1 Leaves either 2-3-foliolate or
palmately 4-11-foliolate (all the
leaflets attached at a common point).
2 Leaves 2-foliolate; fruit a
capsule , opening by a
circumscissile lid
2 Leaves either 3-foliolate or
palmately or pedately 4-11-foliolate.
3 Inflorescence an
involucrate head subtended by
phyllaries , the heads solitary or many and variously arrayed in secondary
inflorescences , the
ovary inferior , the
corolla connate and
tubular at least basally, the
calyx absent, the
stamens 5, the fruit a
cypsela
3 Inflorescence , flower, and fruit structure various, but not with the combination of features as above.
4 Inflorescence various, usually not an
umbel (sometimes an
umbel in
Oxalis in OXALIDACEAE);
ovary superior ; fruit an aggregate,
legume ,
berry , or 2-valved
capsule .
5 Leaflets either
entire or barely and very shallowly
crenulate or notched at the tip (but otherwise
entire ).
6 Inflorescence a
spadix , surrounded by a
spathe ; fruit a
berry ; [
Monocots {illogically keyed here because of the likelihood of being mistaken for a
dicot }]
7 Flowers bilaterally symmetrical; fruit a
legume ; [plant of uplands]
7 Flowers radially symmetrical; fruit a 2-valved or 5-valved
capsule ; [plant of uplands or wetlands]
8 Leaflets not notched at the tip; flowers white; [plants of
saturated saturated or ponded wetlands]
8 Leaflets notched at the tip; flowers pink, white, or yellow; [plants of uplands or
temporarily flooded wetlands]
9 Petals 5 or more;
stamens 10 or more; fruit either a
legume or an aggregate of
achenes or
follicles
10 Stamens many, fused into a
staminal tube;
carpels 10-20, in a ring;
pubescence stellate (sometimes mixed with simple
hairs )
10 Stamens 10-many, separate, or fused but not all into a
staminal tube;
carpel either 1 (FABACEAE), or 3-7 in a ring (RANUNCULACEAE), or many and
spirally arranged on a
conical receptacle (RANUNCULACEAE or ROSACEAE)
11 Leaflets serrulate ; flowers bilaterally symmetrical; fruit a
legume ;
corolla variously colored, including white
11 Leaflets serrate ; flowers radially symmetrical; fruit an aggregate of
achenes or of
follicles ;
corolla white or yellowish or greenish.
12 Fruit an aggregate of
follicles
12 Fruit an aggregate of
achenes (borne on a fleshy, expanded
receptacle in
Fragaria and some
Potentilla )
13 Flowers lacking a
hypanthium
13 Flowers with a
hypanthium
1 Leaves 1-
pinnately compound (all
leaflets attached to a central
rachis ) or more complexly compound (with several orders of branching, some
leaflets at least attached to second-order branches from the
rachis ).
14 Leaves 1-
pinnately compound (all
leaflets attached to a central
rachis ).
15 Inflorescence an
involucrate head subtended by
phyllaries , the heads solitary or many and variously arrayed in secondary
inflorescences , the
ovary inferior , the
corolla connate and
tubular at least basally, the
calyx absent, the
stamens 5, the fruit a
cypsela
15 Inflorescence , flower, and fruit structure various, but not with the combination of features as above.
16 Flowers bilaterally symmetrical; fruit a
legume
16 Flowers radially symmetrical; fruit a
silique /
silicle , or a
schizocarp of
mericarps , or an
achene .
17 Petals 0 or 5 (if 0, the
sepals petaloid );
stamens 2, 4, 5, or many.
18 Stamens 2, 4, or many; fruit an
achene
14 Leaves more complexly compound (with 2 or more orders of branching, some
leaflets at least attached to second-order branches from the
rachis ).
19 Inflorescence an
involucrate head subtended by
phyllaries , the heads solitary or many and variously arrayed in secondary
inflorescences , the
ovary inferior , the
corolla connate and
tubular at least basally, the
calyx absent, the
stamens 5, the fruit a
cypsela
19 Inflorescence , flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head subtended by
bracts , but then with other features differing, such as
stamens 4, or green
calyx present, or
petals separate, or fruit a
schizocarp of
mericarps , etc.).
20 Leaf
segments or ultimate
lobes linear or
lanceolate , > 2× as long as wide, < 4 mm wide.
21 Inflorescence various, but not an
umbel ;
ovary superior ; fruit an aggregate of
follicles or an elongate
capsule .
22 Carpels 2, fused; fruit an elongate
capsule ; flowers bilaterally symmetrical
22 Carpels 5-10 or many, separate; fruit an aggregate; flower radially symmetrical
20 Leaf
segments or ultimate
lobes ovate or
elliptic , < 3× as long as wide, > 5 mm wide.
24 Fruit a 5-seeded
drupe
23 Inflorescence various, but not an
umbel ; fruit an aggregate of
follicles or
achenes , an elongate
capsule , or a
naked seed resembling a
drupe .
25 Leaflets with < 10 ultimate ‘points’ (
lobe or
tooth terminations), these rounded to broadly
acute , often large in comparison to the
leaflet and appearing as “sublobes”;
pistil 1 or 4-many.
26 Corolla bilaterally symmetrical; fruit an elongate
capsule ; [cultivated
exotic , rarely
persistent near gardens]
26 Corolla radially symmetrical; fruit an aggregate of
follicles or
achenes , or a
naked seed resembling a
drupe ; [native plants of moist to dry forests and rock outcrops].
27 Leaflets 5-8 cm long, obviously longer than broad;
pistil 1; fruit a
naked blue seed resembling a
drupe ; flowers mainly 3
-merous
27 Leaflets 1-6 cm long, about as long as broad if > 4 cm long;
pistils 4-many; fruit an aggregate of
follicles or
achenes ; flowers mainly 4-5
-merous
25 Leaflets with >11 ultimate ‘points’ (
lobe or
tooth terminations), these
acuminate to
acute ;
pistils 1-8.
28 Pubescence of the stem and lower leaf surface
glandular ; flowers
unisexual , on the same plant (
monoecious );
stamens 10;
pistils 2, partly fused; fruit an aggregate of
follicles
28 Pubescence of the stem and lower leaf surface non-
glandular (or absent); flowers either
bisexual (the plants
hermaphroditic ), or
unisexual and the male and female flowers on separate plants (the plants
dioecious );
stamens 15 or more;
pistils 1-8, separate; fruit an aggregate of
follicles , a
follicle , or a red or white
berry .
29 Flowers
bisexual (plants
hermaphroditic );
carpels 1-8 per flower;
inflorescence a
raceme , or a
panicle of
racemes with just a few branches; fruit an aggregate of
follicles , a
follicle , or a red or white
berry
29 Flowers
unisexual (plants
dioecious );
carpels 3-4 per
pistillate flower;
inflorescence a
panicle of
racemes , with numerous branches; fruit an aggregate of
follicles
1 Inflorescence an
involucrate head subtended by
phyllaries , the heads solitary or many and variously arrayed in secondary
inflorescences , the
ovary inferior , the
corolla connate and
tubular at least basally, the
calyx absent, the
stamens 5, the fruit a
cypsela
1 Inflorescence , flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head subtended by
bracts , e.g.
Eryngium in APIACEAE, but then with other features differing, such as
stamens 4, or green
calyx present, or
petals separate, or fruit a
schizocarp of
mericarps , etc.).
3 Basal leaves 2-
lobed , hinged between the
lobes , each
lobe with stiff,
marginal , eyelash-like
bristles ; [Coastal Plain of NC and SC, rarely planted and weakly naturalized elsewhere]
3 Basal leaves 3-many-
lobed ,
palmately or
pinnately ; [collectively widespread].
5 Gynoecium of separate
pistils (each with a single
carpel ); fruit an aggregate
5 Gynoecium of a single
pistil (with 2, rarely more,
carpels ); fruit simple.
6 Stamens many;
sepals 2,
petals 4; fresh plants with yellow, orange, or white milky juice
6 Stamens 4, 5, or 6;
sepals 4 or 5;
petals 4 or 5.
7 Petals 4, distinct;
stamens 6
7 Petals 5, fused;
stamens 2, 4, or 5.
8 Corolla radially symmetrical;
stamens 5
8 Corolla 2-lipped, bilaterally symmetrical or asymmetrical;
stamens 2 or 4.
9 Corolla lobes not twisted, the flower bilaterally symmetrical;
stamens 2
9 Corolla lobes twisted so as to make the flower asymmetrical;
stamens 4
10 Petiole attachment
peltate .
11 Leaves < 10 cm in diameter
11 Leaves > 15 cm in diameter
10 Petiole attachment
marginal .
14 Fruit
tuberculate ; leaves 3-
lobed
14 Fruit smooth; leaves with 5 or more
lobes
12 Ovary superior , or
half-inferior by
fusion of a
hypanthium a part of the way up the
ovary wall.
15 Gynoecium of separate
pistils (each with a single
carpel ); fruit an aggregate.
16 Perianth of 5 green
sepals and 5 colored
petals .
17 Carpels partly fused, arrayed in a ring of 10-20
17 Carpels separate,
spiral , many
16 Perianth of a single
whorl of 3-12
petaloid sepals (the
petals absent or small and
rudimentary ).
18 Leaves 2, the single flower terminal and associated with the upper leaf; fruit an aggregate of
berries
18 Leaves normally > 2, flowers not as above; fruit an aggregate of
achenes ,
utricles , or
follicles
15 Gynoecium of a single
pistil (with 1-5
carpels ); fruit simple.
19 Hypanthium present, partially fused or not fused to the
pistil ;
ovary partially
inferior to
superior
20 Petals connate at
base ; leaves sharply
toothed
20 Petals distinct; leaves with rounded
lobes or teeth.
21 Corolla radially symmetrical;
petals 8-12;
capsule fusiform , narrowed to both ends, > 5× as long as wide
21 Corolla bilaterally symmetrical;
petals 5;
capsule ovoid , < 2× as long as wide
2 Basal leaves not
lobed , at most
serrate or
crenate (and sometimes also
cordate ,
hastate ,
auriculate , or
peltate at the
base ).
24 Inflorescence a terminal and/or
axillary raceme ,
panicle , or
cyme of many small flowers; fruit an
achene ;
perianth uniseriate , of 0, 4-5, or 6
tepals .
25 Flowers
unisexual ;
staminate flowers 4-5
tepals ,
pistillate flowers lacking a
perianth
25 Flowers
bisexual ; flowers with 6
tepals
24 Inflorescence either a terminal
spike , or a 1-7-flowered terminal
cyme , or of a solitary
axillary or terminal flower; fruit various;
perianth biseriate (of differentiated
sepals and
petals (except
uniseriate , of 3 fused
sepals in ARISTOLOCHIACEAE).
26 Flowers bilaterally symmetrical;
inflorescence a terminal
spike (with > 20 flowers);
petals 4, usually
scarious , transparent;
sepals 4, green;
stamens 4
26 Flowers radially symmetrical;
inflorescence either of a solitary flower or of a 1-7-flowered terminal
cyme ;
petals 5, 8-12, or 0;
sepals 5 (green), 3 (brown), or 5-9 (yellow);
stamens 5, 12, or many.
27 Gynoecium of separate
pistils (each with a single
carpel ); fruit an aggregate of
achenes or
follicles ; flowers bright yellow, either of 5-9 distinct
petaloid sepals , or of 8-12 distinct
petals subtended by 3-4 green distinct
sepals
27 Gynoecium either of a single
pistil with 6
carpels or of a single
pistil with 4
carpels or of 2 nearly separate
carpels ; fruit a simple
capsule (or deeply 2-
lobed ); flowers white, brown, or greenish, either of 5 fused or distinct white
petals and 5 fused or distinct green
sepals , or of 3 fused brown or greenish
petaloid sepals .
28 Flowers brown or green, of 3 fused brown or greenish
petaloid sepals (and 0
petals );
carpels 6;
stamens 12; leaves 4-10 cm wide
28 Flowers white, of 5 white or whitish
petals and 5 green
sepals ;
carpels 2;
stamens 5; leaves 1-12 (-15) cm wide
29 Petals separate;
sepals separate; plant
glabrous
29 Petals fused (at least basally in
Dichondra );
sepals fused;
carpels 2; plant
pubescent .
30 Fruit a deeply 2-
lobed capsule ;
sepals longer or ca. as long as
petals ;
petals not
undulate , fused at their
bases or
distal ½, the
anthers maroon or brown-colored; [common, widespread in our area]
30 Fruit a fleshy
berry ;
sepals shorter than the
petals ;
petals undulate , fused, the
anthers yellow; [rare
exotic ]
31 Gynoecium of separate
pistils (each with a single
carpel ); fruit an aggregate;
perianth of 5 green
sepals and 5 colored
petals (or of 2 or 4 (-5) white-colored
tepals in
Begonia ).
32 Flowers typically with 2 or 4 (-5) white-colored
tepals ; leaf
bases conspicuously
oblique (sometimes
variegated ); fruit unequally or subequally 3-
winged capsules ; [ornamental
waifs or uncommon non-natives]
32 Flowers with 5 green
sepals and 5 colored
petals (not merged into
tepals ); leaf
bases oblique or not; fruit various but not 3-
winged capsules ; [natives and non-natives].
33 Carpels 10-20, partly fused, arrayed in a ring;
petals white, pink, red, or purplish
33 Carpels many, separate,
spiral ;
petals yellow or white.
34 Flowers lacking a
hypanthium ; fruit an aggregate of
achenes or aggregate of
follicles
34 Flowers with a
hypanthium ; fruit an aggregate of
drupelets or aggregate of
achenes
31 Gynoecium of a single
pistil (with 1-5
carpels ); fruit simple.
35 Flowers bilaterally symmetrical;
inflorescence of a solitary flower; fruit a 3-
locular capsule
35 Flowers radially symmetrical;
inflorescence an
umbel (or
composite of
umbelliform units, or a terminal
panicle .
37 Inflorescence a terminal
raceme ;
perianth of 4 green
sepals and 4 white
petals ; fruit a
silique /
silicle ; fresh foliage in spring and summer with a strong garlic odor; larger leaves < 10 cm in diameter
37 Inflorescence a terminal
panicle ;
perianth of 6 cream-colored
petaloid sepals ; fruit a
winged achene ; fresh foliage lacking a garlic odor; larger leaves typically > 30 cm in diameter
38 Petiole attachment
peltate
39 Leaf
blades longer than wide, sharply V-
cleft at the
base and otherwise shallowly
denticulate
39 Leaf
blades as broad or broader than long,
cleft at the
base and also irregularly
serrate or
crenate around the
margin
22 Basal leaves
petiolate or not, with a
truncate , rounded, or
cuneate leaf
base .
40 Leaves
tubular , with a sutured
ventral flange , erect or
reclining , adapted as a pitfall for insects (flat,
phyllodial leaves sometimes present as well, common in the winter in some species, such as
S. oreophila )
40 Leaves flat, not sutured into a
tubular shape.
41 Stem leaves opposite;
perianth 5
-merous , at least the
corolla bilaterally symmetrical (barely so in VALERIANACEAE), or the parts curved so as to be asymmetrical (
Pedicularis in OROBANCHACEAE);
stamens 2, 3, or 4.
43 Corolla narrowly
tubular , the five
lobes flaring at nearly 90 degrees and nearly radially symmetrical
43 Corolla distinctly 2-lipped (with prominently large upper and lower
corolla lobes ) or hooded (the upper
lip hood -like), distinctly bilaterally symmetrical, or the
lobes twisted so as to make the
corolla asymmetrical.
44 Corolla yellow, the upper
lip often slightly to strongly maroon, hooded but the
corolla lobes twisted so as to make the flower asymmetrical
44 Corolla white, lavender, or blue, 2-lipped and bilaterally symmetrical.
45 Sepals separate to the
base or nearly so, not forming a tube
45 Sepals connate for at least 0.3× their length to form a tube
41 Stem leaves
alternate ;
perianth radially symmetrical (less commonly bilaterally symmetrical);
stamens 5, 6-8, 9, 10 (rarely 4).
49 Pistils many, each with a single
carpel ; fruit an aggregate of
achenes
50 Corolla bilaterally symmetrical (barely so in
Limosella in SCROPHULARIACEAE);
stamens 2, 4, 6, 8, or 10.
52 Petals separate;
stamens 10
52 Petals fused;
stamens 10 or 6-8.
55 [plants of coastal wetlands]
55 [plants of uplands or inland
seeps or
fens ]
56 Flowers (and subtending
bracts ) red or yellow
56 Flowers purple, blue, or lavender
50 Corolla radially symmetrical;
stamens 5, 10, 4-6, or 9.
57 Perianth of green
sepals and more brightly colored
petals ;
stamens 5 or 10;
carpels 2, 3, 4, or 5.
58 Leaves covered with sticky,
gland -tipped
hairs (often red), as flypaper traps for insects
58 Leaves lacking sticky
gland -tipped
hairs .
59 Fruit a
schizocarp of 4
nutlets (
ovary obviously 4-
lobed in flower)
59 Fruit a
capsule or
silique /
silicle (or
utricle in
Limonium in PLUMBAGINACEAE).
60 Inflorescence of a solitary, terminal flower;
carpels 2-3 (-4).
61 Corolla with a long tube and flaring
corolla lobes (united > ½ its length)
61 Corolla of separate
petals or united only for a short length (< ¼ its length).
60 Inflorescence of several to many flowers;
carpels 5 (3 in
Galax in DIAPENSIACEAE).
63 Inflorescence an
umbel ;
petals recurved , pink to almost white
64 Fruit either a
capsule or a
utricle ;
petals 5;
stamens 5 or 10.
65 Inflorescence a somewhat to very
diffuse panicle , with 3 or more orders of branching, not giving at all the impression that the overall
inflorescence is made of
racemose units.
66 Leaves
entire ;
stamens 5; [plants of
tidal marshes ]
66 Leaves
serrate or
crenate ;
stamens 10; [plants of various habitats, especially rock outcrops and bottomland forests and streambanks, never in
tidal marshes ]
65 Inflorescence either a single terminal
raceme (sometimes
spike -like), or of 1 to several terminal and
axillary racemes (these sometimes combined into a
diffuse panicle , but one whose structure is clearly made up of many
racemes ).
67 Inflorescence of 1-several terminal and
axillary racemes , the plant typically well-branched, especially from the
base ;
stamens 5
67 Inflorescence of a single, terminal
raceme , the plant unbranched;
stamens 10 (or 5, with 5
staminodes )
68 Inflorescence spike -like, the flowers many (> 40), barely
pedicelled ;
capsule 3-
locular
68 Inflorescence a
raceme , the flowers fewer (< 20) and distinctly
pedicelled ;
capsule 5-
locular
1 Leaves either 3-foliolate or
palmately 4-11-foliolate (all the
leaflets attached at a common point, or the
leaflets slightly
pedate in
Helleborus foetidus in RANUNCULACEAE).
2 Inflorescence an
involucrate head subtended by
phyllaries , the heads solitary or many and variously arrayed in secondary
inflorescences , the
ovary inferior , the
corolla connate and
tubular at least basally, the
calyx absent, the
stamens 5, the fruit a
cypsela
2 Inflorescence , flower, and fruit structure various, but not with the combination of features as above.
3 Leaflets obviously and sharply
serrate ;
pistils 1 or 5-many; fruit an
achene , or an aggregate of
achenes ,
drupelets , or
follicles .
4 Leaflets 7-11, slightly
pedate in their arrangement, evergreen
5 Pistil 1; fruit an
achene
5 Pistils 5-many; fruit an aggregate of
achenes ,
drupelets , or
follicles
3 Leaflets entire , finely
denticulate , or very obscurely
toothed (or irregularly
serrate or
lobed in
Cardamine in BRASSICACEAE);
pistil 1; fruit simple, a
legume ,
capsule ,
silique , or
berry .
6 Corolla bilaterally symmetrical;
petals 5; fruit a
legume ;
carpel 1
6 Corolla radially symmetrical;
petals 4 or 5; fruit either an elongate
capsule or a
berry ;
carpels 1, 2, or 5.
7 Leaflets radially arranged at the summit of the
petiole , not differentiated in size or placement into a terminal
leaflet and 2
lateral leaflets ;
leaflets prominently notched at the
apex ;
petals 5, yellow;
inflorescence axillary ,
cymose or
umbelliform ;
carpels 5
7 Leaflets differentiated in size and placement into a terminal
leaflet and 2 or more
lateral leaflets ;
leaflets not regularly notched at the
apex (a few may be slightly
retuse );
petals 4, white, pink, or yellow;
inflorescence terminal and
racemose ;
carpels 1 or 2.
8 Carpels 2; fruit a red
berry ;
petals connate , purplish-blue
8 Carpel 1; fruit a
capsule ;
petals separate, white, pink, or yellow.
9 Stem leaves 1-3,
alternate [or
whorled or opposite];
leaflets 3, irregularly
serrate ,
lacerate , or additionally divided or
lobed ; fruit a
silique ;
carpels 2
9 Stem leaves >3,
alternate ;
leaflets (1-) 3-7, each
entire or obscurely
toothed ; fruit a
capsule ;
carpel 1
1 Leaves either 1-
pinnately compound (all
leaflets attached to a central
rachis ) or more complexly compound (with several orders of branching, some
leaflets at least attached to second-order branches from the
rachis ).
10 Inflorescence an
involucrate head subtended by
phyllaries , heads solitary or many, variously arrayed in secondary
inflorescences ; fruit a
cypsela ;
ovary inferior
10 Inflorescence various, but not as above; fruit various, not as above;
ovary superior .
11 Leaves 1-
pinnately compound (all
leaflets attached to a central
rachis ).
12 Flowers bilaterally symmetrical,
papilionaceous ; fruit a
legume ; leaves even-
pinnately or odd-
pinnately compound , the terminal
leaflet sometimes replaced by a
tendril ;
leaflets entire or at most minutely
denticulate
12 Flowers radially symmetrical (or barely bilaterally symmetrical in
Erodium in GERANIACEAE); fruit a
capsule ,
capsular but of 5
mericarps , or an aggregate of
achenes ,
nutlets , or
follicles (in some cases the # of
pistils from many down to 2 or even 1); leaves odd-
pinnately compound , never with
tendrils ;
leaflets serrate (or
entire to
shallowly lobed in
Polemonium in POLEMONIACEAE,
Cardamine in BRASSICACEAE, and
Floerkea in LIMNANTHACEAE).
13 Pistils many (only 1-2 in
Agrimonia ,
Poteridium ,
Poterium , and
Sanguisorba ); fruit an aggregate of
achenes ,
nutlets , or
follicles ;
hypanthium present;
stamens 5-many (only 4 in
Poteridium and
Sanguisorba )
13 Pistil 1 (or deeply 2-3-
lobed in
Floerkea in LIMNANTHACEAE); fruit a
silique ,
capsule ,
schizocarp of 2-3
mericarps , or a
capsular schizocarp of 5
mericarps (
Erodium in GERANIACEAE);
hypanthium absent;
stamens 3-6.
15 Petals 5, fused (distinct in
Erodium in GERANIACEAE);
stamens 5;
inflorescence axillary or terminal,
cymose , consisting of subcapitate,
umbel -like, or
helicoid cymes ; fruit either a
capsule , or a
capsular schizocarp of 5
mericarps (
Erodium in GERANIACEAE).
16 Flowers slightly bilaterally symmetrical (2 of the
petals of different size than the other 3); fruit a
capsular schizocarp of 5
mericarps ;
carpels 5
16 Flowers radially symmetrical; fruit either a
loculicidal capsule or a
berry ;
carpels 2 or 3.
17 Fruit a
berry ; fresh plant rankly fragrant
17 Fruit a
capsule ; fresh plant not aromatic.
18 Capsule 2-
valvate ;
carpels 2;
leaflets prominently
serrate or with some
tooth -like sublobes
11 Leaves more complexly compound (with 2 or more orders of branching, some
leaflets at least attached to second-order branches from the
rachis ).
19 Leaves 2×
even-pinnate ; flowers in
spikes or
spherical heads; fruit a
legume or
loment .
19 Leaves either 2×
odd-pinnate or more complexly 2-4×
ternately or
ternately -
pinnately compound ; flowers in various
inflorescences ; fruits various (not
legumes or
loments ).
20 Leaf
segments linear , less than 2 mm wide.
21 Inflorescence either a terminal solitary flower or terminal
raceme or
panicle ;
ovary superior , either of 2 fused
carpels or of 1-5or many distinct 1-
carpellate pistils ; fruit either a
capsule or an aggregate of
follicles or
achenes .
22 Ovary of 2 fused
carpels ; fruit a
capsule (1-seeded and
indehiscent in
Fumaria )
22 Ovary of 1-5or many distinct 1-
carpellate pistils ; fruit an aggregate of
follicles or
achenes
20 Leaf
segments broader,
lanceolate ,
ovate , or
elliptic , > 5 mm wide.
23 Erect or
sprawling herb;
stamens 5-6 or >15.
24 Leaflets sharply
serrate , with usually many teeth on each
leaflet side, the total number of “points” per
leaflet > 10.
26 Flowers
bisexual ;
carpels 1-8; fruit an aggregate of
follicles , a single
follicle , or an
indehiscent berry -like fruit
26 Flowers
unisexual ;
carpels (in
pistillate flowers) of 3 (-5)
carpels ; fruit an aggregate of
follicles
24 Leaflets entire , or with 1-several, broad,
obtuse , rounded, or broadly
acute “sublobes”, especially towards the tip of the
leaflet , the total number of “points” per
leaflet < 10.
28 Perianth bilaterally symmetrical, the
corolla of 4 fused
petals ; plant a
scandent vine or erect or
sprawling herb
28 Perianth radially symmetrical, of 1-5
whorls of separate
perianth parts; plant an erect herb.
29 Perianth of 4-5
whorls of 3 parts each (some of the
whorls modified into
nectaries );
pistil 1, 1-
carpellate ; fruit a drupelike, blue,
naked seed; largest
leaflets > 6 cm long, obviously longer than wide
29 Perianth of 1
whorl ; of 4-5 parts;
pistils many, each 1-
carpellate ; fruit an aggregate of
achenes or an aggregate of
follicles ; largest
leaflets either < 6 cm long, or if longer than 6 cm, also about as wide as long
1 Inflorescence an
involucrate head subtended by
phyllaries , the heads solitary or many and variously arrayed in secondary
inflorescences , the
ovary inferior , the
corolla connate and
tubular at least basally, the
calyx absent, the
stamens 5, the fruit a
cypsela
1 Inflorescence , flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head, e.g.
Eryngium in APIACEAE, but then with other features differing, such as
stamens 4, or green
calyx present, or fruit a
schizocarp of
mericarps , etc.).
2 Plant an herb, sometimes
sprawling ,
reclining (e.g.
Cymbalaria in PLANTAGINACEAE,
Aconitum in RANUNCULACEAE), but lacking
climbing adaptations such as
tendrils or
twining stems.
6 Ovary inferior ;
inflorescence an
umbel (or flowers solitary or in
dichasia in
Mentzelia ); fruit a
schizocarp of 2
mericarps or a
capsule (
Mentzelia ).
7 Flowers solitary or in
dichasia ; fruit
capsules dehiscing via
apical valves
8 Involucre absent or minute
6 Ovary superior ;
inflorescence various, not an
umbel ; fruit various, a
capsule , an aggregate of
achenes or
follicles , or a ring of (>2)
mericarps .
9 Perianth uniseriate , the
corolla absent (the
calyx petaloid and white in
Cnidoscolus ); flowers
unisexual ; plants either with
stinging hairs or not
9 Perianth biseriate (
uniseriate in
Aphanes in ROSACEAE and in
Trautvetteria in RANUNCULACEAE); flowers
bisexual ; plants lacking
stinging hairs .
10 Pistils many (or 2-3 in
Aphanes in ROSACEAE), each with 1
carpel , arranged
spirally or in a ring (if in a ring, of 2-5); fruit an aggregate of
achenes ,
follicles , or
utricles .
11 Perianth bilaterally symmetrical, either hooded or spurred; fruit an aggregate of
follicles
11 Perianth radially symmetrical, not hooded or spurred; fruit an aggregate of
utricles or
achenes (
plumose achenes in
Geum )
12 Stamens showy, bright white,
dilated towards the tip;
pistils ca. 15; fruit an aggregate of
utricles
12 Stamens not showy, white, or
dilated towards the tip;
pistils many (> 25); fruit an aggregate of
achenes .
13 Flowers lacking a
hypanthium ;
achenes short-beaked
13 Flowers with a prominent
hypanthium ;
achenes with an elongate,
plumose beak
10 Pistil 1, with 1-to many
carpels (in many MALVACEAE, the
carpels loosely united in a ring of more than 5 around the
style ); fruit a
capsule , an
achene , a
follicle , or a ring of 3 or 5-many 1-seeded
mericarps .
14 Perianth biseriate , with well-developed and differentiated
calyx and
corolla
15 Corolla bilaterally symmetrical, the
petals connate (except distinct in
Delphinium in RANUNCULACEAE); fruit a
capsule , a
follicle , or a
schizocarp of 3 1-seeded
mericarps .
16 Corolla not spurred; fruit an elongate (10-20 cm)
capsule with 2 curved
beaks
16 Corolla with a
nectar spur ; fruit < 3 cm long.
17 Petals distinct; fruit a
follicle
18 Carpels 3; fruit a
schizocarp of 3 1-seeded
mericarps ;
stamens 8
15 Corolla radially symmetrical, the
petals distinct (fused and
tubular in
Ipomoea ); fruit a
capsule or a
schizocarp consisting of a ring of 5-many 1-seeded
mericarps .
19 Stem
trailing ;
petals fused and
tubular
19 Stem erect;
petals separate.
20 Stamens many,
connate into a
stamen tube;
carpels 5-many, completely or only loosely fused; fruit a
capsule or a
schizocarp of 5-many
mericarps borne in a ring;
calyx often subtended by an
epicalyx (an additional
calyx -like, green,
foliaceous whorl of
bracts )
20 Stamens 5 or 10, distinct;
carpels 2 or 5, fused; fruit a
capsule or a
schizocarp of 5 1-seeded
mericarps .
21 Fruit a
schizocarp of 5 1-seeded
mericarps ;
carpels 5;
stamens 10