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Key to Lamiaceae

Lamiaceae

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1 Fruit a fleshy drupe; plant a small tree, shrub, or sprawling vine; mature stems terete or obscurely 4-sided (by secondary growth).
  2 Flowers zygomorphic; palmately (3-) 5-7 (-9) foliolate or simple (V. rotundifolia); [subfamily Viticoideae]
  2 Flowers essentially actinomorphic; leaves simple.
    3 Stems pubescent with dendritic hairs; inflorescence axillary; calyx 0.5-2 mm, lobes diminutive to nearly obsolete
    3 Stems glabrous or pubescent with simple (unbranched) hairs; inflorescence terminal (rarely only axillary); calyx lobes conspicuous or diminuitive.
      4 Corolla 5-40 mm long; calyx 5-18 mm, the lobes conspicuous, the calyx length often surpassing the fruit when mature; [subfamily Ajugoideae]
      4 Corolla 2-5 mm long; the calyx lobes inconspicuous and not surpassing the mature fruit; [subfamily Premnoideae]
1 Fruit a schizocarp of 4 dry mericarps; plant either an herb or a shrub to 5 (-20) dm tall; mature stems usually distinctly 4-sided (sometimes terete or obscurely 4-sided).
        5 Calyx with either a distinctly enlarged protuberance on the upper surface, or the upper lobe expanded and “cap-like”.
          6 Calyx with 5 lobes, clearly separate apically, with the upper lobe expanded and “cap-like”
          6 Calyx with 2 entire lobes joined at the margins, rounded apically, with a distinct protuberance on the upper surface
        5 Calyx without an enlarged protuberance or “cap-like” upper lobe.
             7 Upper lip of corolla greatly reduced or lobes laterally disposed, thus the corolla appearing to consist of one large lower lip; [subfamily Ajugoideae].
               8 Lower lip with 2-4 lobes; flowers yellow or deep blue-purple; plants stoloniferous
               8 Lower lip appearing 5-lobed (proximal 2 lateral, erect lobes represent the cryptic upper lip); flowers white to pink; plants cespitose
             7 Upper lip of corolla conspicuous, flaring or galeate.
                 9 Plants distinctly repent and rooting at the nodes, or producing elongate stolons.
                     11 Herbs; leaves cordate-reniform, coarsely crenate, the blade > 1 cm long; inflorescence of axillary cymules; corollas 10-20 mm
                     11 Subshrubs; leaves ovate to elliptic, entire, the blade < 1 cm long; inflorescence a terminal thyrse; corollas 4-5 mm
                              15 Calyx with 3-4 prominent lobes (rarely 5, if one includes small teeth).
                                  17 Calyx with 4 unequal primary lobes; flowers large, 2.5-3.5 cm long, in bracteate terminal racemes
                                  17 Calyx with 3 primary lobes (upper lobe occasionally with 3 apiculate teeth, e.g. Salvia lyrata); flowers 3 cm or less, in terminal thyrses.
                                    18 Stamens 4; calyx enveloped and partially concealed by subtending bracts
                                       19 Herbs; calyx with simple trichomes; leaves non-revolute, margins various, membranaceous
                              15 Calyx usually with 5 prominent lobes (except for Clinopodium with rarely fused upper lobes).
                                                            29 Stamens ascending under the upper corolla lip, either included within the tube (or at least not clearly exserted beyond it).
                                                                       34 Stems often with swollen nodes (areas just below appear dark and sunken upon drying, except G. ladanum); hairs of the stem either exclusively hispid or short recurved and mixed with longer glandular trichomes; corolla 15-28 mm long
                                                                       34 Stems without swollen nodes; hairs of the stem finely and softly retrorse, generally lacking glandular hairs (though sessile glands may be present); corolla 5-14 mm long.

Key N2: herbaceous dicots with mainly basal, simple leaves

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1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head subtended by bracts, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or petals separate, or fruit a schizocarp of mericarps, etc.).
  2 Basal leaves 2-lobed, pinnately lobed, or palmately lobed (not considering cordate, hastate, or auriculate leaf bases as “lobed”).
    3 Basal leaves 2-lobed, hinged between the lobes, each lobe with stiff, marginal, eyelash-like bristles; [Coastal Plain of NC and SC, rarely planted and weakly naturalized elsewhere]
    3 Basal leaves 3-many-lobed, palmately or pinnately; [collectively widespread].
      4 Leaf lobing pinnate.
        5 Gynoecium of separate pistils (each with a single carpel); fruit an aggregate
        5 Gynoecium of a single pistil (with 2, rarely more, carpels); fruit simple.
          6 Stamens many; sepals 2, petals 4; fresh plants with yellow, orange, or white milky juice
          6 Stamens 4, 5, or 6; sepals 4 or 5; petals 4 or 5.
             7 Petals 4, distinct; stamens 6
             7 Petals 5, fused; stamens 2, 4, or 5.
               8 Corolla radially symmetrical; stamens 5
               8 Corolla 2-lipped, bilaterally symmetrical or asymmetrical; stamens 2 or 4.
                 9 Corolla lobes not twisted, the flower bilaterally symmetrical; stamens 2
                 9 Corolla lobes twisted so as to make the flower asymmetrical; stamens 4
      4 Leaf lobing palmate.
                          13 Petals 5; stamens 5; fruit a schizocarp of 2 mericarps.
                            14 Fruit smooth; leaves with 5 or more lobes
                              15 Gynoecium of separate pistils (each with a single carpel); fruit an aggregate.
                                  17 Carpels partly fused, arrayed in a ring of 10-20
                                16 Perianth of a single whorl of 3-12 petaloid sepals (the petals absent or small and rudimentary).
                                    18 Leaves 2, the single flower terminal and associated with the upper leaf; fruit an aggregate of berries
                                    18 Leaves normally > 2, flowers not as above; fruit an aggregate of achenes, utricles, or follicles
                                         20 Petals distinct; leaves with rounded lobes or teeth.
                                           21 Corolla radially symmetrical; petals 8-12; capsule fusiform, narrowed to both ends, > 5× as long as wide
  2 Basal leaves not lobed, at most serrate or crenate (and sometimes also cordate, hastate, auriculate, or peltate at the base).
                                                 24 Inflorescence either a terminal spike, or a 1-7-flowered terminal cyme, or of a solitary axillary or terminal flower; fruit various; perianth biseriate (of differentiated sepals and petals (except uniseriate, of 3 fused sepals in ARISTOLOCHIACEAE).
                                                      26 Flowers bilaterally symmetrical; inflorescence a terminal spike (with > 20 flowers); petals 4, usually scarious, transparent; sepals 4, green; stamens 4
                                                      26 Flowers radially symmetrical; inflorescence either of a solitary flower or of a 1-7-flowered terminal cyme; petals 5, 8-12, or 0; sepals 5 (green), 3 (brown), or 5-9 (yellow); stamens 5, 12, or many.
                                                        27 Gynoecium of separate pistils (each with a single carpel); fruit an aggregate of achenes or follicles; flowers bright yellow, either of 5-9 distinct petaloid sepals, or of 8-12 distinct petals subtended by 3-4 green distinct sepals
                                                        27 Gynoecium either of a single pistil with 6 carpels or of a single pistil with 4 carpels or of 2 nearly separate carpels; fruit a simple capsule (or deeply 2-lobed); flowers white, brown, or greenish, either of 5 fused or distinct white petals and 5 fused or distinct green sepals, or of 3 fused brown or greenish petaloid sepals.
                                                          28 Flowers brown or green, of 3 fused brown or greenish petaloid sepals (and 0 petals); carpels 6; stamens 12; leaves 4-10 cm wide
                                                          28 Flowers white, of 5 white or whitish petals and 5 green sepals; carpels 2; stamens 5; leaves 1-12 (-15) cm wide
                                                              30 Fruit a deeply 2-lobed capsule; sepals longer or ca. as long as petals; petals not undulate, fused at their bases or distal ½, the anthers maroon or brown-colored; [common, widespread in our area]
                                                                 31 Gynoecium of separate pistils (each with a single carpel); fruit an aggregate; perianth of 5 green sepals and 5 colored petals (or of 2 or 4 (-5) white-colored tepals in Begonia).
                                                                   32 Flowers typically with 2 or 4 (-5) white-colored tepals; leaf bases conspicuously oblique (sometimes variegated); fruit unequally or subequally 3-winged capsules; [ornamental waifs or uncommon non-natives]
                                                                   32 Flowers with 5 green sepals and 5 colored petals (not merged into tepals); leaf bases oblique or not; fruit various but not 3-winged capsules; [natives and non-natives].
                                                                              37 Inflorescence a terminal raceme; perianth of 4 green sepals and 4 white petals; fruit a silique/silicle; fresh foliage in spring and summer with a strong garlic odor; larger leaves < 10 cm in diameter
                                                                              37 Inflorescence a terminal panicle; perianth of 6 cream-colored petaloid sepals; fruit a winged achene; fresh foliage lacking a garlic odor; larger leaves typically > 30 cm in diameter
                                                                                    40 Leaves tubular, with a sutured ventral flange, erect or reclining, adapted as a pitfall for insects (flat, phyllodial leaves sometimes present as well, common in the winter in some species, such as S. oreophila)
                                                                                      41 Stem leaves opposite; perianth 5-merous, at least the corolla bilaterally symmetrical (barely so in VALERIANACEAE), or the parts curved so as to be asymmetrical (Pedicularis in OROBANCHACEAE); stamens 2, 3, or 4.
                                                                                           43 Corolla distinctly 2-lipped (with prominently large upper and lower corolla lobes) or hooded (the upper lip hood-like), distinctly bilaterally symmetrical, or the lobes twisted so as to make the corolla asymmetrical.
                                                                                      41 Stem leaves alternate; perianth radially symmetrical (less commonly bilaterally symmetrical); stamens 5, 6-8, 9, 10 (rarely 4).