Colors

Data mode

Account

Login
Sign up

Support FSUS...

We've finished our 2024 fundraiser. Many thanks to those who have given! It's not too late to support us (click here)...

No key was found for the requested taxon, but it is the only child of Asteraceae. Showing where it is keyed below.

X
Keyed in multiple places:

Click the number at the start of a key lead to highlight both that lead and its corresponding lead. Click again to show only the two highlighted leads. Click a third time to return to the full key with the selected leads still highlighted.

Key G4: shrubs and subshrubs with alternate, simple, unlobed, entire leaves

Copy permalink to share | Check for keys that lead to this key

1 Leaves evergreen. {add to 1a: Scaevola in GOODENIACEAE, Morella (inodora) in MYRICACEAE, Ternstroemia in PENTAPHYLACACEAE, Myrsine in PRIMULACEAE, Pyracantha in ROSACEAE, Dodonaea in SAPINDACEAE, Cestrum in SOLANACEAE, Thymelaea in THYMELAEACEAE, Conocarpus in COMBRETACEAE}
  2 Leaves 1-7 mm long, either acicular and spreading or ovate and appressed to the stems
  2 Leaves > 10 mm long.
    3 Leaves linear, > 15× as long as wide; [Monocots]
    3 Leaves broader, < 15× as long as wide; [Eudicots, Basal Angiosperms, or Monocots].
      4 Plant a creeping subshrub, < 1 dm tall
      4 Plant not creeping, > 3 dm tall.
        5 Inflorescence solitary (Illicium in ILLICIACEAE) or variously branched, spicate, racemose, or fascicled, not an involucrate head.
          6 Carpels separate; fruit an aggregate; fresh foliage strongly fragrant; [Basal Angiosperms]
             7 Fruit an aggregate of woody follicles arranged in a whorl
             7 Fruit an aggregate of red to blackish berries, 4-6 mm long, each on a long stipe, giving the aggregate almost the appearance of an umbel
          6 Carpels fused; fruit a berry, drupe, acorn (nut), capsule, or legume; fresh foliage not strongly fragrant; [Eudicots, Monocots, and Basal Angiosperms].
               8 Ovary with 3 carpels; fruit a berry; “leaves” actually cladodes; [Monocots]
               8 Ovary with 1, 2, 3, 4, or 5 carpels; fruit a berry, drupe, capsule, legume, or nut; leaves actually leaves; [Eudicots and Basal Angiosperms].
                 9 Leaves largely covered with silver and/or bronze lepidote scales and/or dense stellate hairs below (visible at 10× or higher magnification), giving the lower leaf surface a slightly shiny to almost metallic appearance. {add Lyonia ferruginea and L. fruticosa in ERICACEAE; add Loropetalum in HAMAMELIDACEAE}
                   10 Petals present, conspicuous, connate, white, the corolla rotate; fruit a berry with several seeds; fresh foliage with a strong, tar-like odor
                   10 Petals absent or inconspicuous, greenish and separate if present (note that the calyx is petaloid and white or yellowish in Elaeagnus of ELAEAGNACEAE); fruit a dry capsule with 3 seeds, or a drupe with a single seed; fresh foliage lacking a strong odor.
                     11 Perianth 4-merous; petals absent; petaloid sepals white to cream, fused and salverform; carpel 1; fruit a fleshy, red drupe, with a single seed
                     11 Perianth 5-merous; petals green and separate, or absent; sepals greenish, separate; carpels 3; fruit a 3-valved capsule with 3 seeds
                 9 Leaves with various vestiture (or glabrous), but not as above.
                       12 Flowers in spikes, these solitary opposite leaves or in axillary umbels
                       12 Flowers in other types of inflorescences, not spikes.
                          13 Leaves 1-foliolate on the upper stems, sometimes 3-foliolate below, or all reduced to phyllodial spines; flowers papilionaceous, bright yellow; fruit a legume; stems bright green
                          13 Leaves simple throughout; flowers either small, inconspicuous, tannish, borne in catkins (Quercus), or larger and urceolate, or with almost separate and spreading petals, white to pink or reddish-orange, in various terminal or axillary, branched inflorescences; fruit either a nut in a cupule (an acorn), or a (3-) 5-valved capsule, or a spherical berry or drupe; stems generally brown or tan (sometimes green).
                            14 Flowers small, inconspicuous, tannish, borne in catkins; fruit a nut in a cupule (an acorn)
                            14 Flowers white to pink or reddish-orange, either urceolate or tubular or with separate and spreading petals, in various terminal or axillary inflorescences, or solitary; fruit either a (3-) 5-valved capsule, or a spherical berry with 10+ seeds, or a 1-8 seeded dry or fleshy drupe.
                              15 Flowers white to pink or reddish-orange, rotate or urceolate (the petals united at least basally), in various terminal or axillary inflorescences or solitary; fruit either a 2-5 valved capsule or a spherical berry with 10+ seeds.
                                16 Flowers reddish-orange; corollas bearing post-staminal hairs (appearing as tufts of hair opposite each anther)
                                16 Flowers white to pink; corollas without obvious tufts of hair opposite each anther.
                                  17 Leaves 1 per node or also paired (on one side of the stem) at some nodes (the leaves then uneven in size); inflorescences leaf-opposed; fruit a berry.
                                  17 Leaves 1 per node; inflorescences terminal or axillary, never leaf-opposed; fruit a valved capsule.
                                    18 Seeds with fleshy aril or attachment; ovaries 2-4 carpellate; flowers rotate; leaves entire or very remotely serrulate.
                                       19 Flowers few, not showy, green to greenish-white, inflorescence not densely arranged (flowers also sometimes solitary); capsules not beaked; leaves not revolute or undulate
                                       19 Flowers numerous, showy, white; inflorescence densely arranged; capsules with short beak; leaf margins revolute or slightly undulate
                              15 Flowers white, petals spreading, separate even at the base, in axillary fascicles or racemes; fruit either a fleshy drupe with 4-8 pyrenes, or a dry single-seeded drupe.
1 Leaves deciduous. {add: Ditrysinia in EUPHORBIACEAE, Glochidion in PHYLLANTHACEAE, Phyllanthopsis in PHYLLANTHACEAE, Nierembergia in SOLANACEAE, Edgeworthia in THYMELAEACEAE, Ipomoea (I. carnea) in CONVOLVULACEAE; Swida (S. alternifolia) in CORNACEAE}
                                             22 Inflorescence branched, spicate, a catkin, or consisting of a solitary flower or axillary clusters or whorls, not an involucrate head.
image of plant
Show caption*© Michelle Wong, some rights reserved (CC BY), uploaded by Michelle W.
                                                      26 Fruit a globose, spinose capsule bearing a longitudinal ridge across 1 or both faces, thus essentially resembling a spikey ball; perianth (4-)5-merous, consisting of 3 upper connate petaloid claws and two lower sessile petals; fresh plants without a strange musky odor; [Eudicots]
                                                      26 Fruit an oblong berry, greenish-yellow when ripe, not bearing small spines; perianth 3-4-merous, without connate petaloid claws; fresh plants fragrant with a strange, musky odor; [Basal Angiosperms]
                                                    25 Inflorescence of 2 or more flowers; perianth 3-5-merous; fresh plants not musky-fragrant; fruits various, not as above.
                                                          28 Leaves elliptic or narrowly elliptic, broadest near the middle; fresh plants strongly fragrant with a citrus-like aroma; stems unarmed; fruit a drupe, with a single seed
                                                          28 Leaves obovate or oblanceolate, broadest near the apex; stems armed with nodal spines; fresh plants not fragrant; fruit a berry, with several seeds.
                                                        27 Flowers 4-5-merous; fruit fleshy or dry, black, blue, brown, tan, or red at maturity; ovary superior or inferior; [Eudicots].
                                                            29 Fruit either a drupe or berry (indehiscent, and variously fleshy or dry) or a dry 3-valved capsule with 1 seed; inflorescence axillary (solitary, clusters, fascicles, or racemes), or in a terminal raceme (Pyrularia in SANTALACEAE).
                                                              30 Leaves largely covered with silver and/or bronze shiny lepidote scales below, giving the lower leaf surface an almost metallic appearance
                                                                         35 Fruit a red or orange berry, 8-20 mm long; leaves usually on spur-shoots; [salty coastal areas, or exotics of disturbed situations]
                                                                           36 Inflorescence a narrowly cylindrical raceme, clustered several to many at the tip of the previous year’s wood and below the current season’s growth; fruit < 3 mm in diameter
                                                                                38 Fruit dark red to black, the pedicel < 10 mm long; leaf venation very neatly pinnate, with the secondary veins nearly straight and parallel to one another

Key G5: shrubs and subshrubs with alternate, simple, unlobed, toothed leaves

Copy permalink to share | Check for keys that lead to this key

1 Subshrubs or dwarf shrubs, aboveground stems creeping or erect, < 15 cm tall; leaves evergreen.
  2 Leaves 1.5-3 cm wide, coarsely toothed; flowers lacking sepals and petals; [exotic species, sparingly naturalized or spreading in suburban situations]
  2 Leaves < 1.5 cm wide, finely toothed or entire; flowers with sepals and petals; [native species, collectively widespread and common].
    3 Leaves fleshy, terete in ×-section; petals 5, bright pink
    3 Leaves flat, not fleshy; petals white or pale pink.
      4 Leaves < 2.5 mm wide; corolla with petals distinct; plant creeping
      4 Leaves > 5 mm wide; corolla with petals fused (distinct in Chimaphila); plant creeping or erect
1 Shrubs, aboveground stems erect, > 30 cm tall; leaves evergreen or deciduous.
        5 Inflorescence an involucrate (composite) head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela
        5 Inflorescence, flower, and fruit structure various, but not with the combination of features as above.
          6 Leaves evergreen. {add to 6a Ardisia in PRIMULACEAE, Rhaphiolepis in ROSACEAE, Camellia in THEACEAE}
             7 Leaves glandular-punctate on one or both surfaces with golden-yellow glands; flowers unisexual, lacking a perianth; fruit a pale gray, waxy drupe with a single seed
             7 Leaves not glandular punctate; flowers bisexual or unisexual, with a white, pink, or yellow perianth; fruit various, a red, blue, or black drupe or berry with several seeds, or a capsule.
               8 Petals connate and urceolate, white to pale pink; flowers bisexual; leaves ovate, lanceolate, or elliptic, broadest near the middle or towards the base, the teeth well-distributed along most of the margin on either side; fruit either a capsule or a red, blue, or black berry
               8 Petals distinct, yellow or white; flowers unisexual or bisexual; leaves oblanceolate or elliptic, broadest towards the tip or near the middle, the teeth usually concentrated in the upper half of the leaf; fruit either a black or red drupe with several pyrenes, a red berry with several seeds, or red or black pomes.
                 9 Plants with nodal, simple or tripartite thorns; flowers bisexual, with a yellow perianth; fruit a red berry with several seeds
                 9 Plants lacking thorns; flowers unisexual or bisexual, with a white perianth; fruit either a black or red drupe with several pyrenes or a red or black pome.
                   10 Flowers unisexual; fruit a black or red drupe with several pyrenes
          6 Leaves deciduous {add [Fagaceae]}.
                     11 Plants lacking thorns; leaf teeth acute, blunt, rounded, or callus-tipped, but not spinulose.
                       12 Leaves crenate-wavy, with 1-2 teeth per cm of leaf margin; leaves usually obliquely cordate or angled-truncate at the base; pubescence of leaves and stems stellate
                       12 Leaves crenulate, serrate or serrulate, with >2 teeth per cm of leaf margin; leaves cuneate, rounded, or subcordate at base, not oblique; pubescence of leaves and stems absent or simple.
                          13 Leaves prominently 3-veined from the base.
                            14 Ovary 5-locular; stamens many or 5, fused or separate; fruit a 5-valved capsule or of 5 mericarps; flowers yellow or pink, or white with a pink blaze
                            14 Ovary 3-locular; stamens 5, separate; fruit a 3-valved capsule or drupe; flowers white or pale green
                              15 Flowers in catkins; perianth absent or very small; fruit a 1-seeded nut, samara, or waxy drupe (capsule in Salix in SALICACEAE).
                                16 Leaves > 4 cm wide, lacking punctate glands; fruit a 1-seeded nut or samara
                                16 Leaves < 3 cm wide, either punctate-glandular on one or both surfaces or lacking punctate glands; fruit a 1-seeded waxy drupe or a capsule.
                              15 Flowers arrayed variously, but not in catkins; perianth present, conspicuous; fruit a 1-many-seeded capsule, pome, berry, or follicle.
                                    18 Ovary superior; fruit either dry and dehiscent, a capsule or an aggregate of follicles or achenes, or fleshy and indehiscent, a drupe with 4-8 pyrenes.
                                             22 Ovary 2-8-locular; fruit fleshy and indehiscent, a drupe with 2-8 pyrenes; flowers mostly functionally unisexual (or sometimes bisexual in RHAMNACEAE).
                                               23 Petals connate at the base; stamens alternate to the petals and opposite to the sepals; fruit 4-8-locular, with 4-8 pyrenes
                                               23 Petals separate (or absent in Rhamnus alnifolia); stamens opposite to the petals (when present) and alternate to the sepals; fruit 2-4-locular, with 2-4 pyrenes
                                                      26 Leaves > 5× as long as wide; stamens 2; ovary and capsule 3-locular; [plants of the Coastal Plain of SC, GA, AL, and FL]
                                                      26 Leaves < 3× as long as wide; stamens 5 or 10; ovary and capsule 2-3-locular; [plants collectively widespread].
                                                        27 Stamens 5; ovary and capsule 2-locular; leaves elliptic (widest near the middle), the teeth fine (usually > 5 points per cm of margin), and along much of the margin; inflorescence a terminal raceme; hairs of the lower leaf surface simple, erect
                                                        27 Stamens 10; ovary and capsule 3-locular; leaves obovate (widest towards the apex), the teeth obscure to coarse (usually < 4 points per cm of margin), and primarily in the upper half of the leaf; inflorescence a terminal or axillary raceme or cyme; hairs of the lower leaf surface either simple and appressed, or stellate.
                                                          28 Leaf margins regularly and evenly serrate in the upper half of the leaf (usually nearly entire towards the base); inflorescence an elongate, many flowered (>30) raceme borne at the end of branchlets of the season; corolla of separate petals, the stamens separate; hairs of the lower leaf surface simple and appressed
                                                          28 Leaf margins wavy or irregularly dentate, mainly in the upper half of the leaf; inflorescence a few flowered (<20) axillary raceme, cyme, or cluster; corolla fused basally into a tube, the stamens adnate to the tube; hairs of the lower leaf surface stellate

Key J4: shrubs and subshrubs with opposite simple leaves with entire margins

Copy permalink to share | Check for keys that lead to this key

1 Aerial and epiphytic, hemiparasitic shrub
1 Terrestrial, autotrophic or hemiparasitic shrub or subshrub.
  2 Leaves succulent, nearly as thick as wide; [brackish to saline situations]
  2 Leaves herbaceous (succulent in Borrichia), much wider than thick; [various habitats].
    3 Creeping or short subshrubs, the stems primarily prostrate, < 2 dm tall.
      4 Well-developed leaves 4-6 per stem; inflorescence a head subtended by 4 large white bracts
      4 Well-developed leaves many per stem; inflorescence of individual flowers axillary in pairs or clusters or in terminal cymes.
        5 Flowers yellow; leaves with pellucid or dark punctate glands (use at least 10× magnification)
        5 Flowers white, pale pink, or deep pink; leaves lacking sessile, punctate glands.
          6 Leaves linear; flowers pale to deep pink, 5-merous
          6 Leaves orbicular or elliptic; flowers white to pale pink, 4-merous or 5-merous
    3 Upright or scrambling shrubs, > 3 dm tall
               8 Scrambling shrubs, armed with recurved paired spines at the nodes
                 9 Inflorescence a terminal head of many flowers.
                   10 Head flattened, either subtended by 4 large white bracts or by an involucre with >5 green phyllaries.
                     11 Head subtended by 4 large white bracts; leaves with prominently parallel-arcing secondary veins; flowers 4-merous
                     11 Head subtended by an involucre of >5 green phyllaries; leaves with venation otherwise; flowers 5-merous
                 9 Inflorescence otherwise, either of a solitary flower, or one of a wide variety of inflorescences with flowers attached at different points along branched or unbranched axes (e.g. axillary). {add: [Lagerstroemia] LYTHRACEAE; [Rosmarinus] LAMIACEAE; [Laguncularia] COMBRETACEAE; [Buxus] BUXACEAE; [Exochorda] ROSACEAE; various other [see spreadsheet]}
                       12 Inflorescence flat-topped (broader than long), terminal, a compound cyme or corymb.
                          13 Flowers bright yellow; stamens many; leaves < 1.5 cm wide; fruit a capsule; leaves with pellucid or dark punctate glands (use at least 10× magnification)
                          13 Flowers white or creamy; stamens 4-5; leaves > 1.5 cm wide; fruit a drupe; leaves lacking sessile, punctate glands.
                       12 Inflorescence either terminal and not flat-topped (longer than wide), or axillary and variously shaped, or terminal and solitary, or leaf-opposed.
                              15 Carpels many (> 9), either separate or fused; stamens many; perianth segments either many and undifferentiated into calyx and corolla, maroon, brown, or yellow (in CALYCANTHACEAE), or differentiated into a fleshy and persistent calyx of 5-9 sepals, and a deciduous corolla of 5-9 red (or white) petals (Punica in LYTHRACEAE).
                                16 Fruit a wrinkled, 3-7 cm long, brown to black, elliptical aggregate of nearly spherical, large achenes; flowers solitary in axils; perianth segments many and undifferentiated into calyx and corolla, maroon, brown, or yellowish; ovary superior; branches unarmed
                                16 Fruit a leathery, 4-15 cm in diameter, reddish, spherical berry with obpyramidal seeds surrounded by a juicy sarcotesta (pomegranate); perianth differentiated, the sepals fleshy and persistent on the fruit, the petals deciduous, 5-9, bright red to white; ovary inferior; branches typically armed with axillary spines
                              15 Carpels 1-5, fused; stamens either 1-5 or 8-10; perianth segments 4-5 or 8, variously colored; fruit a simple capsule, drupe, or berry (including berry-like fruit); flowers 2-many, in axillary or terminal inflorescences (pistillate flowers sometimes solitary in SANTALACEAE and THESIACEAE); [Eudicots].
                                  17 Ovary inferior; corolla absent, radially symmetrical, or bilaterally symmetrical; fruit either a berry or a 1-seeded drupe.
                                    18 Flowers unisexual and plants dioecious; corolla absent; pistillate flowers solitary, staminate flowers in pedunculate umbels or cymes, either terminal or axillary; fruit a 1-seeded drupe; leaves acute to acuminate at the apex
                                    18 Flowers bisexual and plants hermaphroditic; corolla present; flowers paired, terminal or axillary, or in axillary spikes; fruit a berry; leaves rounded, obtuse, to acute (or acuminate in Lonicera maackii) at the apex
                                  17 Ovary superior; corolla radially symmetrical (absent in Forestiera in OLEACEAE); fruit either a 1-4-seeded drupe, or a many-seeded berry (or berry-like fruit), or a capsule.
                                         20 Leaves with a conspicuous mix of silvery stellate hairs (upper) and rusty colored scales (lower); ovaries bearing rusty colored scales; flowers small, yellowish and inconspicuous; [nw. PA northward]
                                         20 Leaves glabrous or nearly so, lacking a conspicuous mix of stellate hairs adaxially and rusty scales abaxially; ovaries not bearing rusty colored scales; flowers of various size and color.
                                           21 Stamens 8-10, of 2 different lengths in each flower; petals separate, 4-5 (-7), pink purple, 10-15 mm long; stems strongly arching, rooting at the tips; [plants of flooded to saturated wetlands]
                                           21 Stamens either (1-) 2 (-4), or 4-5, or 10, all of the same length; petals fused (separate in RHAMNACEAE and BUXACEAE, but then < 5 mm long and white or cream), white, bright-yellow, lilac, or pink; stems erect (or at least not arching and rooting at the tips); [plants of various habitats].
                                               23 Fruit a loculicidal capsule, dehiscing into 3 valves; branches square in ×-section; leaves < 2 cm long; [exotic, cultivated and weakly established, of temperate areas]
                                               23 Fruit a drupe with 2-4 pyrenes; branches round or nearly so in ×-section; leaves > 2 cm long; [natives, of peninsular FL]
                                             22 Petals fused, 4-5, white, bright yellow, lilac, or pink; stamens either (1-) 2 (-4) or 10; fruit either a capsule or a 1-seeded drupe.

Key N1: herbaceous dicots with mainly basal, compound leaves

Copy permalink to share | Check for keys that lead to this key

1 Leaves either 2-3-foliolate or palmately 4-11-foliolate (all the leaflets attached at a common point).
  2 Leaves 2-foliolate; fruit a capsule, opening by a circumscissile lid
  2 Leaves either 3-foliolate or palmately or pedately 4-11-foliolate.
    3 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela
    3 Inflorescence, flower, and fruit structure various, but not with the combination of features as above.
      4 Inflorescence various, usually not an umbel (sometimes an umbel in Oxalis in OXALIDACEAE); ovary superior; fruit an aggregate, legume, berry, or 2-valved capsule.
        5 Leaflets either entire or barely and very shallowly crenulate or notched at the tip (but otherwise entire).
          6 Inflorescence a spadix, surrounded by a spathe; fruit a berry; [Monocots {illogically keyed here because of the likelihood of being mistaken for a dicot}]
          6 Inflorescence a raceme or umbel, not surrounded by a spathe; fruit a capsule or legume; [Eudicots].
             7 Flowers bilaterally symmetrical; fruit a legume; [plant of uplands]
             7 Flowers radially symmetrical; fruit a 2-valved or 5-valved capsule; [plant of uplands or wetlands]
               8 Leaflets not notched at the tip; flowers white; [plants of saturated saturated or ponded wetlands]
               8 Leaflets notched at the tip; flowers pink, white, or yellow; [plants of uplands or temporarily flooded wetlands]
                 9 Petals 5 or more; stamens 10 or more; fruit either a legume or an aggregate of achenes or follicles
                   10 Stamens many, fused into a staminal tube; carpels 10-20, in a ring; pubescence stellate (sometimes mixed with simple hairs)
                   10 Stamens 10-many, separate, or fused but not all into a staminal tube; carpel either 1 (FABACEAE), or 3-7 in a ring (RANUNCULACEAE), or many and spirally arranged on a conical receptacle (RANUNCULACEAE or ROSACEAE)
                     11 Leaflets serrulate; flowers bilaterally symmetrical; fruit a legume; corolla variously colored, including white
                     11 Leaflets serrate; flowers radially symmetrical; fruit an aggregate of achenes or of follicles; corolla white or yellowish or greenish.
                       12 Fruit an aggregate of achenes (borne on a fleshy, expanded receptacle in Fragaria and some Potentilla)
1 Leaves 1-pinnately compound (all leaflets attached to a central rachis) or more complexly compound (with several orders of branching, some leaflets at least attached to second-order branches from the rachis).
                              15 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela
                              15 Inflorescence, flower, and fruit structure various, but not with the combination of features as above.
                                16 Flowers radially symmetrical; fruit a silique/silicle, or a schizocarp of mericarps, or an achene.
                                  17 Petals 0 or 5 (if 0, the sepals petaloid); stamens 2, 4, 5, or many.
                            14 Leaves more complexly compound (with 2 or more orders of branching, some leaflets at least attached to second-order branches from the rachis).
                                       19 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela
                                       19 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head subtended by bracts, but then with other features differing, such as stamens 4, or green calyx present, or petals separate, or fruit a schizocarp of mericarps, etc.).
                                         20 Leaf segments or ultimate lobes linear or lanceolate, > 2× as long as wide, < 4 mm wide.
                                             22 Carpels 2, fused; fruit an elongate capsule; flowers bilaterally symmetrical
                                             22 Carpels 5-10 or many, separate; fruit an aggregate; flower radially symmetrical
                                         20 Leaf segments or ultimate lobes ovate or elliptic, < 3× as long as wide, > 5 mm wide.
                                               23 Inflorescence various, but not an umbel; fruit an aggregate of follicles or achenes, an elongate capsule, or a naked seed resembling a drupe.
                                                    25 Leaflets with < 10 ultimate ‘points’ (lobe or tooth terminations), these rounded to broadly acute, often large in comparison to the leaflet and appearing as “sublobes”; pistil 1 or 4-many.
                                                      26 Corolla radially symmetrical; fruit an aggregate of follicles or achenes, or a naked seed resembling a drupe; [native plants of moist to dry forests and rock outcrops].
                                                        27 Leaflets 5-8 cm long, obviously longer than broad; pistil 1; fruit a naked blue seed resembling a drupe; flowers mainly 3-merous
                                                        27 Leaflets 1-6 cm long, about as long as broad if > 4 cm long; pistils 4-many; fruit an aggregate of follicles or achenes; flowers mainly 4-5-merous
                                                    25 Leaflets with >11 ultimate ‘points’ (lobe or tooth terminations), these acuminate to acute; pistils 1-8.
                                                          28 Pubescence of the stem and lower leaf surface glandular; flowers unisexual, on the same plant (monoecious); stamens 10; pistils 2, partly fused; fruit an aggregate of follicles
                                                          28 Pubescence of the stem and lower leaf surface non-glandular (or absent); flowers either bisexual (the plants hermaphroditic), or unisexual and the male and female flowers on separate plants (the plants dioecious); stamens 15 or more; pistils 1-8, separate; fruit an aggregate of follicles, a follicle, or a red or white berry.
                                                            29 Flowers bisexual (plants hermaphroditic); carpels 1-8 per flower; inflorescence a raceme, or a panicle of racemes with just a few branches; fruit an aggregate of follicles, a follicle, or a red or white berry
                                                            29 Flowers unisexual (plants dioecious); carpels 3-4 per pistillate flower; inflorescence a panicle of racemes, with numerous branches; fruit an aggregate of follicles

Key N2: herbaceous dicots with mainly basal, simple leaves

Copy permalink to share | Check for keys that lead to this key

1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head subtended by bracts, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or petals separate, or fruit a schizocarp of mericarps, etc.).
  2 Basal leaves 2-lobed, pinnately lobed, or palmately lobed (not considering cordate, hastate, or auriculate leaf bases as “lobed”).
    3 Basal leaves 2-lobed, hinged between the lobes, each lobe with stiff, marginal, eyelash-like bristles; [Coastal Plain of NC and SC, rarely planted and weakly naturalized elsewhere]
    3 Basal leaves 3-many-lobed, palmately or pinnately; [collectively widespread].
      4 Leaf lobing pinnate.
        5 Gynoecium of separate pistils (each with a single carpel); fruit an aggregate
        5 Gynoecium of a single pistil (with 2, rarely more, carpels); fruit simple.
          6 Stamens many; sepals 2, petals 4; fresh plants with yellow, orange, or white milky juice
          6 Stamens 4, 5, or 6; sepals 4 or 5; petals 4 or 5.
             7 Petals 4, distinct; stamens 6
             7 Petals 5, fused; stamens 2, 4, or 5.
               8 Corolla radially symmetrical; stamens 5
               8 Corolla 2-lipped, bilaterally symmetrical or asymmetrical; stamens 2 or 4.
                 9 Corolla lobes not twisted, the flower bilaterally symmetrical; stamens 2
                 9 Corolla lobes twisted so as to make the flower asymmetrical; stamens 4
      4 Leaf lobing palmate.
                          13 Petals 5; stamens 5; fruit a schizocarp of 2 mericarps.
                            14 Fruit smooth; leaves with 5 or more lobes
                              15 Gynoecium of separate pistils (each with a single carpel); fruit an aggregate.
                                  17 Carpels partly fused, arrayed in a ring of 10-20
                                16 Perianth of a single whorl of 3-12 petaloid sepals (the petals absent or small and rudimentary).
                                    18 Leaves 2, the single flower terminal and associated with the upper leaf; fruit an aggregate of berries
                                    18 Leaves normally > 2, flowers not as above; fruit an aggregate of achenes, utricles, or follicles
                                         20 Petals distinct; leaves with rounded lobes or teeth.
                                           21 Corolla radially symmetrical; petals 8-12; capsule fusiform, narrowed to both ends, > 5× as long as wide
  2 Basal leaves not lobed, at most serrate or crenate (and sometimes also cordate, hastate, auriculate, or peltate at the base).
                                                 24 Inflorescence either a terminal spike, or a 1-7-flowered terminal cyme, or of a solitary axillary or terminal flower; fruit various; perianth biseriate (of differentiated sepals and petals (except uniseriate, of 3 fused sepals in ARISTOLOCHIACEAE).
                                                      26 Flowers bilaterally symmetrical; inflorescence a terminal spike (with > 20 flowers); petals 4, usually scarious, transparent; sepals 4, green; stamens 4
                                                      26 Flowers radially symmetrical; inflorescence either of a solitary flower or of a 1-7-flowered terminal cyme; petals 5, 8-12, or 0; sepals 5 (green), 3 (brown), or 5-9 (yellow); stamens 5, 12, or many.
                                                        27 Gynoecium of separate pistils (each with a single carpel); fruit an aggregate of achenes or follicles; flowers bright yellow, either of 5-9 distinct petaloid sepals, or of 8-12 distinct petals subtended by 3-4 green distinct sepals
                                                        27 Gynoecium either of a single pistil with 6 carpels or of a single pistil with 4 carpels or of 2 nearly separate carpels; fruit a simple capsule (or deeply 2-lobed); flowers white, brown, or greenish, either of 5 fused or distinct white petals and 5 fused or distinct green sepals, or of 3 fused brown or greenish petaloid sepals.
                                                          28 Flowers brown or green, of 3 fused brown or greenish petaloid sepals (and 0 petals); carpels 6; stamens 12; leaves 4-10 cm wide
                                                          28 Flowers white, of 5 white or whitish petals and 5 green sepals; carpels 2; stamens 5; leaves 1-12 (-15) cm wide
                                                                 31 Gynoecium of separate pistils (each with a single carpel); fruit an aggregate; perianth of 5 green sepals and 5 colored petals.
                                                                           36 Inflorescence a terminal raceme; perianth of 4 green sepals and 4 white petals; fruit a silique/silicle; fresh foliage in spring and summer with a strong garlic odor; larger leaves < 10 cm in diameter
                                                                           36 Inflorescence a terminal panicle; perianth of 6 cream-colored petaloid sepals; fruit a winged achene; fresh foliage lacking a garlic odor; larger leaves typically > 30 cm in diameter
                                                                                  39 Leaves tubular, with a sutured ventral flange, erect or reclining, adapted as a pitfall for insects (flat, phyllodial leaves sometimes present as well, common in the winter in some species, such as S. oreophila)
                                                                                    40 Stem leaves opposite; perianth 5-merous, at least the corolla bilaterally symmetrical (barely so in VALERIANACEAE), or the parts curved so as to be asymmetrical (Pedicularis in OROBANCHACEAE); stamens 2, 3, or 4.
                                                                                        42 Corolla distinctly 2-lipped (with prominently large upper and lower corolla lobes) or hooded (the upper lip hood-like), distinctly bilaterally symmetrical, or the lobes twisted so as to make the corolla asymmetrical.
                                                                                    40 Stem leaves alternate; perianth radially symmetrical (less commonly bilaterally symmetrical); stamens 5, 6-8, 9, 10 (rarely 4).

Key O: herbaceous dicots with alternate, compound leaves on the stem

Copy permalink to share | Check for keys that lead to this key

1 Leaves either 3-foliolate or palmately 4-11-foliolate (all the leaflets attached at a common point, or the leaflets slightly pedate in Helleborus foetidus in RANUNCULACEAE).
  2 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela
  2 Inflorescence, flower, and fruit structure various, but not with the combination of features as above. {add under 2b [Cynosciadium] APIACEAE}
    3 Leaflets obviously and sharply serrate; pistils 1 or 5-many; fruit an achene, or an aggregate of achenes, drupelets, or follicles.
      4 Leaflets 7-11, slightly pedate in their arrangement, evergreen
        5 Pistils 5-many; fruit an aggregate of achenes, drupelets, or follicles
    3 Leaflets entire, finely denticulate, or very obscurely toothed (or irregularly serrate or lobed in Cardamine in BRASSICACEAE); pistil 1; fruit simple, a legume, capsule, silique, or berry.
          6 Corolla bilaterally symmetrical; petals 5; fruit a legume; carpel 1
          6 Corolla radially symmetrical; petals 4 or 5; fruit either an elongate capsule or a berry; carpels 1, 2, or 5.
             7 Leaflets radially arranged at the summit of the petiole, not differentiated in size or placement into a terminal leaflet and 2 lateral leaflets; leaflets prominently notched at the apex; petals 5, yellow; inflorescence axillary, cymose or umbelliform; carpels 5
             7 Leaflets differentiated in size and placement into a terminal leaflet and 2 or more lateral leaflets; leaflets not regularly notched at the apex (a few may be slightly retuse); petals 4, white, pink, or yellow; inflorescence terminal and racemose; carpels 1 or 2.
               8 Carpels 2; fruit a red berry; petals connate, purplish-blue
               8 Carpel 1; fruit a capsule; petals separate, white, pink, or yellow.
                 9 Stem leaves 1-3, alternate [or whorled or opposite]; leaflets 3, irregularly serrate, lacerate, or additionally divided or lobed; fruit a silique; carpels 2
                 9 Stem leaves >3, alternate; leaflets (1-) 3-7, each entire or obscurely toothed; fruit a capsule; carpel 1
1 Leaves either 1-pinnately compound (all leaflets attached to a central rachis) or more complexly compound (with several orders of branching, some leaflets at least attached to second-order branches from the rachis).
                   10 Inflorescence an involucrate head subtended by phyllaries, heads solitary or many, variously arrayed in secondary inflorescences; fruit a cypsela; ovary inferior
                   10 Inflorescence various, but not as above; fruit various, not as above; ovary superior.
                     11 Leaves 1-pinnately compound (all leaflets attached to a central rachis).
                       12 Flowers bilaterally symmetrical, papilionaceous; fruit a legume; leaves even-pinnately or odd-pinnately compound, the terminal leaflet sometimes replaced by a tendril; leaflets entire or at most minutely denticulate
                       12 Flowers radially symmetrical (or barely bilaterally symmetrical in Erodium in GERANIACEAE); fruit a capsule, capsular but of 5 mericarps, or an aggregate of achenes, nutlets, or follicles (in some cases the # of pistils from many down to 2 or even 1); leaves odd-pinnately compound, never with tendrils; leaflets serrate (or entire to shallowly lobed in Polemonium in POLEMONIACEAE, Cardamine in BRASSICACEAE, and Floerkea in LIMNANTHACEAE).
                          13 Pistils many (only 1-2 in Agrimonia, Poteridium, Poterium, and Sanguisorba); fruit an aggregate of achenes, nutlets, or follicles; hypanthium present; stamens 5-many (only 4 in Poteridium and Sanguisorba)
                          13 Pistil 1 (or deeply 2-3-lobed in Floerkea in LIMNANTHACEAE); fruit a silique, capsule, schizocarp of 2-3 mericarps, or a capsular schizocarp of 5 mericarps (Erodium in GERANIACEAE); hypanthium absent; stamens 3-6.
                              15 Petals 5, fused (distinct in Erodium in GERANIACEAE); stamens 5; inflorescence axillary or terminal, cymose, consisting of subcapitate, umbel-like, or helicoid cymes; fruit either a capsule, or a capsular schizocarp of 5 mericarps (Erodium in GERANIACEAE).
                                16 Flowers slightly bilaterally symmetrical (2 of the petals of different size than the other 3); fruit a capsular schizocarp of 5 mericarps; carpels 5
                                16 Flowers radially symmetrical; fruit either a loculicidal capsule or a berry; carpels 2 or 3.
                     11 Leaves more complexly compound (with 2 or more orders of branching, some leaflets at least attached to second-order branches from the rachis).
                                       19 Leaves either 2× odd-pinnate or more complexly 2-4× ternately or ternately-pinnately compound; flowers in various inflorescences; fruits various (not legumes or loments).
                                           21 Inflorescence either a terminal solitary flower or terminal raceme or panicle; ovary superior, either of 2 fused carpels or of 1-5or many distinct 1-carpellate pistils; fruit either a capsule or an aggregate of follicles or achenes.
                                             22 Ovary of 1-5or many distinct 1-carpellate pistils; fruit an aggregate of follicles or achenes
                                                 24 Leaflets sharply serrate, with usually many teeth on each leaflet side, the total number of “points” per leaflet > 10.
                                                 24 Leaflets entire, or with 1-several, broad, obtuse, rounded, or broadly acute “sublobes”, especially towards the tip of the leaflet, the total number of “points” per leaflet < 10.
                                                            29 Perianth of 4-5 whorls of 3 parts each (some of the whorls modified into nectaries); pistil 1, 1-carpellate; fruit a drupelike, blue, naked seed; largest leaflets > 6 cm long, obviously longer than wide
                                                            29 Perianth of 1 whorl; of 4-5 parts; pistils many, each 1-carpellate; fruit an aggregate of achenes or an aggregate of follicles; largest leaflets either < 6 cm long, or if longer than 6 cm, also about as wide as long

Key P1: herbaceous dicots with alternate, simple, and unlobed leaves on the stem

Copy permalink to share | Check for keys that lead to this key

1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.).
  2 Perianth uniseriate (represented only by undifferentiated tepals or sepals) or completely absent; flowers usually unisexual, less commonly bisexual).
    3 Inflorescence a cyathium, consisting of a single pistillate flower (reduced to a single 3-carpellate pistil) and 2 or more staminate flowers (each reduced to 1 stamen), borne in a cup-like involucre, the involucre bearing pointed or rounded glands, these sometimes brightly colored and petaloid, mimicking an individual flower (the cyathia then secondarily arranged in terminal cymes, or solitary and axillary, etc.); fresh plants with milky juice; fruit a 3-lobed, 3-locular capsule
    3 Inflorescence not a cyathium (and staminate or bisexual flowers with > 1 stamen; fresh plants lacking milky juice (except Stillingia in EUPHORBIACEAE); fruit various, not as above.
      4 Leaf margins toothed in some manner (serrate, dentate, crenate, etc.)
        5 Leaf teeth rounded to subacute, resembling shallow lobes, irregular, few (mostly < 6 per leaf side).
          6 Fruit a single-seeded achene or utricle; [plants of various disturbed or saline, usually sunny habitats]
          6 Fruit a 3-lobed, circumscissilely dehiscent capsule; [plants native of rich moist shaded forests or exotics in suburban woodlands]
        5 Leaf teeth sharp to crenate, not lobe-like, regular, many (mostly > 10 per leaf side).
             7 Leaf bases cordate to rounded.
               8 Styles 3; fruit a 3-lobed, 3-carpellate capsule (1 carpel sometimes aborting); inflorescence either a terminal or leaf opposed raceme, or a dense axillary condensed cyme with conspicuous toothed bracts subtending the flowers
               8 Styles 1 or 2; fruit either an achene or a multiple of achenes; inflorescence either an axillary dense cyme (almost a head), or an axillary spike with glomerules, or a terminal or axillary panicle.
                 9 Styles 2; inflorescence a dense axillary cyme (almost a head); fruit a multiple of achenes; plant lacking stinging hairs; [exotic plant of weedy situations]
                 9 Style 1; inflorescence an axillary spikes with glomerules, or a terminal or axillary panicle; plant either with stinging hairs or not; [plant a rare exotic (Boehmeria nivea) or a native of moist forests (Boehmeria cylindrica, Laportea)]
      4 Leaf margins entire.
                     11 Leaf base cordate; calyx 3-lobed, fused into a bilaterally symmetrical, curved brown or yellowish tube; fruit a capsule
                     11 Leaf base cuneate, rounded, or truncate; calyx of 3-4-5 distinct sepals, radially symmetrical, white or yellow; fruit a dry, nutlike drupe or an achene.
                       12 Leaves subsessile or very short petiolate, elliptic or narrowly elliptic, broadest near the middle; [native]
                       12 Leaves distinctly petiolate, rhombic, widest near the base; [rarely naturalized exotics].
                            14 Inflorescence a leaf-opposed (sometimes apparently terminal) spike or raceme; flowers visually white from white petaloid sepals, white bracts, or white stamens.
                              15 Sepals present, 4 or 5; petaloid, white; carpels 1 to many (-12); stamens 4 to many (-25); fruit a berry or an apically 2-lobed achene (as in Petiveria); leaf bases cuneate or rounded (but not cordate); [Eudicots].
                              15 Sepals absent; carpels 3-4; stamens 2-6 (-8); fruit a capsule, a 1-seeded drupe, or a schizocarp of 3-4 mericarps; leaf bases cordate or subcordate; [Basal Angiosperms].
                                  17 Fruit a capsule or schizocarp with 3-4 mericarps; stamens 3 or 6-8; plants terrestrial
                            14 Inflorescence not leaf-opposed, either simpler (single axillary or glomerules of flowers) or more complexly branched (axillary or terminal panicles or complex cymes); flowers white, reddish, scarious, or greenish.
                                    18 Stipules tubular, sheathing (= ocreae); flowers subtended by tubular, sheathing bracteoles (= ocreolae); nodes usually prominently swollen; perianth usually of 5-6 white to pink tepals
                                    18 Stipules not tubular or sheathing; flowers not subtended by ocreolae; nodes not swollen; perianth absent or of 3-5 sepals.
  2 Perianth biseriate (represented by differentiated whorls of sepals and petals, the sepals usually green or drab in color, the petals often brightly colored); flowers nearly always bisexual (there are exceptions).
                                                          28 Petals 4-7; stamens 1× or 2× as many as the petals, 4-7, 8, 10, 12, or14; leaves herbaceous in texture
                                                          28 Petals 5 (or sometimes doubled in horticultural forms); stamens 6-40 (or more); leaves fleshy in texture
                                                            29 Corolla bilaterally symmetrical, petals connate (except distinct in VIOLACEAE); fruit a capsule or legume (except a 1-seeded indehiscent spinose pod in Krameria in KRAMERIACEAE).
                                                              30 Petals connate (at least basally), 4, 5, 6, 7, or 8; carpels 1, 2, 4, 5, or 6 (rarely 3 in Reseda in RESEDACEAE); fruit a legume or 1-, 2-, or 5-loculed capsule (except a 1-seeded indehiscent pod in Krameria in KRAMERIACEAE).
                                                                 31 Stamens 6-10 (-25), more than the number (4 or 5) of petals and the number (4 or 5) of the sepals; fruit a legume or a 1-6-carpellate capsule.
                                                                         35 Pistil 2-carpellate; capsule 2 locular, opening gradually or not at all; inflorescence a terminal spike, raceme or panicle (or solitary, axillary flowers in Chaenorrhinum in PLANTAGINACEAE and Krameria in KRAMERIACEAE).
                                                                              37 Stamens 4; corolla with a distinct spur or sac at the base between the the 2 lower calyx lobes (except not spurred in Digitalis and Schwalbea); capsule loculicidal (only at the summit in Antirrhinum and Chaenorrhinum, and septicidal in Schwalbea); pubescence of the stem and leaves neither gland-tipped (except in Antirrhinum and Chaenorrhinum) nor dendritically branched.
                                                                                    40 Pistils 4-10 (each 1-carpellate) in a ring, these sometimes fused basally, each with its own style/stigma; fruit either an aggregate of achenes or follicles or a 5 (-7) locular capsule.
                                                                                        42 Fruit an aggregate of follicles; leaves fleshy in texture; inflorescence; leaves entire of sparsely and coarsely serrate, with < 12 points per leaf; [plants primarily of dry habitats]
                                                                                    40 Pistil 1, with 1-to many carpels (in many MALVACEAE, the carpels loosely united in a ring [of more than 5] around the single style/stigma); fruit either a 1-, 2-, 3-, 5-, 6-, or 10-locular capsule, or a silique/silicle, or a ring of mericarps.
                                                                                             44 Petals 5 (rarely 4 or 6); sepals 5 (rarely 4 or 6); stamens 5 (or multiples of 5), 6, or 12; fruit a capsule or a ring of mericarps.
                                                                                                                             59 Corolla lobes very short, much shorter than the corolla cup or tube, sometimes barely perceptible and represented only by teeth on the edge of the corolla limb, white or pink; style 1; herbage often with stipitate glands; fresh plants often rankly aromatic
                                                                                                                                      63 Corolla lobes very short, much shorter than the corolla cup or tube, sometimes barely perceptible and represented only by teeth on the edge of the corolla limb, white, yellow, pink, various other colors (rarely including blue).

Key P2: herbaceous dicots with alternate, simple, and palmately lobed leaves on the stem

Copy permalink to share | Check for keys that lead to this key

1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.).
  2 Plant a vine, climbing by tendrils or twining.
      4 Leaf margins entire; flowers bisexual; plants hermaphroditic; petals connate, large and showy
      4 Leaf margins serrate; flowers unisexual; plants dioecious; petals absent
    3 Vine climbing by tendrils.
        5 Ovary superior; petals distinct; flowers bisexual
  2 Plant an herb, sometimes sprawling, reclining (e.g. Cymbalaria in PLANTAGINACEAE, Aconitum in RANUNCULACEAE), but lacking climbing adaptations such as tendrils or twining stems.
             7 Involucre well-developed and obvious
          6 Ovary superior; inflorescence various, not an umbel; fruit various, a capsule, an aggregate of achenes or follicles, a ring of (>2) mericarps.
               8 Perianth uniseriate, the corolla absent (the calyx petaloid and white in Cnidoscolus); flowers unisexual; plants either with stinging hairs or not
               8 Perianth biseriate (uniseriate in Aphanes in ROSACEAE and in Trautvetteria in RANUNCULACEAE); flowers bisexual; plants lacking stinging hairs.
                 9 Pistils many (or 2-3 in Aphanes in ROSACEAE), each with 1 carpel, arranged spirally or in a ring (if in a ring, of 2-5); fruit an aggregate of achenes, follicles, or utricles.
                   10 Perianth bilaterally symmetrical, either hooded or spurred; fruit an aggregate of follicles
                   10 Perianth radially symmetrical, not hooded or spurred; fruit an aggregate of utricles or achenes (plumose achenes in Geum)
                     11 Stamens showy, bright white, dilated towards the tip; pistils ca. 15; fruit an aggregate of utricles
                     11 Stamens not showy, white, or dilated towards the tip; pistils many (> 25); fruit an aggregate of achenes.
                       12 Flowers with a prominent hypanthium; achenes with an elongate, plumose beak
                 9 Pistil 1, with 1-to many carpels (in many MALVACEAE, the carpels loosely united in a ring of more than 5 around the style); fruit a capsule, an achene, a follicle, or a ring of 3 or 5-many 1-seeded mericarps.
                            14 Corolla bilaterally symmetrical, the petals connate (except distinct in Delphinium in RANUNCULACEAE); fruit a capsule, a follicle, or a schizocarp of 3 1-seeded mericarps.
                              15 Corolla not spurred; fruit an elongate (10-20 cm) capsule with 2 curved beaks
                            14 Corolla radially symmetrical, the petals distinct (fused and tubular in Ipomoea); fruit a capsule or a schizocarp consisting of a ring of 5-many 1-seeded mericarps.
                                       19 Stamens many, connate into a stamen tube; carpels 5-many, completely or only loosely fused; fruit a capsule or a schizocarp of 5-many mericarps borne in a ring; calyx often subtended by an epicalyx (an additional calyx-like, green, foliaceous whorl of bracts)
                                       19 Stamens 5 or 10, distinct; carpels 2 or 5, fused; fruit a capsule or a schizocarp of 5 1-seeded mericarps.

Key P3: herbaceous dicots with alternate, simple, and pinnately lobed leaves on the stem

Copy permalink to share | Check for keys that lead to this key

1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.).
  2 Perianth uniseriate, with only undifferentiated tepals; flowers many and small, greenish or brownish, inconspicuous individually; inflorescence of glomerules that are usually further aggregated into racemes or panicles; fruit an achene or utricle
  2 Perianth biseriate, both sepals and petals present and differentiated (except uniseriate and of 2 white to cream-colored sepals 5-10 mm long in Macleaya in PAPAVERACEAE); flowers larger, usually with the petals prominently colored; inflorescence various, but not as above; fruit a capsule, silique/silicle, or a schizocarp of 2 mericarps.
    3 Corolla bilaterally symmetrical and the petals connate into a tube (or the corolla 2-lipped but the corolla lobes twisted so as to make the flower asymmetrical); stamens 4; fruit a 2-locular and loculicidal capsule opening by 2 valves
    3 Corolla radially symmetrical and either connate into a tube or distinct (except Reseda in RESEDACEAE, with bilateral symmetry but separate petals); stamens 5 or more; fruit a silique/silicle, a schizocarp of 4 mericarps, or a 1-, 3-, or 4-locular capsule (2-locular in Ipomoea in CONVOLVULACEAE and Glaucium in PAPAVERACEAE), opening variously.
      4 Ovary inferior; fruit either a schizocarp of 2 mericarps or a 4-locular capsule.
        5 Flowers 5-merous, aggregated into a head; fruit a schizocarp of 2 mericarps
      4 Ovary superior; fruit either a silique/silicle, or a 1-, 2-, or 3-locular capsule, or a berry.
          6 Sepals and petals of different numbers, the sepals 2-3, the petals 0, 4, or 6; stamens many
          6 Sepals and petals the same number, 4-8 each; stamens 5 or 6 (10-25 in Reseda in RESEDACEAE).
             7 Corolla bilaterally symmetrical; stamens 10-25
             7 Corolla radially symmetrical; stamens 5 or 6.
               8 Petals 4, distinct; stamens 6; fruit a silique/silicle
               8 Petals 5, connate into a tube; stamens 5; fruit either a capsule or a berry.
                   10 Corolla with a long tube, much longer than the lobes, scarlet, white, pink, or blue; leaves shallowly to deeply pinnately parted into 3-many lobes
                   10 Corolla with a short tube, the lobes longer than the tube, purplish; leaves with a single large terminal l lobe, and 2 small basal lobes (these almost separate as leaflets)
                     11 Fruit a capsule, 1-locular; corolla white, pink, lavender, or blue, the tube short (< 4 mm long), the lobes flaring, the corolla < 15 mm long or wide
                     11 Fruit either a capsule, 2- or-3 locular, or a berry; corolla scarlet, blue, white, yellow, greenish-yellow, or purple, the tube long (>10 mm long) and cylindrical, the corolla > 10 mm long or wide.
                       12 Stigmas 3; fruit a capsule with 3 valves; leaf lobes 0.5-5 mm wide, often themselves lobed, the sinuses very deep, often nearly to the leaf midvein
                       12 Stigmas 2; fruit either a capsule with 2 valves or a berry; leaf lobes > 5 mm wide, not lobed themselves, the sinuses shallow, often < 1/2 way to the midvein

Key Q: herbaceous dicots with whorled leaves on the stem {add [Platycodon] CAMPANULACEAE}

Copy permalink to share | Check for keys that lead to this key

  2 Cauline leaves essentially sessile, and also palmately cleft to the base, and further lacerately divided into linear or oblanceolate segments
  2 Cauline leaves petiolate, with 3-5, sessile or petiolulate, ovate, elliptic, or obovate leaflets (these serrate and sometimes with additional lobes).
    3 Inflorescence a spherical umbel of many flowers; fruit a drupe with 2-3 seeds; stem leaves 3-5
      4 Leaflets 3 (-5), sessile or subsessile, the petiolules 0-3 mm long; larger leaflets 4-8 cm long, 0.5-2.5 cm wide, averaging about 2.5× as long as wide, the apex obtuse to acute; fruit yellow-green when ripe, longitudinally winged and ribbed in ×-section; petals white (rarely tinged with pink); inflorescence nodding in bud; underground storage organ a spherical tuber
      4 Leaflets (3-) 5, petiolulate, the petiolules (7-) 10-25 mm long; larger leaflets 6-15 cm long, 3.5-7 cm wide, averaging about 1.8× as long as wide, the apex acuminate; fruit bright red when ripe, smoothly elliptical in ×-section; petals light green; inflorescence erect in bud; underground storage organ an +/- elongate root, this vertical or horizontal, and sometimes branched
    3 Inflorescence of single terminal flowers on the 1-several branches; fruit an aggregate of achenes; stem leaves 3
1 Cauline leaves simple.
        5 Inflorescence an involucrate head subtended by phyllaries, heads solitary or many, variously arrayed in secondary inflorescences; fruit a cypsela
        5 Inflorescence various, but not as above; fruit various, not as above (sometimes the flowers tightly grouped, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.).
          6 Fruit a 3-lobed, 3-locular capsule; inflorescence a cyathium, consisting of a single pistillate flower (reduced to a single 3-carpellate pistil) and 2 or more staminate flowers (each reduced to 1 stamen), borne in a cup-like involucre, the involucre bearing pointed or rounded glands, these sometimes brightly colored and petaloid, mimicking an individual flower (the cyathia then secondarily arranged in terminal cymes, or solitary and axillary, etc.); fresh plants with milky juice
          6 Fruit various, not as above; inflorescence not a cyathium (and staminate or bisexual flowers almost always with > 1 stamen); fresh plants lacking milky juice.
             7 Leaves succulent, the terminal whorls closely juxtaposed; pistils 4-5; fruit an aggregate of follicles
             7 Leaves herbaceous, thin in texture, whorls separated; pistil 1, of 2-5 fused carpels; fruit a capsule, achene, or drupe.
               8 Larger whorled leaves on a plant < 10 mm wide [some taxa keyed here and under the second lead].
                   10 Leaves markedly variable in shape or size in each whorl; fruit a capsule; petals 5
                   10 Leaves similar in size and shape in each whorl; fruit dry or fleshy, indehiscent; petals (3-) 4
                     11 Corolla bilaterally symmetrical, the petals connate; carpels 2; stamens 4, 6, or 8.
                     11 Corolla radially symmetrical, the petals separate; carpels 2, 3, or 5; stamens 5, 10, or many.
                          13 Inflorescence an axillary umbel; leaves narrowly linear and more than 10× as long as wide, > 20 mm long and < 2 mm wide; whorls of 3-6 leaves
                          13 Inflorescence a terminal cyme, raceme, panicle, or umbel; leaves as above, or broader in shape, narrower, or shorter; whorls of 3-16 leaves.
               8 Larger whorled leaves on a plant > 10 mm wide.
                                    18 Inflorescence of terminal involucrate clusters; perianth of 6 tepals; fruit an achene; stamens 9; [plants of very dry habitats]
                                    18 Inflorescence various, but not as above; perianth of 2 whorls (the calyx often obsolete in Galium in RUBIACEAE), 3-, 4-, 5-, 6-, or 7-merous; stamens 2-7; [plants of dry-mesic to very wet habitats].
                                           21 Corolla pink-purple, 6-merous, the petals separate and borne on the edge of a hypanthium; stamens 8, 10, or 12; [plants of wetlands]
                                           21 Corolla white, yellow, or greenish, 4-, 5-, or 7-merous, the petals fused at least basally into a tube (falling as a unit), not on a hypanthium; stamens 2, 4, 5, or 7; [plants of mesic habitats].
                                               23 Petals yellowish-white, with prominent green streaks; biennial or monocarpic plant, 10-30 dm tall when fertile; leaves 15-35 cm long
                                               23 Petals white or yellow; perennial plants, 1-15 dm tall; leaves 1-15 cm long.

Key R: herbaceous dicots with opposite, compound leaves on the stem

Copy permalink to share | Check for keys that lead to this key

1 Inflorescence an involucrate head subtended by phyllaries, heads solitary or many, variously arrayed in secondary inflorescences; fruit a cypsela
1 Inflorescence various, but not as above; fruit various, not as above.
      4 Plants vines, climbing or sprawling, the leaves often with tendril like petioles and leaf rachises; corolla 4-parted, radially symmetrical
      4 Plants not vines, more or less erect to arching and lacking adaptations for climbing; corolla 5-parted, radially or bilaterally symmetrical
        5 Plants often with a mix of mostly opposite, deeply 3-lobed leaves or trifoliate leaves (leaves sometimes simple); flowers pink or purplish-white, axillary; plants annual
        5 Plants typically with all leaves with > 3 leaflets; flowers primarily white, borne in dense terminal corymbs/cymes; plants perennial
          6 Cauline leaves essentially sessile, and also palmately cleft to the base, and further lacerately divided into linear or oblanceolate segments
          6 Cauline leaves petiolate, with 3-5, sessile or petiolulate, ovate, elliptic, or obovate leaflets (these serrate and sometimes with additional lobes).
             7 Plants annual; flowers typically unisexual and not showy, green; plants highly resinous and glandular; inflorescences variously arranged
             7 Plants perennial; flowers radially symmetric, showy, variously colored; plants typically not strongly resinous or glandular; inflorescences terminal

Key S1: herbaceous dicots with opposite, simple, and unlobed leaves on the stem

Copy permalink to share | Check for keys that lead to this key

1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head, e.g. Pycnanthemum in LAMIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.).
  2 Leaves scale-like, stems fleshy; flowers embedded in the fleshy stem, no perianth present; [saline environments (coastal or rarely inland)].
  2 Leaves small to large; stems not fleshy; flowers sessile or on pedicels; [collectively of many habitats, saline and not].
      4 Perianth of a single whorl (petals absent) (note that in Mirabilis in NYCTAGINACEAE the petaloid calyx is subtended by a 5-lobed fused set of involucral bracts).
        5 Leaves herbaceous, suborbicular, about as long as wide or wider than long; calyx 3- or 4-merous; stamens 4, 8, or 12.
          6 Plant ascending, with a single node (2 leaves); leaves > 6 cm long and wide; calyx 3-merous, brown to yellowish; stamens 12
          6 Plant creeping, with many nodes; leaves 3-15 mm long and wide; calyx 4-merous, yellow to greenish; stamens 4 or 8
        5 Leaves fleshy, linear, lanceolate, to broadly ovate, at least slightly longer than broad; calyx 5-merous; stamens 3, 5, or 10.
             7 Flowers axillary, sessile or nearly so, solitary or a few; petaloid sepals widely spreading, separate; leaves linear to oblanceolate; stamens 5 or 30-50
             7 Flowers in terminal cymose panicles; petaloid sepals connate into a narrow tube (reminiscent of the corolla of Ipomoea); leaves lanceolate, elliptic, ovate, or broadly ovate; stamens 3 or 5
      4 Perianth in 2 whorls (sepals and petals both present).
                 9 Leaves distinctly 3-veined from the base, the 3 veins converging again at the leaf apex
               8 Petals connate into a tube (at least basally); inflorescence often a head or dense terminal cyme (also axillary, or solitary on long peduncles).
                   10 Petals 5; stamens 3, 4, or 5
                     11 Upright herb; flowers in axils or terminal corymbs
                       12 Flowers in terminal corymbs; fruit dry
                   10 Petals 4 (or 6 or 8 in Richardia in RUBIACEAE); stamens 4, 6, or 8.
                          13 Inflorescence a head or more diffuse (see below), sometimes subtended by green bracts.
                            14 Leaves serrate; corolla bilaterally symmetrical (especially the flowers near the outer edge of the head); inflorescence a head
                            14 Leaves entire; corolla radially symmetrical; inflorescence a head or more diffuse (see below).
                              15 Petals acute; flowers in terminal panicles, cymes, or panicles, or axillary; plant habit various, not simultaneously with all the characters below
                              15 Petals broadly rounded; flowers axillary, solitary; plant a diffusely branched herb with linear leaves
                                16 Perianth of a single whorl (petals absent) or missing entirely (petals and sepals both absent). {key lead number needs adjusting}
                                  17 Inflorescence a cyathium, consisting of a single pistillate flower (reduced to a single 3-carpellate pistil) and 2 or more staminate flowers (each reduced to 1 stamen), borne in a cup-like involucre, the involucre bearing pointed or rounded glands, these sometimes brightly colored and petaloid, mimicking an individual flower (the cyathia then secondarily arranged in terminal cymes, or solitary and axillary, etc.); fresh plants with milky juice; fruit a 3-lobed, 3-locular capsule
                                  17 Inflorescence not a cyathium (and staminate or bisexual flowers with > 1 stamen, except Callitriche in PLANTAGINACEAE); fresh plants lacking milky juice; fruit various, not as above.
                                    18 Flowers many, in axillary spikes, cymes, or glomerules, or in terminal spikes, heads, cymes, or panicles; leaves entire or serrate.
                                             22 Leaves entire, or with a few very obscure crenations (Iresine) or basally disposed rounded lobe-like teeth (Atriplex); plants without stinging hairs.
                                                 24 Style 1; leaves generally either longer than 30 mm, or wider than 8 mm (if linear and smaller than those dimensions, then fleshy).
                                16 Perianth in 2 whorls (sepals and petals both present).
                                                        27 Pistils 4-5, in a single whorl; stamens 4, 5, 8, or 10; fruit an aggregate of follicles
                                                      26 Gynoecium either of 1 pistil (with 1 or more carpels), or of 2 pistils, united only by the style and stigma (APOCYNACEAE).
                                                            29 Leaves with pellucid punctate glands (most easily visible with transmitted light); stamens often fascicled into 3, 4, or 5 fascicles; petals yellow or pinkish
                                                                 31 Petals 3; sepals 5, dimorphic, the 2 outer sepals narrower than the 3 inner and concave sepals; stamens (3-) 5-15 (-25)
                                                                 31 Petals 4-7; sepals 4-7, normally monomorphic; stamens 4, 5, 6, 8, 10, or 12 (or sometimes rarely 2 or 3).
                                                                     33 Capsule 1-locular, dehiscent apically by teeth or valves; sepals connate into a tube or separate; styles 2-5; perianth 4-5-merous; stamens 4, 5, 8, or 10 (or rarely 2 or 3)
                                                                       34 Corolla radially symmetrical (or so slightly bilaterally symmetrical as to be mistakable as radially symmetrical); stamens as many as the corolla lobes (or 1 less in Ruellia in ACANTHACEAE, Buchnera in OROBANCHACEAE, Trichostema in LAMIACEAE, and Verbena in VERBENACEAE); carpels 2 or 3.
                                                                         35 Pistils 2, united only by the style and stigma; fruit a schizocarp of 2 1-carpellate follicles (often single by abortion); plant with milky juice when fresh (except Catharanthus); leaves entire
                                                                         35 Pistil 1 (of 2-5 fused carpels); fruit either a 2-5-carpellate capsule or of 2 or 4 1-seeded nutlets derived from 2 carpels; plant lacking milky juice; leaves entire or serrate.
                                                                                  39 Stamens either 4, 1 fewer than the 5 corolla lobes, or 2 (with 2 staminodes); corolla usually slightly bilaterally symmetrical (the flower as a whole made bilaterally symmetrical by the 2 or 4 stamens).
                                                                       34 Corolla bilaterally symmetrical (or the corolla 2-lipped but the corolla lobes twisted so as to make the flower asymmetrical); fertile stamens fewer than the corolla lobes (except Plantago in PLANTAGINACEAE, which is equal, with 4 each; a few genera have a 5th, sterile, stamen which is obviously different in form than the 4 fertile stamens) (note that many corollas are bilabiate and the number of corolla lobes, 4 or 5, may be difficult to interpret); carpels 2.
                                                                                             44 Carpels 2, each carpel slightly to deeply lobed, separating at maturity into 4 half-carpellate units (not separating in Phyla in VERBENACEAE); fruit a schizocarp of 4 mericarps (or 2 nutlets in Phyla in VERBENACEAE).
                                                                                               45 Inflorescence a thyrse, verticillaster, or terminal cyme, the flowers borne in cymose lateral branches; corolla strongly bilaterally symmetrical (rarely nearly radially symmetrical); stems square in ×-section (or sometimes rounded, especially on older growth); fresh plants often (but not always) aromatic
                                                                                               45 Inflorescence of spikes, racemes, or heads, the flowers or fruits single at nodes; corolla often nearly radially symmetrical; stems rounded in X-section (rarely square); fresh plants usually not aromatic
                                                                                                                     55 Inflorescence of 1 or more terminal (and sometimes upper axillary) spikes or racemes; corolla 10-50 mm long (6-8 mm long in Phryma in PHRYMACEAE), white, pink, blue, purple, or yellow; fruit either a loculicidal capsule (OROBANCHACEAE) or a single seeded achene (Phryma in PHRYMACEAE).
                                                                                                                53 Flowers axillary and solitary, borne in the axils of normally-sized leaves or somewhat reduced but still large and leaf-like bracts [some taxa keyed here and below].
                                                                                                                                      63 Corolla red or orange, with a very narrow, cylindrical tube, the lobes then flaring into a limb about 1 cm across; plants blackening on drying; [rare exotic, in crop fields, a noxious hemiparasitic weed under quarantine]

Key S2: herbaceous dicots with opposite, simple, and palmately lobed leaves on the stem {add [Humulus] CANNABINACEAE}

Copy permalink to share | Check for keys that lead to this key

1 Leaf lobes very narrow, < 3 mm wide; inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela
1 Leaf lobes broad, >20 mm wide; inflorescence, flower, and fruit structure various, but not with the combination of features as above.
  2 Leaves >4 per above-ground stem; perianth 5-merous; flowers bilaterally symmetrical, the corolla with connate petals, lavender-white with yellow markings in the throat; fruit a large curved capsule
  2 Leaves 1-2 per above-ground stem; perianth 3-merous; flowers radially symmetrical, the corolla absent or with distinct petals, white; fruit a berry or aggregate of berries.
    3 Leaves with peltate petiole attachment; carpel 1; petals present, white
    3 Leaves with petiole attached marginally; carpels many, as separate pistils; petals absent

Key S3: herbaceous dicots with opposite, simple, and pinnately lobed leaves on the stem

Copy permalink to share | Check for keys that lead to this key

1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers tightly grouped, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.).
  2 Flowers tiny, individually inconspicuous; perianth absent or vestigial; fruit a utricle
  2 Flowers larger, individually conspicuous; perianth present, the petals or sepals brightly colored; fruit a capsule (or aggregate of achenes in Clematis in RANUNCULACEAE or schizocarp of 4 mericarps in Glandularia in VERBENACEAE).
    3 Flowers radially symmetrical; stamens 5 or many; fruit a capsule or aggregate of achenes.
      4 Stamens many; fruit an aggregate of plumose achenes
    3 Flowers bilaterally symmetrical (sometimes only slightly so); stamens 4 (or 2 in Veronica in PLANTAGINACEAE); fruit a capsule or schizocarp of mericarps.
        5 Inflorescence of cymosely arranged spikes or heads; fruit a schizocarp of 4 nutlets
        5 Inflorescence of solitary axillary flowers or terminal racemes.
          6 Corolla yellow, orange, or red; plants often drying black (but not Striga); sepals connate into a tube at least 1/3 as long as the corolla lobes; calyx 5-merous
          6 Corolla white, pink, lavender, or blue; plants not drying black; sepals distinct or only shortly connate into a short tube, the calyx lobes much longer than the tube; calyx 5- or 4-merous