1 Culm nodes , at least the lower,
bearded (often
retrorsely ).
2 Nodes with a
viscid band below the
beard ;
internodes and
sheaths velvety-
pubescent ;
vernal culm blades 9-20 cm long, 9-20 mm wide
2 Nodes without a
viscid band below the
beard ;
internodes and
sheaths glabrous , the lower sometimes sparsely hairy;
vernal culm blades 3.5-14 cm long, 7-15 mm wide
3 Second
glume and
sterile lemma acute to short-
acuminate , some in a given specimen 0.3 mm or more longer than fertile
lemma .
4 Ligule an eciliate membrane;
peduncle scabrous ; first
glume 0.3-0.6 (-0.8) mm long,
reniform to suborbicular
4 Ligule a
ciliate membrane or entirely
ciliate ;
peduncle smooth; first
glume 0.7-1.6 mm long, broadly
ovate to
lanceolate .
5 Ligule a
ciliate membrane; larger leaves 10-20 cm long;
spikelets pubescent , (2.6-) 2.8-3.4 (-3.6) mm long; first
glume (0.9-) 1.0-1.6 mm long
5 Ligule entirely
ciliate ; larger leaves 9-14 cm long;
spikelets glabrous , 2.1-2.6 mm long; first
glume 0.7-1.2 mm long
3 Second
glume and
sterile lemma blunt to subacute, shorter than, equaling, or barely exceeding fertile
lemma , never by as much as 0.3 mm.
6 Ligule ciliate , 0.2-1.1 (-2.1) mm long; leaf
blades 5-14 cm by 5-17 mm; second
glume 0-0.3 mm shorter than fertile
lemma .
7 Culms 30-75 cm long; lower
sheaths glabrous to sparsely
papillose -
pilose ;
panicle pulvini glabrous to sparsely
pilose ;
spikelets 1.8-2.6 mm long; first
glume narrowly
triangular (
triangular -
ovate ); second
glume 0-0.1 mm shorter than fertile
lemma
7 Culms 60-150 cm long; lower
sheaths papillose -
pilose ;
panicle pulvini pubescent ;
spikelets 2.2-2.8 (-3.0) mm long; first
glume broadly
ovate ; second
glume 0.05-0.3 mm shorter than fertile
lemma
6 Ligule a
ciliate or eciliate membrane (0-) 0.3-0.9 mm long; leaf
blades 5-28 cm by 5-40 mm; second
glume 0-0.1 mm shorter than fertile
lemma .
8 Lower
sheaths papillose -
hispid with spreading
hairs ;
ligule a membrane 0.4-0.9 mm long; leaf
blades 10-28 cm long
8 Lower
sheaths glabrous ,
puberulent , or softly
villous basally;
ligule either a membrane 0-0.3 mm long or a
ciliate membrane 0.4-0.7 mm long; leaf
blades 5-18 cm long.
9 Ligule a
ciliate membrane 0.4-0.7 mm long; leaves 15-40 mm wide;
spikelets 1.6-2.0 mm wide
9 Ligule a membrane 0-0.3 mm long; leaves 5-25 mm wide;
spikelets 1.1-1.3 mm wide
10 Cauline blades 5-18 cm long, 6-14 mm wide, averaging about 10 × as long as wide, the
margins nearly parallel;
spikelets 2.9-3.7 mm long; first
glumes 1.1-2.6 mm long, 2/5 or more as long as the
spikelets
10 Cauline blades 7-12 cm long, (6-) 12-25 mm wide,
ovate -
lanceolate , averaging about 6 × as long as wide, the
margins curved most of their length;
spikelets 2.0-3.2 mm long; first
glumes 0.5-1.0 mm long, about 1/4 as long as
spikelets .
11 Culms erect;
blades more nearly symmetrical;
spikelets 2.0-3.2 mm long;
sterile lemma apex usually blunt to rounded
2 Leaf
blades , at least the lower,
cordate or subcordate at the
base , 4-25 mm wide.
3 Culm blades 4-12 mm wide;
spikelets obpyriform when viewed dorsally, strongly plano-
convex when viewed laterally, usually markedly reddish-purple basally; fertile
lemma papillose
3 Culm blades 5-25 mm wide;
spikelets elliptic to
elliptic -
obovoid when viewed dorsally or laterally, greenish to faintly purple-tinged basally; fertile
lemma not
papillose .
4 Ligule ciliate , 0.2-0.8 mm long;
culm blades 5-14 mm wide;
spikelets 1.8-2.6 mm long; first
glumes 0.5-1 mm long
4 Ligule an eciliate membrane 0-0.3 mm long;
culm blades 5-25 cm wide;
spikelets 2.0-3.7 mm long; first
glumes 0.5-2.6 mm long.
5 Internodes crisp-
puberulent ; larger
culm blades 4-8 (-11) cm long, 5-10 (-12) mm wide, broadest near the
base ;
spikelets 2.1-2.7 mm long; first
glumes 0.7-0.9 mm long
5 Internodes glabrous to sparsely
pubescent , larger
culm blades 5-18 cm long, 5-25 mm wide, broadest near the middle or broadly
linear ;
spikelets 2.0-3.7 mm long; first
glumes 0.5-2.6 mm long.
6 Cauline blades 5-18 cm long, 6-14 mm wide, averaging about 10 × as long as wide, the
margins nearly parallel;
spikelets 2.9-3.7 mm long; first
glumes 11-26 mm long, 2/5 or more as long as the
spikelets
6 Cauline blades 7-12 cm long, (6-) 12-25 mm wide,
ovate -
lanceolate , averaging about 6 × as long as wide, the
margins curved most of their length;
spikelets 2.0-3.2 mm long; first
glumes 5-10 mm long, about 1/4 as long as
spikelets .
7 Culms erect;
blades more nearly symmetrical;
spikelets 2.0-3.2 mm long;
sterile lemma apex usually blunt to rounded
2 Leaf
blades tapering to the
base , 2-12 mm wide.
8 Ligule a stramineous to light brown membrane, with or without terminal
cilia .
9 Ligule a
ciliate membrane;
spikelets (2.6) 2.8-3.4 (3.6) mm long; first
glume (0.9) 1.1-1.6 mm long
9 Ligule an eciliate membrane;
spikelets 1.6-2.8 (3.0) mm long; first
glume 0.3-1.1 mm long.
10 Panicle rachis smooth,
pellucid -
punctate ; first
glume 0.3-0.6 (-0.8 ) mm long, as broad as or broader than long,
truncate to
obtuse ; larger leaves 10-25 cm long, 8-15 mm wide;
ligule 0.5-1.3 mm long; lowest elongate
culm internode > 2 mm in diameter; lowest
nodes usually
glabrous or
pubescent
10 Panicle rachis scabrous or smooth, not
pellucid -
punctate ; first
glume 0.5-1.1 mm long, longer than wide, rounded to
acute ; larger leaves 3.5-12 cm long, 3-9 mm wide;
ligule 0.1-0.6 mm long; lowest elongate
culm internode < 2 mm in diameter; lowest
nodes retrorsely bearded or
glabrous .
11 Lowest
nodes usually
retrorsely bearded ;
ligules (0.1-) 0.3-0.6 mm long; largest
vernal blades 7-12 cm long, (4.5-) 6-9 mm wide;
panicle peduncle scabrous ;
spikelets ovate -
lanceolate ,
acute , 2.0-2.4 mm long; first
glume lanceolate , blunt to
acute ; fertile
lemma smooth
11 Lowest
nodes usually
glabrous ;
ligules 0.1-0.2 (-0.3) mm long; largest
vernal blades 3.5-7 cm long, 3-6 mm wide;
panicle peduncle smooth;
spikelets elliptic , blunt to
acute , 1.6-2.2 mm long; first
glume ovate to
rotund , rounded to
acute ; fertile
lemma papillose
8 Ligule of short white
hairs or absent.
12 Leaves basally disposed, usually matted or cushion-forming, larger than the mid and upper
culm leaves;
blade margins uniformly
papillose -
ciliate ;
culms branching only at the
base , 0.5-3.5 dm tall;
internodes glabrous or sparsely
pubescent
12 Basal leaves usually
rosette -forming, not matted or cushion-forming, usually much smaller than
culm leaves;
blade margins glabrous , or
ciliate only below the middle (or sometimes
papillose -
ciliate throughout in
D. portoricense ssp.
patulum , which has densely
puberulent internodes );
culms branching at the
nodes in age, 1.5-7.5 dm tall.
13 Blades progressively longer upwards on
culm , the larger 9-20 cm long and 3-8 mm wide, (15-) 20 × or more as long as wide, firm but not stiff, not longitudinally
grooved or
wrinkled ;
blade adaxial surface (weakly-) strongly
scabrous ;
sheaths glabrous ; second
glume and
sterile lemma exceeding fertile
lemma
13 Blades similar or more often decreasing in size upwards on
culm , the larger 3-16 cm long and 2-25 mm wide; if 15 × or more as long as wide, then
blades stiff and longitudinally
grooved or
wrinkled (sect.
Angustifolia );
blade adaxial surface smooth to moderately
scabrous ;
sheaths glabrous or
pilose ; second
glume and
sterile lemma shorter than to exceeding fertile
lemma .
14 Blades of mid-
culm leaves typically long and stiff,
acuminate ,
linear or narrowly
lanceolate , usually > 10× as long as wide, only 2-6 mm wide when < 8 cm long.
15 Vegetative parts
glabrous (
spikelets pubescent , lowest
internodes and
sheaths sometimes sparsely
pubescent ,
blades and
sheaths sometimes
ciliate ); mature
panicles less than ¼ as wide as long, the branches erect-
ascending , the
spikelets often subsecund;
autumnal blades 4-10 cm long,
involute , < 2 mm wide;
spikelets 1.8-2.2 mm long.
16 Larger
vernal blades 5-8 cm long and 3-5 mm wide;
vernal panicles usually less than ¼ as wide as long,
spikelets often subsecund along the strongly
ascending to erect branches;
spikelets 1.8-2.2 mm long, second
glume and
sterile lemma pubescent , summits blunt, about equaling fertile
lemma ; [of coastal wetlands and Piedmont
prairies from NC and AR south to the Caribbean and Central America]
16 Larger
vernal blades 5-9 cm long and 2-3 mm wide;
vernal panicles usually ¼ to ½ as wide as long, the branches
ascending ;
spikelets 2.3-3.0 mm long, second
glume and
sterile lemma sparsely
pubescent to
glabrous , summits pointed to blunt, exceeding fertile
lemma ; [of c. and s. FL]
15 Vegetative parts
pubescent , at least in the lower portion of plant; mature
panicles usually more than half as wide as long, the branches spreading-
ascending , the
spikelets not noticeably subsecund;
autumnal blades 1-6 cm long,
involute and < 2 mm wide in
D. aciculare and
D. arenicoloides , or to 9 cm long, flat and 2-4 mm wide in
D. angustifolium (
autumnal blade dimensions not yet known for
D. wilcoxianum );
spikelets 1.5-3.2 mm long.
17 Culms 15-35 cm long; all
culm leaf
blades long-
hirsute on both surfaces, their
sheaths densely
papillose -
hirsute ; leaf
blades of the primary branches often exceeding the immature
vernal panicles
17 Culms 15-75 cm long;
culm leaf
blades glabrous to sparsely
pubescent , or the lower softly
villous abaxially in
D. arenicoloides ; lower
sheaths usually
villous , mid-
culm and upper
sheaths glabrous (though often
ciliate basally); leaf
blades of primary branches not exceeding immature
vernal panicles .
18 Spikelets (3.0-) 3.2-3.8 mm long,
fusiform , pointed at summit,
attenuate at
base , with both
glumes attached 0.3-0.5 mm below
sterile lemma , the
autumnal spikelets 3.5-3.8 mm long; larger
vernal leaf
blades 3-6 mm wide, the lower and mid-
culm leaf
blades of similar width;
autumnal leaf
blades involute
18 Spikelets 1.8-3.1 mm long,
obovate , blunt, and the
base not
attenuate (except in
D. arenicoloides with
autumnal spikelets 2.3-3.1 mm long); larger
vernal leaf
blades 2-8 mm wide, the lower usually wider and often shorter than mid-
culm leaf
blades ;
autumnal leaf
blades involute or flat.
19 Longest
vernal blades to 16 cm, widest
vernal leaf
blades 4-8 mm, usually longitudinally
wrinkled ;
vernal and
autumnal spikelets 2.3-3.1 mm long;
autumnal leaf
blades flat, the larger to 9 cm by 2-4 mm
19 Longest
vernal leaf
blades to 6 cm (
D. aciculare ) or 12 cm (
D. arenicoloides ) long, 2-5 mm wide, not noticeably
wrinkled ;
vernal spikelets 1.5-2.8 mm long,
autumnal spikelets either 1.5-2.3 (
D. aciculare ) or 2.3-3.1 (
D. arenicoloides );
autumnal leaf
blades involute , the larger to 6 cm by 1.5 mm.
20 Longest
vernal leaf
blades to 12 cm, widest
vernal leaf
blades 3-4 (-5) mm;
vernal spikelets 2.1-2.8 mm long;
autumnal spikelets 2.3-3.1 mm long, pointed,
attenuate , the
glumes attached 0.3-0.5 mm below
sterile lemma ; first
glumes 0.7-1.5 mm long; larger
autumnal leaf
blades to 6 cm by 1.5 mm
20 Longest
vernal leaf
blades to 6 (-8) cm, widest
vernal leaf
blades 2-5 mm;
vernal and
autumnal spikelets 1.7-2.3 mm long, blunt, not
attenuate , the
glumes attached < 0.2 mm below
sterile lemma ; first
glumes 0.6-0.9 mm long; larger
autumnal leaf
blades to 3 cm by 1 mm
14 Blades of mid-
culm leaves
lanceolate , thin or firm but not stiff, usually < 10× as long as wide, usually 7 mm or more wide when as much as 8 cm long.
21 Spikelet bases strongly
attenuate ,
apices blunt to beaked; first
glume (2/5-) ½-2/3 (-4/5) as long as
spikelet , attached 0.3-0.5 (-0.7) mm below expansion of second
glume and
sterile lemma
21 Spikelet bases not noticeably
attenuate ,
apices rounded to
acute ; first
glume < ½ as long as
spikelet , attached < 0.3 mm below expansion of second
glume and
sterile lemma .
22 Spikelets 2.9-3.8 mm long, broadly
elliptic , rounded at the summit, with broad and thick
nerves
22 Spikelets 2.1-2.9 mm long,
elliptic or
obovate , rounded or pointed at the summit, the
nerves often raised, but not broad and thick.
24 Culms 50-150 cm long; at least lowest
sheaths papillose -
pilose .
25 Lowest
internodes usually purple-
villous ;
nodes retrorsely (-spreading)
bearded ;
spikelets 1.8-2.2 mm long, moderately densely
pubescent ; first
glume broadly
ovate , subacute, 0.4-0.7 mm long, usually < 1.5 × as long as wide; second
glume about as long as
sterile and fertile
lemmas ; fertile
lemma 1.7-1.8 mm long
25 Lowest
internodes glabrous or
pubescent or spreading
villous , usually not purplish;
nodes glabrous , sparsely
pubescent to sparsely
bearded ;
spikelets 2.2-2.8 (-3.0) mm long, sparsely
pubescent ; first
glume 0.7-1.1 mm long, usually > 1.5 × as long as wide; second
glume shorter than
sterile and fertile
lemmas ; fertile
lemma 2.1-2.2 mm long
24 Culms 18-100 cm long;
sheaths glabrous or the lowest softly
pubescent but not
papillose -
pilose .
26 Spikelets strongly plano-
convex when viewed laterally,
obpyriform when viewed dorsally, (2.1-) 2.2-2.6 mm long; fertile
lemma and
palea papillose or smooth; first
glume usually broadened above
base , rounded or broadly tapered distally;
glume bases often reddish-purple.
27 Culm leaves 2-5 mm wide; fertile
lemma and
palea smooth, shiny
27 Culm leaves 4-12 mm wide; fertile
lemma and
palea papillose -roughened
26 Spikelets biconvex to
elliptic when viewed laterally, variously shaped but not
obpyriform when viewed dorsally; fertile
lemma and
palea smooth or
reticulate (or
papillose in D. lucidum with weak and
sprawling culms and
spikelets 1.7-2.3 mm long); first
glume usually tapered from near
base to
apex ;
glume bases various
23 Culm internodes crisp-
puberulent (sparsely so in
D. webberianum) ;
sheaths puberulent or
glabrous .
28 Spikelets elliptic , sub-
acute to pointed, greenish or faintly purple-tinged basally
28 Spikelets strongly plano-
convex when viewed laterally,
obpyriform when viewed dorsally, broadly rounded, usually markedly reddish-purple basally.
29 Fertile
lemma and
palea papillose ;
spikelets 2.2-2.6 mm long; lower
culm blades 6-12 mm wide,
glabrous
29 Fertile
lemma and
palea smooth (minutely
reticulate but not
papillose );
spikelets (1.8) 1.9-2.2 (-2.5) mm long; lower
culm blades 2-8 mm wide,
glabrous ,
glabrate , or
puberulent (especially abaxially)