1 Leaves either 3-foliolate or
palmately 4-11-foliolate (all the
leaflets attached at a common point, or the
leaflets slightly
pedate in
Helleborus foetidus in RANUNCULACEAE).
2 Inflorescence , flower, and fruit structure various, but not with the combination of features as above.
3 Leaflets obviously and sharply
serrate ;
pistils 1 or 5-many; fruit an
achene , or an aggregate of
achenes ,
drupelets , or
follicles .
4 Leaflets 7-11, slightly
pedate in their arrangement, evergreen
5 Pistil 1; fruit an
achene
5 Pistils 5-many; fruit an aggregate of
achenes ,
drupelets , or
follicles
3 Leaflets entire , finely
denticulate , or very obscurely
toothed (or irregularly
serrate or
lobed in
Cardamine in BRASSICACEAE);
pistil 1; fruit simple, a
legume ,
capsule ,
silique , or
berry .
6 Corolla bilaterally symmetrical;
petals 5; fruit a
legume ;
carpel 1
6 Corolla radially symmetrical;
petals 4 or 5; fruit either an elongate
capsule or a
berry ;
carpels 1, 2, or 5.
7 Leaflets radially arranged at the summit of the
petiole , not differentiated in size or placement into a terminal
leaflet and 2
lateral leaflets ;
leaflets prominently notched at the
apex ;
petals 5, yellow;
inflorescence axillary ,
cymose or
umbelliform ;
carpels 5
7 Leaflets differentiated in size and placement into a terminal
leaflet and 2 or more
lateral leaflets ;
leaflets not regularly notched at the
apex (a few may be slightly
retuse );
petals 4, white, pink, or yellow;
inflorescence terminal and
racemose ;
carpels 1 or 2.
8 Carpels 2; fruit a red
berry ;
petals connate , purplish-blue
8 Carpel 1; fruit a
capsule ;
petals separate, white, pink, or yellow.
9 Stem leaves 1-3,
alternate [or
whorled or opposite];
leaflets 3, irregularly
serrate ,
lacerate , or additionally divided or
lobed ; fruit a
silique ;
carpels 2
9 Stem leaves >3,
alternate ;
leaflets (1-) 3-7, each
entire or obscurely
toothed ; fruit a
capsule ;
carpel 1
1 Leaves either 1-
pinnately compound (all
leaflets attached to a central
rachis ) or more complexly compound (with several orders of branching, some
leaflets at least attached to second-order branches from the
rachis ).
10 Inflorescence an
involucrate head subtended by
phyllaries , heads solitary or many, variously arrayed in secondary
inflorescences ; fruit a
cypsela ;
ovary inferior
10 Inflorescence various, but not as above; fruit various, not as above;
ovary superior .
11 Leaves 1-
pinnately compound (all
leaflets attached to a central
rachis ).
12 Flowers bilaterally symmetrical,
papilionaceous ; fruit a
legume ; leaves even-
pinnately or odd-
pinnately compound , the terminal
leaflet sometimes replaced by a
tendril ;
leaflets entire or at most minutely
denticulate
12 Flowers radially symmetrical (or barely bilaterally symmetrical in
Erodium in GERANIACEAE); fruit a
capsule ,
capsular but of 5
mericarps , or an aggregate of
achenes ,
nutlets , or
follicles (in some cases the # of
pistils from many down to 2 or even 1); leaves odd-
pinnately compound , never with
tendrils ;
leaflets serrate (or
entire to
shallowly lobed in
Polemonium in POLEMONIACEAE,
Cardamine in BRASSICACEAE, and
Floerkea in LIMNANTHACEAE).
13 Pistils many (only 1-2 in
Agrimonia ,
Poteridium ,
Poterium , and
Sanguisorba ); fruit an aggregate of
achenes ,
nutlets , or
follicles ;
hypanthium present;
stamens 5-many (only 4 in
Poteridium and
Sanguisorba )
13 Pistil 1 (or deeply 2-3-
lobed in
Floerkea in LIMNANTHACEAE); fruit a
silique ,
capsule ,
schizocarp of 2-3
mericarps , or a
capsular schizocarp of 5
mericarps (
Erodium in GERANIACEAE);
hypanthium absent;
stamens 3-6.
15 Petals 5, fused (distinct in
Erodium in GERANIACEAE);
stamens 5;
inflorescence axillary or terminal,
cymose , consisting of subcapitate,
umbel -like, or
helicoid cymes ; fruit either a
capsule , or a
capsular schizocarp of 5
mericarps (
Erodium in GERANIACEAE).
16 Flowers slightly bilaterally symmetrical (2 of the
petals of different size than the other 3); fruit a
capsular schizocarp of 5
mericarps ;
carpels 5
16 Flowers radially symmetrical; fruit either a
loculicidal capsule or a
berry ;
carpels 2 or 3.
17 Fruit a
berry ; fresh plant rankly fragrant
17 Fruit a
capsule ; fresh plant not aromatic.
18 Capsule 2-
valvate ;
carpels 2;
leaflets prominently
serrate or with some
tooth -like sublobes
11 Leaves more complexly compound (with 2 or more orders of branching, some
leaflets at least attached to second-order branches from the
rachis ).
19 Leaves 2×
even-pinnate ; flowers in
spikes or
spherical heads; fruit a
legume or
loment .
19 Leaves either 2×
odd-pinnate or more complexly 2-4×
ternately or
ternately -
pinnately compound ; flowers in various
inflorescences ; fruits various (not
legumes or
loments ).
20 Leaf
segments linear , less than 2 mm wide.
21 Inflorescence either a terminal solitary flower or terminal
raceme or
panicle ;
ovary superior , either of 2 fused
carpels or of 1-5or many distinct 1-
carpellate pistils ; fruit either a
capsule or an aggregate of
follicles or
achenes .
22 Ovary of 2 fused
carpels ; fruit a
capsule (1-seeded and
indehiscent in
Fumaria )
22 Ovary of 1-5or many distinct 1-
carpellate pistils ; fruit an aggregate of
follicles or
achenes
20 Leaf
segments broader,
lanceolate ,
ovate , or
elliptic , > 5 mm wide.
23 Erect or
sprawling herb;
stamens 5-6 or >15.
24 Leaflets sharply
serrate , with usually many teeth on each
leaflet side, the total number of “points” per
leaflet > 10.
26 Flowers
bisexual ;
carpels 1-8; fruit an aggregate of
follicles , a single
follicle , or an
indehiscent berry -like fruit
26 Flowers
unisexual ;
carpels (in
pistillate flowers) of 3 (-5)
carpels ; fruit an aggregate of
follicles
24 Leaflets entire , or with 1-several, broad,
obtuse , rounded, or broadly
acute “sublobes”, especially towards the tip of the
leaflet , the total number of “points” per
leaflet < 10.
28 Perianth bilaterally symmetrical, the
corolla of 4 fused
petals ; plant a
scandent vine or erect or
sprawling herb
28 Perianth radially symmetrical, of 1-5
whorls of separate
perianth parts; plant an erect herb.
29 Perianth of 4-5
whorls of 3 parts each (some of the
whorls modified into
nectaries );
pistil 1, 1-
carpellate ; fruit a drupelike, blue,
naked seed; largest
leaflets > 6 cm long, obviously longer than wide
29 Perianth of 1
whorl ; of 4-5 parts;
pistils many, each 1-
carpellate ; fruit an aggregate of
achenes or an aggregate of
follicles ; largest
leaflets either < 6 cm long, or if longer than 6 cm, also about as wide as long
1 Cypselae (at least of the inner
florets of the head) beaked, the
beak usually > ½ the length of the
cypselae body.
2 Heads solitary and terminal at the end of a stem unbranched to its
base .
3 Flower color pink to purple or white
3 Flower color yellow to yellowish-white.
4 Leaves
basal and
cauline , grasslike (untoothed and
unlobed ); stem leafy
4 Leaves
basal only, variously
toothed to
pinnately lobed ; stem
scapiform (leafless).
5 Pappus either of outer
scales and inner
bristles or entirely of
aristate scales .
6 Pappus of outer
scales and inner
plumose bristles ; leaves
oblanceolate to
oblong ; plants annual or perennial
6 Pappus of
aristate scales ; leaves
linear to narrowly
lanceolate ; plants annual
2 Heads several per stem, in various
corymbiform ,
umbelliform ,
spiciform , or
paniculiform arrays (rarely solitary and terminal in the smallest and most
depauperate individuals in a population.
7 Achenes distinctly flattened or
compressed .
8 Florets (6-)10-50+ per head
9 Plants taprooted annuals or
biennials , without
rhizomes
9 Plants
rhizomatous perennials, the
rhizomes deeply rooted in the substrate
10 Pappus of
plumose bristles , at least the inner series; plant an annual or
biennial .
10 Pappus of simple
capillary bristles ; plant an annual,
biennial , or perennial.
12 Beak of the
cypsela with a ring of soft white
reflexed hairs at the summit (just below the
pappus );
phyllary apices usually with
keel -like flap
12 Beak of the
cypsela lacking a ring of
hairs as described;
phyllary apices lacking
keel (
phyllary bases can be
keeled in
Crepis )
13 Pappus of 80-150
barbellulate bristles ; plant an annual or
biennial
13 Pappus either of 40-50 (or more) smooth
bristles or of 20-30
barbellulate bristles ; plant a perennial.
14 Phyllaries tomentose ;
pappus of 40-50 (or more) smooth
bristles
1 Cypselae beakless, the
apex typically rounded or
truncate (sometimes tapered but lacking a distinct
beak and conspicuously < ½ the length of
cypselae body).
15 Leaves basally disposed (stem leaves few or none, if present generally smaller in size than the
basal leaves, which are
persistent into flowering and fruiting);
corollas yellow, orange, or red.
17 Heads solitary or 2-3;
pappus absent, the
cypselae never bearing
scales or
bristles ;
cypselae with 8-10
ribs (3-5 prominent); [
waif , ne. US]
17 Heads solitary;
pappus present (occasionally absent in
Krigia );
cypselae with 10-20
ribs ; [widespread natives]
18 Pappus of 40-80 smooth to
barbellulate bristles and sometimes a few
subulate to
setiform scales (but
pappus never absent); [Great Plains, midwest US]
18 Pappus absent, of
scales , or of 5-45
bristles and 5+
scales ; [collectively widespread]
16 Pappus of
bristles only (these
barbellulate ,
plumose , or rarely smooth).
19 Pappus of
plumose bristles (these somewhat flattened at their
bases )
20 Leaves with
entire margins ; plants perennials, from short
rhizomes , a short
caudex with
fibrous roots , or a thick
taproot .
20 Leaves coarsely
toothed or
pinnately lobed ; plants annuals, from a
taproot .
22 Flowers usually pink to lavender; [TX; tribe Mutisieae]
22 Flowers usually yellow to orange; [widespread, including TX; tribe Cichorieae].
23 Involucre 5-12 mm high;
achenes usually > 2.5 mm long;
pappus bristles distinct, 3-7 mm long
23 Involucre 3-5 mm high;
achenes 1.5-2.5 mm long;
pappus bristles basally
connate (unified basally into a ring-like structure at the junction of
achene summit), 2.5-3.5 mm long
15 Leaves
basal and
cauline (plant often beginning with a
basal rosette , but by flowering bearing well-developed stem leaves about as large as the
basal leaves, the
basal rosette often withering prior to flowering and fruiting);
corollas yellow, orange, red, blue, pink, white, or lavender.
24 Pappus absent or of
scales .
25 Corollas pale blue (rarely pink or white)
25 Corollas yellow (rarely orange).
26 Phyllaries enfolding the outer
cypselae
26 Phyllaries not enfolding the outer
cypselae .
27 Stem
winged and
spiny ; leaves
spiny -margined;
receptacle with
paleae
27 Stem not
winged or
spiny ; leaves not
spiny -margined;
receptacle lacking
paleae .
28 Cypselae 1.2-2.8 mm long; heads borne single at the ends of
scapiform stems that are unbranched (rarely few-branched near the
base ); plants to 7 dm tall
28 Cypselae 3-5 mm long; heads borne in
corymbiform or
thyrsiform arrays; plants to 15 dm tall
24 Pappus of numerous smooth,
barbellate , or
plumose bristles .
29 Cypselae more or less strongly flattened.
30 Pappus of 30-45+
bristles , these at least partly
plumose
29 Cypselae terete or
prismatic , slightly or not at all flattened.
31 Corollas pink, purple, lavender, white, or creamy-yellow.
32 Leaves
linear ,
entire , < 3 mm wide, or reduced to
scales ;
cypselae 11-14 mm long; [of the Coastal Plain of GA and se. AL south to s. FL]
32 Leaves broader, of various shapes, usually
hastate , irregularly
lobed , and/or
serrate ;
cypselae 3.5-10 mm long; [collectively widespread in our area, south to n. FL]
31 Corollas bright yellow, orange, or red.
33 Leaves finely
prickly ;
pappus of frizzy outer
bristles and coarse, straight inner
bristles
33 Leaves not finely
prickly ;
pappus of
barbellulate bristles (sometimes of uneven thickness but not of 2 distinct types)
34 Plants taprooted annuals and
biennials (rarely perennials);
pappus bristles white and soft in texture
34 Plants
fibrous -rooted perennials;
pappus bristles white, light to medium tan, or sordid, stiff.
35 Cypselae (2-) 2.5-7 mm long;
pappus of (30-) 40-80 white, tan, or sordid
bristles , in 1-2+ series; plants
cespitose ;
corollas yellow
35 Cypselae 1-2.5 mm long;
pappus of 25-40+ white to sordid
bristles , in 1 series; plants
stoloniferous (
cespitose in a few species);
corollas yellow or orange