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5 Inflorescence solitary (Illicium in ILLICIACEAE) or variously branched, spicate, racemose, or fascicled, not an involucrate head. | |
9 Leaves largely covered with silver and/or bronze lepidote scales and/or dense stellate hairs below (visible at 10× or higher magnification), giving the lower leaf surface a slightly shiny to almost metallic appearance. {add Lyonia ferruginea and L. fruticosa in ERICACEAE; add Loropetalum in HAMAMELIDACEAE} | |
13 Leaves 1-foliolate on the upper stems, sometimes 3-foliolate below, or all reduced to phyllodial spines; flowers papilionaceous, bright yellow; fruit a legume; stems bright green | |
13 Leaves simple throughout; flowers either small, inconspicuous, tannish, borne in catkins (Quercus), or larger and urceolate, or with almost separate and spreading petals, white to pink or reddish-orange, in various terminal or axillary, branched inflorescences; fruit either a nut in a cupule (an acorn), or a (3-) 5-valved capsule, or a spherical berry or drupe; stems generally brown or tan (sometimes green). | |
17 Leaves 1 per node or also paired (on one side of the stem) at some nodes (the leaves then uneven in size); inflorescences leaf-opposed; fruit a berry. | |
17 Leaves 1 per node; inflorescences terminal or axillary, never leaf-opposed; fruit a valved capsule. | |
19 Flowers few, not showy, green to greenish-white, inflorescence not densely arranged (flowers also sometimes solitary); capsules not beaked; leaves not revolute or undulate | |
19 Flowers numerous, showy, white; inflorescence densely arranged; capsules with short beak; leaf margins revolute or slightly undulate | |
1 Leaves deciduous. {add: Ditrysinia in EUPHORBIACEAE, Glochidion in PHYLLANTHACEAE, Phyllanthopsis in PHYLLANTHACEAE, Nierembergia in SOLANACEAE, Edgeworthia in THYMELAEACEAE, Ipomoea (I. carnea) in CONVOLVULACEAE; Swida (S. alternifolia) in CORNACEAE} | |
22 Inflorescence branched, spicate, a catkin, or consisting of a solitary flower or axillary clusters or whorls, not an involucrate head. | |
25 Inflorescences axillary, solitary flowers | |
25 Inflorescence of 2 or more flowers; perianth 3-5-merous; fresh plants not musky-fragrant; fruits various, not as above. | |
27 Flowers 3-merous; fruit fleshy, red or greenish-yellow at maturity; ovary superior; [Basal Angiosperms or Eudicots]. | |
29 Fruit either a drupe or berry (indehiscent, and variously fleshy or dry) or a dry 3-valved capsule with 1 seed; inflorescence axillary (solitary, clusters, fascicles, or racemes), or in a terminal raceme (Pyrularia in SANTALACEAE). | |
30 Leaves with various vestiture, but not as above. | |
31 Ovary inferior or half-inferior; inflorescence an axillary cluster or raceme, or a terminal raceme. | |
32 Fruit an elongate drupe (definitely longer than thick), with 1 seed. | |
34 Fruits spherical, < 10 mm long. | |
36 Inflorescence a narrowly cylindrical raceme, clustered several to many at the tip of the previous year’s wood and below the current season’s growth; fruit < 3 mm in diameter | |
37 Fruit fleshy, with 4-8 seeds; leaf pubescence simple or absent. | |
38 Fruit yellow to red, the pedicel 10-30 mm long; leaf venation pinnate, but irregular and reticulated | |
38 Fruit dark red to black, the pedicel < 10 mm long; leaf venation very neatly pinnate, with the secondary veins nearly straight and parallel to one another |
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1 Subshrubs or dwarf shrubs, aboveground stems creeping or erect, < 15 cm tall; leaves evergreen. | |
1 Shrubs, aboveground stems erect, > 30 cm tall; leaves evergreen or deciduous. | |
5 Inflorescence an involucrate (composite) head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
5 Inflorescence, flower, and fruit structure various, but not with the combination of features as above. | |
8 Petals distinct, yellow or white; flowers unisexual or bisexual; leaves oblanceolate or elliptic, broadest towards the tip or near the middle, the teeth usually concentrated in the upper half of the leaf; fruit either a black or red drupe with several pyrenes, a red berry with several seeds, or red or black pomes. | |
6 Leaves deciduous {add [Fagaceae]}. | |
12 Leaves crenate-wavy, with 1-2 teeth per cm of leaf margin; leaves usually obliquely cordate or angled-truncate at the base; pubescence of leaves and stems stellate | |
12 Leaves crenulate, serrate or serrulate, with >2 teeth per cm of leaf margin; leaves cuneate, rounded, or subcordate at base, not oblique; pubescence of leaves and stems absent or simple. | |
13 Leaves prominently 3-veined from the base. | |
13 Leaves pinnately veined. | |
23 Petals separate (or absent in Rhamnus alnifolia); stamens opposite to the petals (when present) and alternate to the sepals; fruit 2-4-locular, with 2-4 pyrenes | |
27 Stamens 10; ovary and capsule 3-locular; leaves obovate (widest towards the apex), the teeth obscure to coarse (usually < 4 points per cm of margin), and primarily in the upper half of the leaf; inflorescence a terminal or axillary raceme or cyme; hairs of the lower leaf surface either simple and appressed, or stellate. | |
28 Leaf margins regularly and evenly serrate in the upper half of the leaf (usually nearly entire towards the base); inflorescence an elongate, many flowered (>30) raceme borne at the end of branchlets of the season; corolla of separate petals, the stamens separate; hairs of the lower leaf surface simple and appressed | |
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3 Creeping or short subshrubs, the stems primarily prostrate, < 2 dm tall. | |
4 Well-developed leaves 4-6 per stem; inflorescence a head subtended by 4 large white bracts | |
4 Well-developed leaves many per stem; inflorescence of individual flowers axillary in pairs or clusters or in terminal cymes. | |
8 Upright shrubs, unarmed. | |
9 Inflorescence a terminal head of many flowers. | |
10 Head flattened, either subtended by 4 large white bracts or by an involucre with >5 green phyllaries. | |
11 Head subtended by an involucre of >5 green phyllaries; leaves with venation otherwise; flowers 5-merous | |
9 Inflorescence otherwise, either of a solitary flower, or one of a wide variety of inflorescences with flowers attached at different points along branched or unbranched axes (e.g. axillary). {add: [Lagerstroemia] LYTHRACEAE; [Rosmarinus] LAMIACEAE; [Laguncularia] COMBRETACEAE; [Buxus] BUXACEAE; [Exochorda] ROSACEAE; various other [see spreadsheet]} | |
12 Inflorescence either terminal and not flat-topped (longer than wide), or axillary and variously shaped, or terminal and solitary, or leaf-opposed. | |
15 Carpels many (> 9), either separate or fused; stamens many; perianth segments either many and undifferentiated into calyx and corolla, maroon, brown, or yellow (in CALYCANTHACEAE), or differentiated into a fleshy and persistent calyx of 5-9 sepals, and a deciduous corolla of 5-9 red (or white) petals (Punica in LYTHRACEAE). | |
16 Fruit a leathery, 4-15 cm in diameter, reddish, spherical berry with obpyramidal seeds surrounded by a juicy sarcotesta (pomegranate); perianth differentiated, the sepals fleshy and persistent on the fruit, the petals deciduous, 5-9, bright red to white; ovary inferior; branches typically armed with axillary spines | |
15 Carpels 1-5, fused; stamens either 1-5 or 8-10; perianth segments 4-5 or 8, variously colored; fruit a simple capsule, drupe, or berry (including berry-like fruit); flowers 2-many, in axillary or terminal inflorescences (pistillate flowers sometimes solitary in SANTALACEAE and THESIACEAE); [Eudicots]. | |
21 Stamens 8-10, of 2 different lengths in each flower; petals separate, 4-5 (-7), pink purple, 10-15 mm long; stems strongly arching, rooting at the tips; [plants of flooded to saturated wetlands] | |
23 Fruit a loculicidal capsule, dehiscing into 3 valves; branches square in ×-section; leaves < 2 cm long; [exotic, cultivated and weakly established, of temperate areas] | |
23 Fruit a drupe with 2-4 pyrenes; branches round or nearly so in ×-section; leaves > 2 cm long; [natives, of peninsular FL] | |
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1 Leaves either 2-3-foliolate or palmately 4-11-foliolate (all the leaflets attached at a common point). | |
2 Leaves either 3-foliolate or palmately or pedately 4-11-foliolate. | |
3 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
3 Inflorescence, flower, and fruit structure various, but not with the combination of features as above. | |
7 Flowers radially symmetrical; fruit a 2-valved or 5-valved capsule; [plant of uplands or wetlands] | |
8 Leaflets notched at the tip; flowers pink, white, or yellow; [plants of uplands or temporarily flooded wetlands] | |
9 Petals 5 or more; stamens 10 or more; fruit either a legume or an aggregate of achenes or follicles | |
10 Stamens many, fused into a staminal tube; carpels 10-20, in a ring; pubescence stellate (sometimes mixed with simple hairs) | |
10 Stamens 10-many, separate, or fused but not all into a staminal tube; carpel either 1 (FABACEAE), or 3-7 in a ring (RANUNCULACEAE), or many and spirally arranged on a conical receptacle (RANUNCULACEAE or ROSACEAE) | |
12 Fruit an aggregate of achenes (borne on a fleshy, expanded receptacle in Fragaria and some Potentilla) | |
1 Leaves 1-pinnately compound (all leaflets attached to a central rachis) or more complexly compound (with several orders of branching, some leaflets at least attached to second-order branches from the rachis). | |
15 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
15 Inflorescence, flower, and fruit structure various, but not with the combination of features as above. | |
16 Flowers radially symmetrical; fruit a silique/silicle, or a schizocarp of mericarps, or an achene. | |
17 Petals 0 or 5 (if 0, the sepals petaloid); stamens 2, 4, 5, or many. | |
14 Leaves more complexly compound (with 2 or more orders of branching, some leaflets at least attached to second-order branches from the rachis). | |
19 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
19 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head subtended by bracts, but then with other features differing, such as stamens 4, or green calyx present, or petals separate, or fruit a schizocarp of mericarps, etc.). | |
21 Inflorescence various, but not an umbel; ovary superior; fruit an aggregate of follicles or an elongate capsule. | |
23 Inflorescence various, but not an umbel; fruit an aggregate of follicles or achenes, an elongate capsule, or a naked seed resembling a drupe. | |
26 Corolla bilaterally symmetrical; fruit an elongate capsule; [cultivated exotic, rarely persistent near gardens] | |
27 Leaflets 1-6 cm long, about as long as broad if > 4 cm long; pistils 4-many; fruit an aggregate of follicles or achenes; flowers mainly 4-5-merous | |
28 Pubescence of the stem and lower leaf surface glandular; flowers unisexual, on the same plant (monoecious); stamens 10; pistils 2, partly fused; fruit an aggregate of follicles | |
28 Pubescence of the stem and lower leaf surface non-glandular (or absent); flowers either bisexual (the plants hermaphroditic), or unisexual and the male and female flowers on separate plants (the plants dioecious); stamens 15 or more; pistils 1-8, separate; fruit an aggregate of follicles, a follicle, or a red or white berry. | |
29 Flowers bisexual (plants hermaphroditic); carpels 1-8 per flower; inflorescence a raceme, or a panicle of racemes with just a few branches; fruit an aggregate of follicles, a follicle, or a red or white berry | |
29 Flowers unisexual (plants dioecious); carpels 3-4 per pistillate flower; inflorescence a panicle of racemes, with numerous branches; fruit an aggregate of follicles |
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1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head subtended by bracts, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or petals separate, or fruit a schizocarp of mericarps, etc.). | |
2 Basal leaves 2-lobed, pinnately lobed, or palmately lobed (not considering cordate, hastate, or auriculate leaf bases as “lobed”). | |
4 Leaf lobing pinnate. | |
8 Corolla 2-lipped, bilaterally symmetrical or asymmetrical; stamens 2 or 4. | |
4 Leaf lobing palmate. | |
13 Petals 5; stamens 5; fruit a schizocarp of 2 mericarps. | |
16 Perianth of 5 green sepals and 5 colored petals. | |
18 Leaves 2, the single flower terminal and associated with the upper leaf; fruit an aggregate of berries | |
19 Hypanthium present, partially fused or not fused to the pistil; ovary partially inferior to superior | |
23 Leaf margins entire. | |
24 Inflorescence a terminal and/or axillary raceme, panicle, or cyme of many small flowers; fruit an achene; perianth uniseriate, of 0, 4-5, or 6 tepals. | |
24 Inflorescence either a terminal spike, or a 1-7-flowered terminal cyme, or of a solitary axillary or terminal flower; fruit various; perianth biseriate (of differentiated sepals and petals (except uniseriate, of 3 fused sepals in ARISTOLOCHIACEAE). | |
26 Flowers bilaterally symmetrical; inflorescence a terminal spike (with > 20 flowers); petals 4, usually scarious, transparent; sepals 4, green; stamens 4 | |
26 Flowers radially symmetrical; inflorescence either of a solitary flower or of a 1-7-flowered terminal cyme; petals 5, 8-12, or 0; sepals 5 (green), 3 (brown), or 5-9 (yellow); stamens 5, 12, or many. | |
27 Gynoecium either of a single pistil with 6 carpels or of a single pistil with 4 carpels or of 2 nearly separate carpels; fruit a simple capsule (or deeply 2-lobed); flowers white, brown, or greenish, either of 5 fused or distinct white petals and 5 fused or distinct green sepals, or of 3 fused brown or greenish petaloid sepals. | |
28 Flowers brown or green, of 3 fused brown or greenish petaloid sepals (and 0 petals); carpels 6; stamens 12; leaves 4-10 cm wide | |
29 Petals fused; sepals fused; carpels 2; plant pubescent. | |
32 Carpels many, separate, spiral; petals yellow or white. | |
33 Flowers lacking a hypanthium; fruit an aggregate of achenes or aggregate of follicles | |
34 Flowers radially symmetrical; inflorescence an umbel (or composite of umbelliform units, or a terminal panicle. | |
36 Inflorescence a terminal raceme; perianth of 4 green sepals and 4 white petals; fruit a silique/silicle; fresh foliage in spring and summer with a strong garlic odor; larger leaves < 10 cm in diameter | |
35 Ovary inferior; inflorescence an umbel (or a composite of umbellate units); fruit a schizocarp of mericarps. | |
39 Leaves tubular, with a sutured ventral flange, erect or reclining, adapted as a pitfall for insects (flat, phyllodial leaves sometimes present as well, common in the winter in some species, such as S. oreophila) | |
39 Leaves flat, not sutured into a tubular shape. | |
43 Corolla white, lavender, or blue, 2-lipped and bilaterally symmetrical. | |
40 Stem leaves alternate; perianth radially symmetrical (less commonly bilaterally symmetrical); stamens 5, 6-8, 9, 10 (rarely 4). | |
49 Corolla bilaterally symmetrical (barely so in Limosella in SCROPHULARIACEAE); stamens 2, 4, 6, 8, or 10. | |
49 Corolla radially symmetrical; stamens 5, 10, 4-6, or 9. | |
56 Perianth of green sepals and more brightly colored petals; stamens 5 or 10; carpels 2, 3, 4, or 5. | |
57 Leaves lacking sticky gland-tipped hairs. | |
59 Inflorescence of a solitary, terminal flower; carpels 2-3 (-4). | |
60 Corolla of separate petals or united only for a short length (< ¼ its length). | |
59 Inflorescence of several to many flowers; carpels 5 (3 in Galax in DIAPENSIACEAE). | |
64 Inflorescence a somewhat to very diffuse panicle, with 3 or more orders of branching, not giving at all the impression that the overall inflorescence is made of racemose units. | |
66 Inflorescence of 1-several terminal and axillary racemes, the plant typically well-branched, especially from the base; stamens 5 | |
66 Inflorescence of a single, terminal raceme, the plant unbranched; stamens 10 (or 5, with 5 staminodes) | |
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2 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
2 Inflorescence, flower, and fruit structure various, but not with the combination of features as above. {add under 2b [Cynosciadium] APIACEAE} | |
7 Leaflets radially arranged at the summit of the petiole, not differentiated in size or placement into a terminal leaflet and 2 lateral leaflets; leaflets prominently notched at the apex; petals 5, yellow; inflorescence axillary, cymose or umbelliform; carpels 5 | |
1 Leaves either 1-pinnately compound (all leaflets attached to a central rachis) or more complexly compound (with several orders of branching, some leaflets at least attached to second-order branches from the rachis). | |
10 Inflorescence an involucrate head subtended by phyllaries, heads solitary or many, variously arrayed in secondary inflorescences; fruit a cypsela; ovary inferior | |
12 Flowers bilaterally symmetrical, papilionaceous; fruit a legume; leaves even-pinnately or odd-pinnately compound, the terminal leaflet sometimes replaced by a tendril; leaflets entire or at most minutely denticulate | |
12 Flowers radially symmetrical (or barely bilaterally symmetrical in Erodium in GERANIACEAE); fruit a capsule, capsular but of 5 mericarps, or an aggregate of achenes, nutlets, or follicles (in some cases the # of pistils from many down to 2 or even 1); leaves odd-pinnately compound, never with tendrils; leaflets serrate (or entire to shallowly lobed in Polemonium in POLEMONIACEAE, Cardamine in BRASSICACEAE, and Floerkea in LIMNANTHACEAE). | |
13 Pistils many (only 1-2 in Agrimonia, Poteridium, Poterium, and Sanguisorba); fruit an aggregate of achenes, nutlets, or follicles; hypanthium present; stamens 5-many (only 4 in Poteridium and Sanguisorba) | |
13 Pistil 1 (or deeply 2-3-lobed in Floerkea in LIMNANTHACEAE); fruit a silique, capsule, schizocarp of 2-3 mericarps, or a capsular schizocarp of 5 mericarps (Erodium in GERANIACEAE); hypanthium absent; stamens 3-6. | |
15 Petals 5, fused (distinct in Erodium in GERANIACEAE); stamens 5; inflorescence axillary or terminal, cymose, consisting of subcapitate, umbel-like, or helicoid cymes; fruit either a capsule, or a capsular schizocarp of 5 mericarps (Erodium in GERANIACEAE). | |
16 Flowers slightly bilaterally symmetrical (2 of the petals of different size than the other 3); fruit a capsular schizocarp of 5 mericarps; carpels 5 | |
17 Fruit a capsule; fresh plant not aromatic. | |
11 Leaves more complexly compound (with 2 or more orders of branching, some leaflets at least attached to second-order branches from the rachis). | |
19 Leaves either 2× odd-pinnate or more complexly 2-4× ternately or ternately-pinnately compound; flowers in various inflorescences; fruits various (not legumes or loments). | |
20 Leaf segments linear, less than 2 mm wide. | |
21 Inflorescence either a terminal solitary flower or terminal raceme or panicle; ovary superior, either of 2 fused carpels or of 1-5or many distinct 1-carpellate pistils; fruit either a capsule or an aggregate of follicles or achenes. | |
23 Erect or sprawling herb; stamens 5-6 or >15. | |
25 Inflorescence an umbel; ovary inferior, of 2 fused carpels; fruit a schizocarp of 2 mericarps; inflorescence an umbel | |
25 Inflorescence a panicle or raceme; ovary superior, of 1-8 carpels; fruit an aggregate of follicles, a single follicle, or an indehiscent berry-like fruit. | |
26 Flowers bisexual; carpels 1-8; fruit an aggregate of follicles, a single follicle, or an indehiscent berry-like fruit | |
26 Flowers unisexual; carpels (in pistillate flowers) of 3 (-5) carpels; fruit an aggregate of follicles | |
27 Inflorescence a raceme, panicle, or cyme; ovary superior, of either 1-2 fused carpels or of many separate 1-carpellate pistils. | |
29 Perianth of 4-5 whorls of 3 parts each (some of the whorls modified into nectaries); pistil 1, 1-carpellate; fruit a drupelike, blue, naked seed; largest leaflets > 6 cm long, obviously longer than wide | |
29 Perianth of 1 whorl; of 4-5 parts; pistils many, each 1-carpellate; fruit an aggregate of achenes or an aggregate of follicles; largest leaflets either < 6 cm long, or if longer than 6 cm, also about as wide as long |
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1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.). | |
2 Perianth uniseriate (represented only by undifferentiated tepals or sepals) or completely absent; flowers usually unisexual, less commonly bisexual). | |
3 Inflorescence a cyathium, consisting of a single pistillate flower (reduced to a single 3-carpellate pistil) and 2 or more staminate flowers (each reduced to 1 stamen), borne in a cup-like involucre, the involucre bearing pointed or rounded glands, these sometimes brightly colored and petaloid, mimicking an individual flower (the cyathia then secondarily arranged in terminal cymes, or solitary and axillary, etc.); fresh plants with milky juice; fruit a 3-lobed, 3-locular capsule | |
3 Inflorescence not a cyathium (and staminate or bisexual flowers with > 1 stamen; fresh plants lacking milky juice (except Stillingia in EUPHORBIACEAE); fruit various, not as above. | |
7 Leaf bases cordate to rounded. | |
8 Styles 3; fruit a 3-lobed, 3-carpellate capsule (1 carpel sometimes aborting); inflorescence either a terminal or leaf opposed raceme, or a dense axillary condensed cyme with conspicuous toothed bracts subtending the flowers | |
4 Leaf margins entire. | |
11 Leaf base cuneate, rounded, or truncate; calyx of 3-4-5 distinct sepals, radially symmetrical, white or yellow; fruit a dry, nutlike drupe or an achene. | |
14 Inflorescence a leaf-opposed (sometimes apparently terminal) spike or raceme; flowers visually white from white petaloid sepals, white bracts, or white stamens. | |
15 Sepals absent; carpels 3-4; stamens 2-6 (-8); fruit a capsule, a 1-seeded drupe, or a schizocarp of 3-4 mericarps; leaf bases cordate or subcordate; [Basal Angiosperms]. | |
14 Inflorescence not leaf-opposed, either simpler (single axillary or glomerules of flowers) or more complexly branched (axillary or terminal panicles or complex cymes); flowers white, reddish, scarious, or greenish. | |
24 Petals connate. | |
25 Inflorescence various, either a terminal head, or axillary and solitary, or variously axillary or terminal and more diffuse. | |
28 Petals 4-7; stamens 1× or 2× as many as the petals, 4-7, 8, 10, 12, or14; leaves herbaceous in texture | |
30 Petals connate (at least basally), 4, 5, 6, 7, or 8; carpels 1, 2, 4, 5, or 6 (rarely 3 in Reseda in RESEDACEAE); fruit a legume or 1-, 2-, or 5-loculed capsule (except a 1-seeded indehiscent pod in Krameria in KRAMERIACEAE). | |
31 Stamens 6-10 (-25), more than the number (4 or 5) of petals and the number (4 or 5) of the sepals; fruit a legume or a 1-6-carpellate capsule. | |
35 Pistil 5-carpellate; capsule 5-locular, explosively dehiscent; inflorescence axillary, small clusters of flowers | |
35 Pistil 2-carpellate; capsule 2 locular, opening gradually or not at all; inflorescence a terminal spike, raceme or panicle (or solitary, axillary flowers in Chaenorrhinum in PLANTAGINACEAE and Krameria in KRAMERIACEAE). | |
36 Fruit 1-seeded, indehiscent; sepals petaloid, red-purple; petals dimorphic, the upper 3 long-clawed, the lower 2 small, thickened, and glandlike | |
37 Stamens 5; corolla not spurred; capsule septicidal; pubescence of the stem and leaves either gland-tipped or dendritically branched | |
37 Stamens 4; corolla with a distinct spur or sac at the base between the the 2 lower calyx lobes (except not spurred in Digitalis and Schwalbea); capsule loculicidal (only at the summit in Antirrhinum and Chaenorrhinum, and septicidal in Schwalbea); pubescence of the stem and leaves neither gland-tipped (except in Antirrhinum and Chaenorrhinum) nor dendritically branched. | |
40 Pistils 4-10 (each 1-carpellate) in a ring, these sometimes fused basally, each with its own style/stigma; fruit either an aggregate of achenes or follicles or a 5 (-7) locular capsule. | |
42 Fruit an aggregate of follicles; leaves fleshy in texture; inflorescence; leaves entire of sparsely and coarsely serrate, with < 12 points per leaf; [plants primarily of dry habitats] | |
44 Petals 5 (rarely 4 or 6); sepals 5 (rarely 4 or 6); stamens 5 (or multiples of 5), 6, or 12; fruit a capsule or a ring of mericarps. | |
47 Flowers 6-merous (the petals and sepals 6, the stamens 6 or 12); corolla pink or purplish (rarely white); fruit a septicidal capsule | |
47 Flowers 5-merous (the petals and sepals 5, stamens 5 or various multiples of 5); corolla yellow, reddish, or blue; fruit a loculicidal or septicidal capsule. | |
48 Stamens 5; corolla yellow or blue; capsule either 10-locular and septicidal or 1-locular (with 3 carpels) and loculicidal. | |
48 Stamens (5-) 10, 15, 20, 30 (-many); corolla white, pink, yellow, or reddish; capsule 3-, 5- (-10)-locular, loculicidal. | |
50 Stamens (5-) 10, 15, 20, 30 (-many); corolla yellow or reddish; capsule 3 (-10)-locular, loculicidal | |
51 Pistils 2, united only by the style and stigma; fruit a schizocarp of 2 follicles (often single by abortion); plant with milky juice when fresh; leaves entire; inflorescence an umbel | |
55 Plant an herb, erect or sprawling; leaves > 1.5 cm long. | |
61 Flowers sessile or very-short pedicelled, solitary in the leaf axils. | |
61 Flowers either solitary and obviously pedicelled, or several in an axillary or lateral inflorescence. | |
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1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.). | |
6 Ovary superior; inflorescence various, not an umbel; fruit various, a capsule, an aggregate of achenes or follicles, a ring of (>2) mericarps. | |
8 Perianth biseriate (uniseriate in Aphanes in ROSACEAE and in Trautvetteria in RANUNCULACEAE); flowers bisexual; plants lacking stinging hairs. | |
10 Perianth bilaterally symmetrical, either hooded or spurred; fruit an aggregate of follicles | |
11 Stamens showy, bright white, dilated towards the tip; pistils ca. 15; fruit an aggregate of utricles | |
11 Stamens not showy, white, or dilated towards the tip; pistils many (> 25); fruit an aggregate of achenes. | |
14 Corolla bilaterally symmetrical, the petals connate (except distinct in Delphinium in RANUNCULACEAE); fruit a capsule, a follicle, or a schizocarp of 3 1-seeded mericarps. | |
14 Corolla radially symmetrical, the petals distinct (fused and tubular in Ipomoea); fruit a capsule or a schizocarp consisting of a ring of 5-many 1-seeded mericarps. | |
19 Stamens many, connate into a stamen tube; carpels 5-many, completely or only loosely fused; fruit a capsule or a schizocarp of 5-many mericarps borne in a ring; calyx often subtended by an epicalyx (an additional calyx-like, green, foliaceous whorl of bracts) | |
19 Stamens 5 or 10, distinct; carpels 2 or 5, fused; fruit a capsule or a schizocarp of 5 1-seeded mericarps. | |
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1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.). | |
2 Perianth uniseriate, with only undifferentiated tepals; flowers many and small, greenish or brownish, inconspicuous individually; inflorescence of glomerules that are usually further aggregated into racemes or panicles; fruit an achene or utricle | |
2 Perianth biseriate, both sepals and petals present and differentiated (except uniseriate and of 2 white to cream-colored sepals 5-10 mm long in Macleaya in PAPAVERACEAE); flowers larger, usually with the petals prominently colored; inflorescence various, but not as above; fruit a capsule, silique/silicle, or a schizocarp of 2 mericarps. | |
3 Corolla radially symmetrical and either connate into a tube or distinct (except Reseda in RESEDACEAE, with bilateral symmetry but separate petals); stamens 5 or more; fruit a silique/silicle, a schizocarp of 4 mericarps, or a 1-, 3-, or 4-locular capsule (2-locular in Ipomoea in CONVOLVULACEAE and Glaucium in PAPAVERACEAE), opening variously. | |
7 Corolla radially symmetrical; stamens 5 or 6. | |
9 Plant an erect or sprawling herb. | |
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4 Leaflets 3 (-5), sessile or subsessile, the petiolules 0-3 mm long; larger leaflets 4-8 cm long, 0.5-2.5 cm wide, averaging about 2.5× as long as wide, the apex obtuse to acute; fruit yellow-green when ripe, longitudinally winged and ribbed in ×-section; petals white (rarely tinged with pink); inflorescence nodding in bud; underground storage organ a spherical tuber | |
4 Leaflets (3-) 5, petiolulate, the petiolules (7-) 10-25 mm long; larger leaflets 6-15 cm long, 3.5-7 cm wide, averaging about 1.8× as long as wide, the apex acuminate; fruit bright red when ripe, smoothly elliptical in ×-section; petals light green; inflorescence erect in bud; underground storage organ an +/- elongate root, this vertical or horizontal, and sometimes branched | |
3 Inflorescence of single terminal flowers on the 1-several branches; fruit an aggregate of achenes; stem leaves 3 | |
1 Cauline leaves simple. | |
5 Inflorescence an involucrate head subtended by phyllaries, heads solitary or many, variously arrayed in secondary inflorescences; fruit a cypsela | |
5 Inflorescence various, but not as above; fruit various, not as above (sometimes the flowers tightly grouped, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.). | |
6 Fruit a 3-lobed, 3-locular capsule; inflorescence a cyathium, consisting of a single pistillate flower (reduced to a single 3-carpellate pistil) and 2 or more staminate flowers (each reduced to 1 stamen), borne in a cup-like involucre, the involucre bearing pointed or rounded glands, these sometimes brightly colored and petaloid, mimicking an individual flower (the cyathia then secondarily arranged in terminal cymes, or solitary and axillary, etc.); fresh plants with milky juice | |
6 Fruit various, not as above; inflorescence not a cyathium (and staminate or bisexual flowers almost always with > 1 stamen); fresh plants lacking milky juice. | |
7 Leaves herbaceous, thin in texture, whorls separated; pistil 1, of 2-5 fused carpels; fruit a capsule, achene, or drupe. | |
11 Corolla radially symmetrical, the petals separate; carpels 2, 3, or 5; stamens 5, 10, or many. | |
13 Inflorescence an axillary umbel; leaves narrowly linear and more than 10× as long as wide, > 20 mm long and < 2 mm wide; whorls of 3-6 leaves | |
13 Inflorescence a terminal cyme, raceme, panicle, or umbel; leaves as above, or broader in shape, narrower, or shorter; whorls of 3-16 leaves. | |
8 Larger whorled leaves on a plant > 10 mm wide. | |
18 Inflorescence of terminal involucrate clusters; perianth of 6 tepals; fruit an achene; stamens 9; [plants of very dry habitats] | |
21 Corolla pink-purple, 6-merous, the petals separate and borne on the edge of a hypanthium; stamens 8, 10, or 12; [plants of wetlands] | |
21 Corolla white, yellow, or greenish, 4-, 5-, or 7-merous, the petals fused at least basally into a tube (falling as a unit), not on a hypanthium; stamens 2, 4, 5, or 7; [plants of mesic habitats]. | |
22 Stamens 4, 5, or 7; corolla radially symmetrical. | |
23 Petals yellowish-white, with prominent green streaks; biennial or monocarpic plant, 10-30 dm tall when fertile; leaves 15-35 cm long | |
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1 Inflorescence an involucrate head subtended by phyllaries, heads solitary or many, variously arrayed in secondary inflorescences; fruit a cypsela | |
1 Inflorescence various, but not as above; fruit various, not as above. | |
2 Leaves pinnately compound. | |
3 Leaves odd-pinnate. | |
5 Plants often with a mix of mostly opposite, deeply 3-lobed leaves or trifoliate leaves (leaves sometimes simple); flowers pink or purplish-white, axillary; plants annual | |
2 Leaves palmately compound. | |
7 Plants perennial; flowers radially symmetric, showy, variously colored; plants typically not strongly resinous or glandular; inflorescences terminal |
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1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head, e.g. Pycnanthemum in LAMIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.). | |
5 Leaves herbaceous, suborbicular, about as long as wide or wider than long; calyx 3- or 4-merous; stamens 4, 8, or 12. | |
5 Leaves fleshy, linear, lanceolate, to broadly ovate, at least slightly longer than broad; calyx 5-merous; stamens 3, 5, or 10. | |
4 Perianth in 2 whorls (sepals and petals both present). | |
8 Petals connate into a tube (at least basally); inflorescence often a head or dense terminal cyme (also axillary, or solitary on long peduncles). | |
14 Leaves serrate; corolla bilaterally symmetrical (especially the flowers near the outer edge of the head); inflorescence a head | |
17 Inflorescence a cyathium, consisting of a single pistillate flower (reduced to a single 3-carpellate pistil) and 2 or more staminate flowers (each reduced to 1 stamen), borne in a cup-like involucre, the involucre bearing pointed or rounded glands, these sometimes brightly colored and petaloid, mimicking an individual flower (the cyathia then secondarily arranged in terminal cymes, or solitary and axillary, etc.); fresh plants with milky juice; fruit a 3-lobed, 3-locular capsule | |
17 Inflorescence not a cyathium (and staminate or bisexual flowers with > 1 stamen, except Callitriche in PLANTAGINACEAE); fresh plants lacking milky juice; fruit various, not as above. | |
18 Flowers 1 (or 2) in leaf axils; leaves entire. | |
19 Flowers bisexual; sepals 4; flowers with 2, 4, or 6 stamens. | |
16 Perianth in 2 whorls (sepals and petals both present). | |
27 Pistils 4-5, in a single whorl; stamens 4, 5, 8, or 10; fruit an aggregate of follicles | |
28 Petals not at all connate, not even at their bases. | |
31 Petals 4-7; sepals 4-7, normally monomorphic; stamens 4, 5, 6, 8, 10, or 12 (or sometimes rarely 2 or 3). | |
28 Petals connate at least for a short distance at their bases. | |
35 Pistils 2, united only by the style and stigma; fruit a schizocarp of 2 1-carpellate follicles (often single by abortion); plant with milky juice when fresh (except Catharanthus); leaves entire | |
35 Pistil 1 (of 2-5 fused carpels); fruit either a 2-5-carpellate capsule or of 2 or 4 1-seeded nutlets derived from 2 carpels; plant lacking milky juice; leaves entire or serrate. | |
38 Inflorescence a terminal cyme; corolla salverform, with an elongated and very narrow tube, pink or white; capsule 3-locular | |
38 Inflorescence various but not cymose, of terminal or axillary racemes or panicles, or of solitary axillary flowers; corolla connate only at the base, the petals appearing nearly separate (not salverform); capsule 1-locular | |
36 Ovary and capsule 2-carpellate; fruit either a 2-locular capsule or of 2 or 4 1-seeded nutlets derived from 2 carpels. | |
40 Capsule septicidal; corolla white, pink, blue, yellowish white, or greenish white; inflorescence either a terminal or axillary cyme, or a terminal panicle or raceme, or a terminal or axillary cyme reduced to 1 or a few flowers | |
40 Capsule loculicidal and also deeply 2-lobed; corolla white, pink, or scarlet with a yellow interior; inflorescence of cymosely arranged spikes | |
42 Fruit a schizocarp of 4 1-seeded nutlets; inflorescence terminal, of cymes; corolla ca. 5 mm long; leaves prominently 3-veined | |
42 Fruit a capsule; inflorescence axillary, of cymes or clusters (often reduced to a solitary flower); corolla > 12 mm long; leaves with single primary vein | |
41 Leaves serrate; corolla salverform, the tube narrow and nearly the same diameter for most of its length; inflorescence a terminal spike, raceme, raceme of racemes, or head. | |
34 Corolla bilaterally symmetrical (or the corolla 2-lipped but the corolla lobes twisted so as to make the flower asymmetrical); fertile stamens fewer than the corolla lobes (except Plantago in PLANTAGINACEAE, which is equal, with 4 each; a few genera have a 5th, sterile, stamen which is obviously different in form than the 4 fertile stamens) (note that many corollas are bilabiate and the number of corolla lobes, 4 or 5, may be difficult to interpret); carpels 2. | |
44 Carpels 2, each carpel slightly to deeply lobed, separating at maturity into 4 half-carpellate units (not separating in Phyla in VERBENACEAE); fruit a schizocarp of 4 mericarps (or 2 nutlets in Phyla in VERBENACEAE). | |
45 Inflorescence a thyrse, verticillaster, or terminal cyme, the flowers borne in cymose lateral branches; corolla strongly bilaterally symmetrical (rarely nearly radially symmetrical); stems square in ×-section (or sometimes rounded, especially on older growth); fresh plants often (but not always) aromatic | |
45 Inflorescence of spikes, racemes, or heads, the flowers or fruits single at nodes; corolla often nearly radially symmetrical; stems rounded in X-section (rarely square); fresh plants usually not aromatic | |
47 Corolla 4-5-lobed, either strongly bilabiate or salverform (Pseuderanthemum in ACANTHACEAE); white, blue, or yellow. | |
53 Flowers in terminal racemes, panicles, or spikes, the inflorescence not interspersed with large, leaf-like bracts. | |
54 Sepals connate for at least 0.3× their length to form a tube (this cup-like and flaring in Scrophularia in SCROPHULARIACEAE). | |
55 Inflorescence a diffuse panicle; corolla 5-11 mm long, reddish-brown (sometimes with some yellow); fruit a septicidal capsule | |
55 Inflorescence of 1 or more terminal (and sometimes upper axillary) spikes or racemes; corolla 10-50 mm long (6-8 mm long in Phryma in PHRYMACEAE), white, pink, blue, purple, or yellow; fruit either a loculicidal capsule (OROBANCHACEAE) or a single seeded achene (Phryma in PHRYMACEAE). | |
56 Corolla 6-8 mm long, white to pale pink; fruit a 1-seeded achene contained in the accrescent calyx, this “lopping down” against the inflorescence axis | |
57 Sepals separate to the base or nearly so, not forming a tube. | |
59 Leaves entire, either mostly larger or smaller [see below]; plant not blackening on drying | |
60 Leaves 3-30 cm long, narrowly elliptic to lanceolate, > 2× as long as wide; plants erect or the stems sprawling; [plants mostly of uplands] | |
57 Sepals connate for at least 0.3× their length to form a tube. | |
61 Corolla yellow, orange, or red. | |
63 Corolla red or orange, with a very narrow, cylindrical tube, the lobes then flaring into a limb about 1 cm across; plants blackening on drying; [rare exotic, in crop fields, a noxious hemiparasitic weed under quarantine] | |
63 Corolla yellow, not narrowly cylindrical, the lower lip arched; plants not blackening on drying; [rare, in seepage wetlands] | |
61 Corolla white, pale blue, lavender, or pink (sometimes with some yellow). | |
64 Corolla white, blue, or combinations of blue and white (sometimes with some yellow markings); leaves broader, mostly lanceolate; plants not blackening on drying; corolla strongly bilabiate. | |
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1 Leaf lobes very narrow, < 3 mm wide; inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
1 Leaf lobes broad, >20 mm wide; inflorescence, flower, and fruit structure various, but not with the combination of features as above. | |
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1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers tightly grouped, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.). | |
2 Flowers larger, individually conspicuous; perianth present, the petals or sepals brightly colored; fruit a capsule (or aggregate of achenes in Clematis in RANUNCULACEAE or schizocarp of 4 mericarps in Glandularia in VERBENACEAE). | |
3 Flowers radially symmetrical; stamens 5 or many; fruit a capsule or aggregate of achenes. | |
3 Flowers bilaterally symmetrical (sometimes only slightly so); stamens 4 (or 2 in Veronica in PLANTAGINACEAE); fruit a capsule or schizocarp of mericarps. | |