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1 Receptacles paleate (with chaff); [tribe Heliantheae]. |
2 Stems not winged; leaves simple; disc florets and ray florets differing in color |
2 Stems usually winged; leaves simple or pinnately to palmately lobed; disc florets concolorous (of similar color) with ray florets |
1 Receptacles epaleate (lacking chaff). |
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4 Plants arachnose or wooly pubescent throughout; [ waif, SC; tribe Arctotideae] |
4 Plants glabrous or densely viscid but not wooly tomentose throughout (if hirsuto- pilose, this usually concentrated distally). |
5 Receptacles flat or broadly conic, not conspicuously pitted; heads in corymbiform arrays; pappus a short crown; rays white, pink, or purple adaxially; [widespread non-natives, tribe Anthemideae] |
5 Receptacles conic, pitted; heads borne singly ( Aphanostephus) or in loose corymbiform arrays ( Egletes); rays white adaxially; [natives primarily w. of MS river; tribe Astereae]. |
6 Plants aromatic, but not densely viscid, instead sometimes hairy distally; ray laminae not shorter than involucre; phyllary margins broadly scarious; pappus coroniform or a mix of variously shaped scales; [widespread w. of MS river] |
6 Plants densely viscid with stipitate- glandular trichomes; ray laminae sometimes shorter than involucre; phyllary margins scarious or not; pappus of uneven rings; [TX] |
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7 Plants 1-3 cm tall, densely pin-cushioned ( pulvinate); flowering heads sessile, borne singly, conspicuously nestled within the rosette leaves and proportionately of similar size; pappus persistent, of 12-35+ setiform scales in 1 series; phyllaries in 4-6+ series |
7 Plants of various size, but not densely pin-cushioned; flowering heads borne on stems, if peduncles short the heads not conspicuously nestled within similar-sized leaf rosettes; pappus various, phyllaries in 2-6 series. |
8 Pappus of 5-40+ longer inner bristles plus shorter outer scales or setae (these sometimes falling); stems and leaves often (but not always) stipitate- glandular. |
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9 Inner bristles barbellate or thicker, fewer (ca. 5); stems and leaves glandular ( C. asteroides) or eglandular ( C. asteroides; C. belliloides); plants tap-rooted annuals |
8 Pappus primarily of awns, scales, crowns, or thickened rings (these minute, 0.1-0.8 mm long), sometimes also accompanied by shorter inner scales or bristles; stems and leaves eglandular (sometimes minutely glandular in Chaetopappa bellidifolia). |
10 Cypsela margins narrowly winged or winglike. |
11 Receptacles obscurely pitted; pappus of 2-3 sclerified awns plus often with 7-12 shorter bristles or scales; [collectively widespread] |
11 Receptacles smooth; pappus of 2 thin awns plus an elliptic ring of shorter awns; [s. TX] |
10 Cypsela margins not narrowly winged (4-angled in Aphanostephus but lacking marginal wings). |
12 Receptacles conic, pitted; cypselae 4-angled; pappus of setiform to awn-tipped scales or short ciliate crowns (0.1-0.2 mm long, minute, thus requiring magnification) |
12 Receptacles flat or slightly convex, smooth (not pitted); cypselae variously shaped but not conspicuously 4-angled; pappus of minute, thickened rings ( C. bellidifolia) or erose cuplike crowns ( C. imberbis) |
1 Receptacles chaffy (paleate). |
2 Phyllaries dry, scarious-margined. |
3 Ray florets 1-5 mm long; heads small in corymbiform arrays |
3 Ray florets > 5 mm long; heads large, terminating the branches |
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4 Ray florets < 2.5 mm long; disk florets sterile, with an undivided style |
4 Ray florets > 5 mm long; disk florets fertile, with a divided style. |
5 Ray florets pink or light purple, > 15 mm long; heads single |
5 Ray florets white, 5-10 mm long; heads 20-100 in a compound corymb |
1 Receptacles naked (epaleate). |
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7 Achene faces minutely glandular- glochidiate (bearing small gland-like bristles) |
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8 Leaves basal; cypselae marginally 2- ribbed; mature plants < 2 dm tall |
8 Leaves cauline (and sometimes also basal); cypselae with ± 10 ribs; mature plants > 2 dm tall |
9 Heads solitary or 2-3 in corymbiform arrays |
9 Heads 5-60+ in corymbiform arrays |
6 Pappus present, of bristles or scales. |
10 Heads primarily discoid with reduced ray florets |
10 Heads radiate with conspicuous ray florets. |
11 Taprooted annuals; ray florets 1-7 mm long. |
12 Leaves and stems not fleshy, rarely glabrous; cypselas < 1.4 mm long |
12 Leaves and stems fleshy, mostly glabrous; cypselas > 1.4 mm long |
11 Not taprooted and mostly perennials; ray florets > 3 mm long. |
13 Ray florets usually > 60; blooming Apr-Oct |
13 Ray florets usually < 60; blooming late May-Nov. |
14 Receptacles hemispheric to conic; pappus often with 2-4 awns ( Boltonia) or lacking awns and coroniform ( Astranthium). |
15 Achenes not narrowly winged, the surfaces minutely glandular- glochidiate (bearing small gland-like bristles) |
15 Achenes narrowly winged, the surfaces sometimes puberulent but not glandular- glochidiate |
14 Receptacles flat to slightly convex; pappus lacking awns. |
16 At least the basal and lower leaves both petiolate and cordate/subcordate at base. |
17 Often colonial; inflorescence corymbiform, flat-topped or rounded; outer phyllaries > 1 mm broad |
17 Not colonial; inflorescence paniculiform, often elongate; outer phyllaries < 1 mm broad |
16 Basal and lower leaves not both petiolate and cordate/subcordate at base. |
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19 Cypselas glandular; pappus double |
19 Cypselas not glandular; pappus single or double or in four series. |
20 Leaves silvery-silky on both sides (at least when young), entire |
20 Leaves not silvery-silky, entire or toothed. |
21 Pappus double, with inner bristles distinctly longer than outer bristles. |
22 Leaves not rigid, veiny, lanceolate to elliptic or ovate, > 6 mm wide |
22 Leaves rigid, 1-nerved, linear to linear- spatulate, < 5 mm wide |
21 Pappus simple with all bristles often about the same length. |
23 Ray florets white, few (usually 3-8); cypselas densely silky |
23 Ray florets white to pink or blue or purple, more numerous (usually 8-30); cypselas glabrous to pubescent but not densely silky. |
24 Ray florets white; involucres < 6 mm long; phyllaries < 1 mm wide |
24 Ray florets white or pink to blue or purple; involucres 7-12 mm long; phyllaries usually > 1 mm wide. |
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26 Phyllaries long- attenuate or loose and spreading |
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27 Plants coarse-hairy; lowest leaves > 5 cm wide |
27 Plants glabrous to hairy; lowest leaves < 3 cm wide |