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3 Leaves lanceolate or broader, 1-5× as long as wide; receptacle paleate; [collectively widespread]. | |
4 Leaves lanceolate to oblanceolate, unlobed or obscurely 3-lobed; leaf blade margins toothed or entire; leaves strictly opposite; [tribe Heliantheae; subtribe Ecliptinae]. | |
6 Leaves lanceolate, sometimes 3-lobed, usually serrate but sometimes nearly entire; heads 4-8 mm in diameter; cypselas flattened and winged; [TX only] | |
7 Disc flowers pink, purple, or white. | |
8 Heads small (involucres 2-8 mm high), solitary, axillary in the axils of leaves or leafy bracts (similar to the leaves but smaller) or interpretable as arrayed in bracteate racemes; heads nodding, the involucre 2-7 mm high; [collectively widespread in coastal portions of our area, of maritime situations]; [tribe Heliantheae; subtribe Ambrosiinae] | |
8 Heads small to larger (involucres 3-15+ mm high), many, terminal on the branches of corymbiform or paniculiform arrays; heads erect, the involucre 4-12 mm high; [of Coastal Plain of FL, s. GA, and TX]. | |
10 Leaf blades triangular or lanceolate, truncate, cordate, or cuneate at the base, acute to acuminate at the apex; leaf margins serrate; leaves opposite throughout. | |
11 Phyllaries broad, 2-3× as long as wide, the inner phyllaries translucent-scarious white to pinkish at their broadly rounded apices, falling as the head goes to fruit | |
11 Phyllaries narrow, 4-7× as long as wide, the inner phyllaries green at their acute to acuminate apices, persistent on the head in age. | |
12 Involucres 2-3 mm in diameter; phyllaries 7-16 in 1-2 series; disc flowers 3-13 per head; [s. FL only (in our region)] | |
12 Involucres 3-7 mm in diameter; phyllaries 25-35 in 2-3 series; disc flowers 10-60 per head; [TX only (in our region)]. | |
17 Rays white, 21-34+; leaves obovate to oblanceolate, averaging ca. 4× as long as wide, dentate; [non-native, known in our region only from NY] | |
18 Heads pseudoradiate, with 11-25 pseudoray flowers and 0 disc flowers; leaves 8-70 mm wide | |
19 Phyllaries in 1-2 series and equal or subequal in length to one another, the phyllaries often coherent/fused (the involucre thus appearing as a cylinder with ribs extending from base to top; calyculus (of bracts subtending the phyllaries and differing from them in texture, color, or orientation) present; [tribe Senecioneae] | |
19 Phyllaries in 3-17 series and unequal in length (imbricated); calyculus absent. | |
20 Rays bright yellow, (0-) 1-9; involucre 1-2.5 (-3) mm in diameter. | |
21 Pappus of 40-60 barbellate bristles; ray blades 4-6 mm long; leaf surfaces minutely pebbled; intricately branched shrubs to 1 m tall, with persistent sterile shoots with evergreen leaves and annual flowering shoots (the entire shoot and leaves deciduous); [Coastal Plain, se. NC south to Panhandle FL and s. MS] | |
24 Leaves 1-pinnately lobed, the many (>21) lobes mostly < 1.5 mm long and wide | |
25 Plants dioecious, either male or female; heads either staminate, with 10-50 flowers, or pistillate, with 20-150 flowers | |
27 Leaves lanceolate to ovate (broadest below the midpoint); leaf margins entire to coarsely toothed. | |
28 Heads in leafy, paniculate arrays; leaf blades coarsely dentate, 0.5-4 cm long; shrubs to 12 dm tall; [c. and e. TX] | |
28 Heads borne in leafless, corymbiform arrays; leaf blades entire to denticulate, 5-16 (-20) cm long; shrubs to 40 dm tall; [FL and s. TX] | |
29 Leaf blades elliptic, acute to acuminate at the apex; leaf blade strongly bicolored above and below, the upper surface dark green and glabrate, the lower surface densely and finely tomentose and strongly whitened, the upper surface dark green and glabrate or glabrous; disc flowers 4-25 per head; [s. TX] |
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1 Receptacle paleate (bearing receptacular bracts which individually subtend at least some of the flowers of the head). | |
2 Involucres absent or vestigial, usually with 0 phyllaries; heads very small, < 6 mm high, < 4 mm wide; annuals, < 4 dm tall; disc corollas hidden or dingy in color; [tribe Gnaphalieae]. | |
2 Involucres present, conspicuous, of many phyllaries; heads larger, 2-20+ mm wide; perennials or coarse annuals, 2-25 dm tall; disc corollas apparent, white, or pink to purplish or reddish. | |
5 Cypselae with 10 ribs; involucre 3.5-12 (-15) mm tall and in diameter; pappus of 35-40 barbellulate bristles; [mainly Coastal Plain; VA south to s. FL, west to e. LA] | |
6 Heads larger, the involucre 6-15 mm high, with 15-40 phyllaries; leaves with conspicuous (at least at 10× magnification) resin dots. | |
7 Stem with punctate glands; peduncles punctate-glandular and also hirsutulous; pappus bristles in 2 series; [of FL peninsula] | |
1 Receptacle epaleate (lacking receptacular bracts individually subtending flowers, but the receptacle sometimes bearing bristles, setae, hairs, or pits with laciniate margins; some taxa lacking paleae on many or most flowers of the head are keyed here as well, as a failsafe). | |
8 Phyllaries in 1-2 series and equal or subequal in length to one another, the phyllaries often coherent/fused (the involucre thus appearing as a cylinder with ribs extending from base to top, except in species in which the phyllaries are radially winged); calyculus (of bracts subtending the phyllaries and differing from them in texture, color, or orientation) present or not; [tribe Senecioneae] | |
10 Plants annual, upright or ascending, not purple-hairy. | |
9 Perennials (robust annuals in Erechtites), 2-30 dm tall; leaves basally disposed and alternate on the stem (except only alternate in Erechtites); disc flowers cream or yellow (rarely pale lavender in Arnoglossum). | |
12 Plants annual, with leaves primarily on the stem; leaves sharply and raggedly toothed or even pinnately lobed; heads disciform (the outer flowers female, the middle bisexual, and the inner functionally male) | |
13 Disc flowers yellow. | |
14 Annuals; leaves not basally disposed, evenly distributed on the stem, the lower stem leaves more-or-less pinnatifid | |
13 Disc flowers cream, white, or greenish white (to pale lavender in some Arnoglossum). | |
15 Phyllaries 7-21, not radially winged; disc flowers 10-55 (-80+); calyculus absent or of 1-9+ bracts. | |
8 Phyllaries in 3-17 series and unequal in length (imbricated); calyculus absent. | |
19 Plants perennial, mat-forming by stolons, with basal rosettes of spatulate leaves, these usually with obvious appressed hairiness on (at least) the lower and (sometimes also) the upper blade surfaces; erect stems (with scattered alternate leaves) 4-45 cm tall; plants usually dioecious (pistillate and staminate heads on separate plants) | |
19 Plants annual, biennial, or perennial, not mat-forming or stoloniferous, erect, the leaves usually lanceolate or linear (spatulate in some species), leaf surface hairiness variable; erect stems (with alternate leaves) 5-80 cm tall; plants hermaphroditic (with heads containing both pistillate and functionally staminate flowers) or dioecious or subdioecious (in Anaphalis). | |
20 Plants annuals (perennials in Omalotheca sylvatica); heads disciform (each bisexual); plants hermaphroditic. | |
21 Leaves oblanceolate, 1.5-4 mm wide, notched at the tip or with a minute apiculus in the notch; pappus bristles persistent, plumose | |
21 Leaves various in shape, the lager > 4 mm wide, acuminate, acute, obtuse or rounded at the tip; pappus readily falling, the bristles barbellate to barbellulate. | |
22 Heads in capitate clusters arrayed in corymbiform or paniculiform arrays; phyllaries white or silvery; pappus bristles distinct, falling individually | |
24 Perennial; phyllaries in 2-3 series; cypselas strigillose; involucre light to dark brown; [waif, in PA] | |
24 Annual; phyllaries in 3-7 series; cypselas papillate or glabrous; involucre white to red-tinged or brown; [widespread in our region] | |
26 Disc corollas yellow, pale yellow, orange, reddish, or brown. | |
27 Phyllaries 5, in one series; phyllaries streaked with elongate glands filled with strong-scented oil | |
29 Erect or ascending annual or perennial, 1-15 dm tall; heads in corymbiform, paniculiform, or racemiform arrays. | |
30 Plants dioecious, either male or female; heads either staminate, with 10-50 flowers, or pistillate, with 20-150 flowers; heads pale yellow to white | |
31 Leaves (at least the larger) toothed or lobed; flowers 5-35 per head; heads in racemiform, paniculiform, or corymbiform arrays, but if corymbiform, irregular. | |
33 Inflorescences in elongate, racemiform to paniculiform arrays; leaves moderately to densely hairy with both glandular and non-glandular hairs | |
26 Disc corollas pink, purple, lavender, blue, or white. | |
34 Phyllaries either spine-tipped, or appendaged with a marginal zone that is strikingly different in color and texture than the phyllary body (except Leuzea); receptacle epaleate, but densely beset with bristles or subulate scales (flattened bristles). | |
35 Phyllaries scarious-margined, neither spine-tipped nor appendaged with dentate or fringed appendages | |
35 Plyllaries either spine-tipped or appendaged. | |
34 Phyllaries not armed, also not appendaged with a marginal zone that is strikingly different in color and texture than the phyllary body. | |
39 Plants dioecious, either male or female; heads either staminate, with 10-50 flowers, or pistillate, with 20-150 flowers; heads pale yellow to white | |
40 Leaf blades linear, lanceolate, or ovate; leaf margin entire, serrulate, or serrate; [collectively common and widespread]. | |
45 All leaves entire (or at most with a few obscure crenations); phyllaries lacking elongate striations, the nerving not apparent. | |
46 Heads borne in spiciform or racemiform arrays (all heads directly attached to a central access of the capitulescence) | |
46 Heads borne in corymbiform, paniculiform, or thyrsiform arrays (at least some heads borne on secondary branches of the capitulescence. | |
47 Heads 3-20 mm tall; cypselas 10-ribbed. | |
48 Heads 3.5-6 mm tall; phyllaries 5-12 in 1-2 (-3) series; [Coastal Plain, NC to FL to e. LA]. | |
49 Stem with punctate glands; peduncles punctate-glandular and also hirsutulous; pappus bristles in 2 series; [of FL peninsula] | |
48 Heads 3-20 mm tall; phyllaries 15-50+ in 3-7 series; [collectively widespread]. | |
50 Heads borne in corymbiform arrays; involucres 6-12 (15) mm tall; receptacles paleate, at least towards the periphery of the receptacle; pappus of 35-40 barbellulate to barbellate bristles; [nearly restricted to the Coastal Plain] | |
50 Heads borne in paniculiform or thyrsiform arrays; involucres 3-20 mm tall; receptacles epaleate; pappus of 12-40 coarsely barbellate, subplumose, or plumose bristles; [widespread in our region] |