Colors

Data mode

Account

Login
Sign up

Collapse this

Support the Flora of the Southeastern US

2024 has been a banner year for making the best flora we can imagine. We've created:
With financial support from people like you, we are aiming even higher in 2025. Together we can accomplish all this: Vote on our 2025 priorities
  • Add Global Conservation Ranks (GRanks) vote
  • Professional graphic keys (polyclaves) to individual families/genera vote
  • 2 new FloraQuest apps: Florida & Mid-South vote
  • Image overlays highlighting diagnostic characters with arrows vote
  • iNaturalist integration in FloraQuest vote
Write-in vote: vote
We've set a goal of recruiting 200 ongoing supporters to donate $15 or more each month in 2025. Please help us reach this goal and make next year's flora even better:

No key was found for the requested taxon, but it has only one child: Chamaepericlymenum canadense. Showing where it is keyed below.

X
Keyed in multiple places:

Click the number at the start of a key lead to highlight both that lead and its corresponding lead. Click again to show only the two highlighted leads. Click a third time to return to the full key with the selected leads still highlighted.

Key to Cornaceae

Copy permalink to share

1 Leaves alternate (the internodes typically short and therefore the leaves apprearing nearly whorled)
1 Leaves opposite.
  2 Herb or dwarf shrub from a woody rhizome, to 2 dm tall; leaves in 2-4 pairs below the inflorescence; [of NJ and montane VA and WV northward]
  2 Shrub or tree, much taller than 2 dm when mature; leaves many; [collectively widespread].
    3 Inflorescence a head, subtended by 4 showy (white, creamy, or pink) bracts.
    3 Inflorescence a compound corymb, with green, tan, or brown (non-petaloid) bracts.
      4 Inflorescence a simple umbel; fruit red when ripe; inflorescence bracts brownish, 4, 5-10 mm long, subtending the umbel and enclosing it in winter
      4 Inflorescence a compound corymb; fruit blue or white when ripe; inflorescence bracts greenish, many, < 2 mm long, subtending the primary and secondary branches of the corymb

Key H: woody plants with whorled leaves

Copy permalink to share | Check for keys that lead to this key

1 Leaves tiny, bract-like, triangular, 6-14 (-17) per node
1 Leaves either needle-like, scale-like, or flattened and large, (2-) 3-4 (-6) per node.
  2 Leaves needle-like or scale-like, terete, angled, or flat in ×-section, < 2 cm long; leaves (2-) 3-4 (-6) per node
  2 Leaves flat, > 3 cm long; leaves (2-) 3 per node; [Eudicots].
    3 Plant a subshrub, < 3 dm tall, with < 10 leaves per stem.
      4 Leaves entire, broadly elliptic; flowers numerous, in a hemispherical head, subtended by 4 large white bracts
      4 Leaves serrate, narrowly ovate or narrowly obovate; flowers (1-) 2-8 in a long-peduncled umbel or corymb, not subtended by bracts
    3 Plant a shrub or tree, > 3 dm tall, with many > 10 leaves per stem.
          6 Leaves cordate at base; leaves about as long as wide; medium to large tree.
             7 Flowers white to yellow; capsules linear, >10× as long as wide; leaf undersurface with curly simple hairs; nectar glands present in the main vein axils on the undersurface of the leaf (visible from the underside or the upperside in fresh leaves and herbarium specimens as a triangle 1-4 mm on a side)
             7 Flowers lavender; capsules ellipsoid, < 2× as long as wide; leaf undersurface with branched (dendritic or stellate) hairs; nectar glands absent
          6 Leaves cuneate to rounded at base; leaves > 1.5× as long as wide; shrub to small tree.
               8 Leaves rounded at the tip
                 9 Leaves lanceolate or oblong-elliptic (> 2.5× as long as wide), the secondary venation not prominent; inflorescences axillary or terminal; flowers pink or white.
                   10 Inflorescences axillary; flowers pink; leaves thin and herbaceous; [native plants of wetlands]
                   10 Inflorescences terminal or axillary; flowers pink or white; leaves thick and leathery; [exotics of uplands or wetlands, persistent or weakly naturalized]
                     11 Fruit follicles; flowers variously colored, showy and salverform; inflorescence terminal; leaves alternate to whorled (pseudo-whorled)
                 9 Leaves ovate (< 2× as long as wide), the secondary venation prominent and arching-parallel; inflorescences terminal; flowers white, greenish-yellow, red, or orange.
                       12 Flowers in a spherical or hemispherical head; corollas white or greenish-yellow
                       12 Flowers in a monochasial helicoid cyme; corollas red to orange

Key J4: shrubs and subshrubs with opposite simple leaves with entire margins

Copy permalink to share | Check for keys that lead to this key

1 Aerial and epiphytic, hemiparasitic shrub
1 Terrestrial, autotrophic or hemiparasitic shrub or subshrub.
  2 Leaves succulent, nearly as thick as wide; [brackish to saline situations]
  2 Leaves herbaceous (succulent in Borrichia), much wider than thick; [various habitats].
    3 Creeping or short subshrubs, the stems primarily prostrate, < 2 dm tall.
      4 Well-developed leaves 4-6 per stem; inflorescence a head subtended by 4 large white bracts
      4 Well-developed leaves many per stem; inflorescence of individual flowers axillary in pairs or clusters or in terminal cymes.
        5 Flowers yellow; leaves with pellucid or dark punctate glands (use at least 10× magnification)
        5 Flowers white, pale pink, or deep pink; leaves lacking sessile, punctate glands.
          6 Leaves linear; flowers pale to deep pink, 5-merous
          6 Leaves orbicular or elliptic; flowers white to pale pink, 4-merous or 5-merous
    3 Upright or scrambling shrubs, > 3 dm tall
               8 Scrambling shrubs, armed with recurved paired spines at the nodes
                 9 Inflorescence a terminal head of many flowers.
                   10 Head flattened, either subtended by 4 large white bracts or by an involucre with >5 green phyllaries.
                     11 Head subtended by 4 large white bracts; leaves with prominently parallel-arcing secondary veins; flowers 4-merous
                     11 Head subtended by an involucre of >5 green phyllaries; leaves with venation otherwise; flowers 5-merous
                 9 Inflorescence otherwise, either of a solitary flower, or one of a wide variety of inflorescences with flowers attached at different points along branched or unbranched axes (e.g. axillary). {add: [Lagerstroemia] LYTHRACEAE; [Rosmarinus] LAMIACEAE; [Laguncularia] COMBRETACEAE; [Buxus] BUXACEAE; [Exochorda] ROSACEAE; various other [see spreadsheet]}
                       12 Inflorescence flat-topped (broader than long), terminal, a compound cyme or corymb.
                          13 Flowers bright yellow; stamens many; leaves < 1.5 cm wide; fruit a capsule; leaves with pellucid or dark punctate glands (use at least 10× magnification)
                          13 Flowers white or creamy; stamens 4-5; leaves > 1.5 cm wide; fruit a drupe; leaves lacking sessile, punctate glands.
                       12 Inflorescence either terminal and not flat-topped (longer than wide), or axillary and variously shaped, or terminal and solitary, or leaf-opposed.
                              15 Carpels many (> 9), either separate or fused; stamens many; perianth segments either many and undifferentiated into calyx and corolla, maroon, brown, or yellow (in CALYCANTHACEAE), or differentiated into a fleshy and persistent calyx of 5-9 sepals, and a deciduous corolla of 5-9 red (or white) petals (Punica in LYTHRACEAE).
                                16 Fruit a wrinkled, 3-7 cm long, brown to black, elliptical aggregate of nearly spherical, large achenes; flowers solitary in axils; perianth segments many and undifferentiated into calyx and corolla, maroon, brown, or yellowish; ovary superior; branches unarmed
                                16 Fruit a leathery, 4-15 cm in diameter, reddish, spherical berry with obpyramidal seeds surrounded by a juicy sarcotesta (pomegranate); perianth differentiated, the sepals fleshy and persistent on the fruit, the petals deciduous, 5-9, bright red to white; ovary inferior; branches typically armed with axillary spines
                              15 Carpels 1-5, fused; stamens either 1-5 or 8-10; perianth segments 4-5 or 8, variously colored; fruit a simple capsule, drupe, or berry (including berry-like fruit); flowers 2-many, in axillary or terminal inflorescences (pistillate flowers sometimes solitary in SANTALACEAE and THESIACEAE); [Eudicots].
                                  17 Ovary inferior; corolla absent, radially symmetrical, or bilaterally symmetrical; fruit either a berry or a 1-seeded drupe.
                                    18 Flowers unisexual and plants dioecious; corolla absent; pistillate flowers solitary, staminate flowers in pedunculate umbels or cymes, either terminal or axillary; fruit a 1-seeded drupe; leaves acute to acuminate at the apex
                                    18 Flowers bisexual and plants hermaphroditic; corolla present; flowers paired, terminal or axillary, or in axillary spikes; fruit a berry; leaves rounded, obtuse, to acute (or acuminate in Lonicera maackii) at the apex
                                  17 Ovary superior; corolla radially symmetrical (absent in Forestiera in OLEACEAE); fruit either a 1-4-seeded drupe, or a many-seeded berry (or berry-like fruit), or a capsule.
                                         20 Leaves with a conspicuous mix of silvery stellate hairs (upper) and rusty colored scales (lower); ovaries bearing rusty colored scales; flowers small, yellowish and inconspicuous; [nw. PA northward]
                                         20 Leaves glabrous or nearly so, lacking a conspicuous mix of stellate hairs adaxially and rusty scales abaxially; ovaries not bearing rusty colored scales; flowers of various size and color.
                                           21 Stamens 8-10, of 2 different lengths in each flower; petals separate, 4-5 (-7), pink purple, 10-15 mm long; stems strongly arching, rooting at the tips; [plants of flooded to saturated wetlands]
                                           21 Stamens either (1-) 2 (-4), or 4-5, or 10, all of the same length; petals fused (separate in RHAMNACEAE and BUXACEAE, but then < 5 mm long and white or cream), white, bright-yellow, lilac, or pink; stems erect (or at least not arching and rooting at the tips); [plants of various habitats].
                                               23 Fruit a loculicidal capsule, dehiscing into 3 valves; branches square in ×-section; leaves < 2 cm long; [exotic, cultivated and weakly established, of temperate areas]
                                               23 Fruit a drupe with 2-4 pyrenes; branches round or nearly so in ×-section; leaves > 2 cm long; [natives, of peninsular FL]
                                             22 Petals fused, 4-5, white, bright yellow, lilac, or pink; stamens either (1-) 2 (-4) or 10; fruit either a capsule or a 1-seeded drupe.

Key S1: herbaceous dicots with opposite, simple, and unlobed leaves on the stem

Copy permalink to share | Check for keys that lead to this key

1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head, e.g. Pycnanthemum in LAMIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.).
  2 Leaves scale-like, stems fleshy; flowers embedded in the fleshy stem, no perianth present; [saline environments (coastal or rarely inland)].
  2 Leaves small to large; stems not fleshy; flowers sessile or on pedicels; [collectively of many habitats, saline and not].
      4 Perianth of a single whorl (petals absent) (note that in Mirabilis in NYCTAGINACEAE the petaloid calyx is subtended by a 5-lobed fused set of involucral bracts).
        5 Leaves herbaceous, suborbicular, about as long as wide or wider than long; calyx 3- or 4-merous; stamens 4, 8, or 12.
          6 Plant ascending, with a single node (2 leaves); leaves > 6 cm long and wide; calyx 3-merous, brown to yellowish; stamens 12
          6 Plant creeping, with many nodes; leaves 3-15 mm long and wide; calyx 4-merous, yellow to greenish; stamens 4 or 8
        5 Leaves fleshy, linear, lanceolate, to broadly ovate, at least slightly longer than broad; calyx 5-merous; stamens 3, 5, or 10.
             7 Flowers axillary, sessile or nearly so, solitary or a few; petaloid sepals widely spreading, separate; leaves linear to oblanceolate; stamens 5 or 30-50
             7 Flowers in terminal cymose panicles; petaloid sepals connate into a narrow tube (reminiscent of the corolla of Ipomoea); leaves lanceolate, elliptic, ovate, or broadly ovate; stamens 3 or 5
      4 Perianth in 2 whorls (sepals and petals both present).
                 9 Leaves distinctly 3-veined from the base, the 3 veins converging again at the leaf apex
               8 Petals connate into a tube (at least basally); inflorescence often a head or dense terminal cyme (also axillary, or solitary on long peduncles).
                   10 Petals 5; stamens 3, 4, or 5
                     11 Upright herb; flowers in axils or terminal corymbs
                       12 Flowers in terminal corymbs; fruit dry
                   10 Petals 4 (or 6 or 8 in Richardia in RUBIACEAE); stamens 4, 6, or 8.
                          13 Inflorescence a head or more diffuse (see below), sometimes subtended by green bracts.
                            14 Leaves serrate; corolla bilaterally symmetrical (especially the flowers near the outer edge of the head); inflorescence a head
                            14 Leaves entire; corolla radially symmetrical; inflorescence a head or more diffuse (see below).
                              15 Petals acute; flowers in terminal panicles, cymes, or panicles, or axillary; plant habit various, not simultaneously with all the characters below
                              15 Petals broadly rounded; flowers axillary, solitary; plant a diffusely branched herb with linear leaves
                                16 Perianth of a single whorl (petals absent) or missing entirely (petals and sepals both absent). {key lead number needs adjusting}
                                  17 Inflorescence a cyathium, consisting of a single pistillate flower (reduced to a single 3-carpellate pistil) and 2 or more staminate flowers (each reduced to 1 stamen), borne in a cup-like involucre, the involucre bearing pointed or rounded glands, these sometimes brightly colored and petaloid, mimicking an individual flower (the cyathia then secondarily arranged in terminal cymes, or solitary and axillary, etc.); fresh plants with milky juice; fruit a 3-lobed, 3-locular capsule
                                  17 Inflorescence not a cyathium (and staminate or bisexual flowers with > 1 stamen, except Callitriche in PLANTAGINACEAE); fresh plants lacking milky juice; fruit various, not as above.
                                    18 Flowers many, in axillary spikes, cymes, or glomerules, or in terminal spikes, heads, cymes, or panicles; leaves entire or serrate.
                                             22 Leaves entire, or with a few very obscure crenations (Iresine) or basally disposed rounded lobe-like teeth (Atriplex); plants without stinging hairs.
                                                 24 Style 1; leaves generally either longer than 30 mm, or wider than 8 mm (if linear and smaller than those dimensions, then fleshy).
                                16 Perianth in 2 whorls (sepals and petals both present).
                                                        27 Pistils 4-5, in a single whorl; stamens 4, 5, 8, or 10; fruit an aggregate of follicles
                                                      26 Gynoecium either of 1 pistil (with 1 or more carpels), or of 2 pistils, united only by the style and stigma (APOCYNACEAE).
                                                            29 Leaves with pellucid punctate glands (most easily visible with transmitted light); stamens often fascicled into 3, 4, or 5 fascicles; petals yellow or pinkish
                                                                 31 Petals 3; sepals 5, dimorphic, the 2 outer sepals narrower than the 3 inner and concave sepals; stamens (3-) 5-15 (-25)
                                                                 31 Petals 4-7; sepals 4-7, normally monomorphic; stamens 4, 5, 6, 8, 10, or 12 (or sometimes rarely 2 or 3).
                                                                     33 Capsule 1-locular, dehiscent apically by teeth or valves; sepals connate into a tube or separate; styles 2-5; perianth 4-5-merous; stamens 4, 5, 8, or 10 (or rarely 2 or 3)
                                                                       34 Corolla radially symmetrical (or so slightly bilaterally symmetrical as to be mistakable as radially symmetrical); stamens as many as the corolla lobes (or 1 less in Ruellia in ACANTHACEAE, Buchnera in OROBANCHACEAE, Trichostema in LAMIACEAE, and Verbena in VERBENACEAE); carpels 2 or 3.
                                                                         35 Pistils 2, united only by the style and stigma; fruit a schizocarp of 2 1-carpellate follicles (often single by abortion); plant with milky juice when fresh (except Catharanthus); leaves entire
                                                                         35 Pistil 1 (of 2-5 fused carpels); fruit either a 2-5-carpellate capsule or of 2 or 4 1-seeded nutlets derived from 2 carpels; plant lacking milky juice; leaves entire or serrate.
                                                                                  39 Stamens either 4, 1 fewer than the 5 corolla lobes, or 2 (with 2 staminodes); corolla usually slightly bilaterally symmetrical (the flower as a whole made bilaterally symmetrical by the 2 or 4 stamens).
                                                                       34 Corolla bilaterally symmetrical (or the corolla 2-lipped but the corolla lobes twisted so as to make the flower asymmetrical); fertile stamens fewer than the corolla lobes (except Plantago in PLANTAGINACEAE, which is equal, with 4 each; a few genera have a 5th, sterile, stamen which is obviously different in form than the 4 fertile stamens) (note that many corollas are bilabiate and the number of corolla lobes, 4 or 5, may be difficult to interpret); carpels 2.
                                                                                             44 Carpels 2, each carpel slightly to deeply lobed, separating at maturity into 4 half-carpellate units (not separating in Phyla in VERBENACEAE); fruit a schizocarp of 4 mericarps (or 2 nutlets in Phyla in VERBENACEAE).
                                                                                               45 Inflorescence a thyrse, verticillaster, or terminal cyme, the flowers borne in cymose lateral branches; corolla strongly bilaterally symmetrical (rarely nearly radially symmetrical); stems square in ×-section (or sometimes rounded, especially on older growth); fresh plants often (but not always) aromatic
                                                                                               45 Inflorescence of spikes, racemes, or heads, the flowers or fruits single at nodes; corolla often nearly radially symmetrical; stems rounded in X-section (rarely square); fresh plants usually not aromatic
                                                                                                                     55 Inflorescence of 1 or more terminal (and sometimes upper axillary) spikes or racemes; corolla 10-50 mm long (6-8 mm long in Phryma in PHRYMACEAE), white, pink, blue, purple, or yellow; fruit either a loculicidal capsule (OROBANCHACEAE) or a single seeded achene (Phryma in PHRYMACEAE).
                                                                                                                53 Flowers axillary and solitary, borne in the axils of normally-sized leaves or somewhat reduced but still large and leaf-like bracts [some taxa keyed here and below].
                                                                                                                                      63 Corolla red or orange, with a very narrow, cylindrical tube, the lobes then flaring into a limb about 1 cm across; plants blackening on drying; [rare exotic, in crop fields, a noxious hemiparasitic weed under quarantine]