves 1-3,
alternate [or
whorled or opposite];
leaflets 3, irregularly
serrate ,
lacerate , or additionally divided or
lobed ; fruit a
silique ;
carpels 2
9 Stem leaves >3,
alternate ;
leaflets (1-) 3-7, each
entire or obscurely
toothed ; fruit a
capsule ;
carpel 1
1 Leaves either 1-
pinnately compound (all
leaflets attached to a central
rachis ) or more complexly compound (with several orders of branching, some
leaflets at least attached to second-order branches from the
rachis ).
10 Inflorescence an
involucrate head subtended by
phyllaries , heads solitary or many, variously arrayed in secondary
inflorescences ; fruit a
cypsela ;
ovary inferior
10 Inflorescence various, but not as above; fruit various, not as above;
ovary superior .
11 Leaves 1-
pinnately compound (all
leaflets attached to a central
rachis ).
12 Flowers bilaterally symmetrical,
papilionaceous ; fruit a
legume ; leaves even-
pinnately or odd-
pinnately compound , the terminal
leaflet sometimes replaced by a
tendril ;
leaflets entire or at most minutely
denticulate
12 Flowers radially symmetrical (or barely bilaterally symmetrical in
Erodium in GERANIACEAE); fruit a
capsule ,
capsular but of 5
mericarps , or an aggregate of
achenes ,
nutlets , or
follicles (in some cases the # of
pistils from many down to 2 or even 1); leaves odd-
pinnately compound , never with
tendrils ;
leaflets serrate (or
entire to
shallowly lobed in
Polemonium in POLEMONIACEAE,
Cardamine in BRASSICACEAE, and
Floerkea in LIMNANTHACEAE).
13 Pistils many (only 1-2 in
Agrimonia ,
Poteridium ,
Poterium , and
Sanguisorba ); fruit an aggregate of
achenes ,
nutlets , or
follicles ;
hypanthium present;
stamens 5-many (only 4 in
Poteridium and
Sanguisorba )
13 Pistil 1 (or deeply 2-3-
lobed in
Floerkea in LIMNANTHACEAE); fruit a
silique ,
capsule ,
schizocarp of 2-3
mericarps , or a
capsular schizocarp of 5
mericarps (
Erodium in GERANIACEAE);
hypanthium absent;
stamens 3-6.
15 Petals 5, fused (distinct in
Erodium in GERANIACEAE);
stamens 5;
inflorescence axillary or terminal,
cymose , consisting of subcapitate,
umbel -like, or
helicoid cymes ; fruit either a
capsule , or a
capsular schizocarp of 5
mericarps (
Erodium in GERANIACEAE).
16 Flowers slightly bilaterally symmetrical (2 of the
petals of different size than the other 3); fruit a
capsular schizocarp of 5
mericarps ;
carpels 5
16 Flowers radially symmetrical; fruit either a
loculicidal capsule or a
berry ;
carpels 2 or 3.
17 Fruit a
berry ; fresh plant rankly fragrant
17 Fruit a
capsule ; fresh plant not aromatic.
18 Capsule 2-
valvate ;
carpels 2;
leaflets prominently
serrate or with some
tooth -like sublobes
11 Leaves more complexly compound (with 2 or more orders of branching, some
leaflets at least attached to second-order branches from the
rachis ).
19 Leaves 2×
even-pinnate ; flowers in
spikes or
spherical heads; fruit a
legume or
loment .
19 Leaves either 2×
odd-pinnate or more complexly 2-4×
ternately or
ternately -
pinnately compound ; flowers in various
inflorescences ; fruits various (not
legumes or
loments ).
20 Leaf
segments linear , less than 2 mm wide.
21 Inflorescence either a terminal solitary flower or terminal
raceme or
panicle ;
ovary superior , either of 2 fused
carpels or of 1-5or many distinct 1-
carpellate pistils ; fruit either a
capsule or an aggregate of
follicles or
achenes .
22 Ovary of 2 fused
carpels ; fruit a
capsule (1-seeded and
indehiscent in
Fumaria )
22 Ovary of 1-5or many distinct 1-
carpellate pistils ; fruit an aggregate of
follicles or
achenes
20 Leaf
segments broader,
lanceolate ,
ovate , or
elliptic , > 5 mm wide.
23 Erect or
sprawling herb;
stamens 5-6 or >15.
24 Leaflets sharply
serrate , with usually many teeth on each
leaflet side, the total number of “points” per
leaflet > 10.
26 Flowers
unisexual ;
carpels (in
pistillate flowers) of 3 (-5)
carpels ; fruit an aggregate of
follicles
24 Leaflets entire , or with 1-several, broad,
obtuse , rounded, or broadly
acute “sublobes”, especially towards the tip of the
leaflet , the total number of “points” per
leaflet < 10.
28 Perianth bilaterally symmetrical, the
corolla of 4 fused
petals ; plant a
scandent vine or erect or
sprawling herb
28 Perianth radially symmetrical, of 1-5
whorls of separate
perianth parts; plant an erect herb.
29 Perianth of 4-5
whorls of 3 parts each (some of the
whorls modified into
nectaries );
pistil 1, 1-
carpellate ; fruit a drupelike, blue,
naked seed; largest
leaflets > 6 cm long, obviously longer than wide
29 Perianth of 1
whorl ; of 4-5 parts;
pistils many, each 1-
carpellate ; fruit an aggregate of
achenes or an aggregate of
follicles ; largest
leaflets either < 6 cm long, or if longer than 6 cm, also about as wide as long
1 Leaflets linear , < 1.5 mm wide.
2 Flowers bilaterally symmetrical; [subfamily
Ranunculoideae , tribe
Delphineae ]
2 Flowers radially symmetrical.
3 Aquatic; [native]; [subfamily
Ranunculoideae , tribe
Ranunculeae ]
4 Flower closely subtended by a finely
dissected involucre ;
pistils compound; [subfamily
Ranunculoideae , tribe
Nigelleae ]
4 Flower lacking
involucre ;
pistils simple.
5 Petals 8-15 mm long, red, purple, or orange; [subfamily
Ranunculoideae , tribe
Adonideae ]
5 Petals 3-5 mm long, yellow; [subfamily
Ranunculoideae , tribe
Ranunculeae ]
1 Leaflets broader, rounded,
lobed , or
toothed .
6 Leaves all
cauline , opposite; stems somewhat
woody at
base ; [subfamily
Ranunculoideae , tribe
Anemoneae ]
7 Petals present, conspicuous
8 Flowers dangling;
petals red, orange with yellow, or blue, spurred; [subfamily
Thalictroideae ]
8 Flowers not dangling;
petals yellow, not spurred; [subfamily
Ranunculoideae , tribe
Ranunculeae ]
7 Petals absent or inconspicuous (soon
deciduous or altered into a
nectary -bearing
clavate structure);
sepals sometimes
petaloid and conspicuous.
9 Sepals petaloid , conspicuous, white (or cream, rose, pink, or tinged with green).
10 Involucre absent, all leaves on the stem
alternate ;
petaloid sepals 5, white; [subfamily
Thalictroideae ]
10 Involucre of opposite or
whorled , leaflike
bracts present;
petaloid sepals (4-) 5-20 (-30), white, cream, rose, or green.
11 Basal leaves with 3-5
leaflets , these
toothed or
incised ;
petaloid sepals white, cream, rose, or green; [subfamily
Ranunculoideae , tribe
Anemoneae ]
11 Basal leaves with > 5
leaflets ; these with 0-3 rounded
lobes at the tip;
petaloid sepals white to pale pink; [subfamily
Thalictroideae ]
9 Sepals absent, or inconspicuous in comparison to the
stamens or
pistils .
12 Leaflets 3; flower solitary; [subfamily
Coptidoideae ]
12 Leaflets many; flowers many, in a
panicle or
raceme .
13 Inflorescence a
panicle ; [subfamily
Thalictroideae ]
1 Fruit a
follicle or
capsular (or fleshy and berrylike in some
Actaea ).
2 Mature leaves > 4 dm wide; [subfamily
Ranunculoideae , tribe
Cimicifugeae ]
2 Mature leaves < 3 dm wide.
4 Flowers in a
raceme , not subtended by an
involucre ; fruit
follicular , each with a 1-2 mm long
beak ; [subfamily
Ranunculoideae , tribe
Delphineae ]
4 Flower solitary, subtended by a finely divided
involucre ; fruit a
spherical capsule -like structure composed of 5 or 10 partially
connate follicles , each
follicle terminated by a
linear beak 13-20 mm long; [subfamily
Ranunculoideae , tribe
Nigelleae ]
3 Leaflets broad, rounded; [mostly natives].
5 Follicles borne on
stipes , forming an
umbel -like cluster;
rhizomes yellow or orange; [subfamily
Coptidoideae ]
5 Follicles sessile ;
rhizomes brown or tan; [subfamily
Thalictroideae ].
6 Follicles 15-31 mm long, with
beaks 7-18 mm long
6 Follicles 3.5-6.5 mm long, with
beaks 1.5-3 mm long
7 Leaves divided into numerous
linear segments , all of which are < 1 mm wide.
8 Plant aquatic (if leaves divided into numerous
linear segments ); [subfamily
Ranunculoideae , tribe
Ranunculeae ]
9 Achenes 3-5 mm long,
glabrous , with a
persistent beak 0.5-1 mm long; [subfamily
Ranunculoideae , tribe
Adonideae ]
9 Achenes 1.6-2 mm long,
tomentose , with a
persistent beak 3.5-4.5 mm long; [subfamily
Ranunculoideae , tribe
Ranunculeae ]
7 Leaf
segments rounded or
cleft , > 1 mm wide.
10 Leaves
cauline , opposite; [subfamily
Ranunculoideae , tribe
Anemoneae ]
10 Leaves
basal and/or
cauline ,
cauline leaves (if present)
alternate (leaflike
involucral bracts sometimes present and opposite or
whorled ).
11 Leaflike
involucral bracts present, opposite or
whorled .
12 Achenes not
ribbed or veined on
lateral surfaces; leaf texture moderate to distinctly thick and
leathery ; [subfamily
Ranunculoideae , tribe
Anemoneae ]
12 Achenes conspicuously
ribbed or veined on
lateral surfaces; leaf texture thin, delicate; [subfamily
Thalictroideae ]
11 Leaflike
involucral bracts not present.
13 Leaflets 3-many, if many the
leaflets typically with teeth, or sharp
lobes ; [subfamily
Ranunculoideae , tribe
Ranunculeae ]
13 Leaflets many,
unlobed or typically with 3-9 rounded
lobes ; [subfamily
Thalictroideae ]