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Key to Combretaceae
https://fsus.ncbg.unc.edu/main.php?pg=show-key.php&keyid=40097
2 Mangrove, a tree or erect shrub, often with pneumatophores; petiole with 2 glands near blade junction; petals ca. 1 mm long, greenish white; leaf blades with glands in sunken pits, several mm in from the leaf margin; flowers with shallowly cupulate hypanthium, inconspicuous | |
3 Trees or shrubs, either with or without pneumatophores; flowers in axillary spikes. | |
Key G4: shrubs and subshrubs with alternate, simple, unlobed, entire leaves
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3 Leaves broader, < 15× as long as wide, leaf apices variously shaped, if pointed usually not conspicuously sharpened; [Eudicots, Basal Angiosperms, or Monocots]. | |
5 Inflorescence an involucrate head or a raceme or a panicle consisting of spherical heads (Conocarpus in COMBRETACEAE). | |
6 Inflorescence an involucrate head, the heads consisting of a receptacle bearing few-many cypselae; plants of various habitats, but not forming dense stands of shrubs in coastal habitats | |
5 Inflorescence not an involucrate head, instead either solitary (Illicium in ILLICIACEAE) or variously branched, spicate, racemose, umbellate, or fascicled. | |
8 Leaves alternate, but usually clustered densely towards branch tips; leaves narrowly oblanceolate, 1-4 cm long; flowers yellow, 5-merous, the petals diminuitive (ca. 4 mm) and clawed basally (the petals often falling off by midday); [of coastal beaches, dunes, and hammocks; FL peninsula] | |
8 Plants not with the above combination of traits: leaves longer and variously shaped (if narrowly oblanceolate, leaves much longer than 40 cm long) usually well-spaced throughout the stems (if clustered, then other characters lacking); [plants widely distributed in a variety of habitats]. | |
12 Leaves largely covered with silver and/or bronze lepidote scales and/or dense stellate hairs below (visible at 10× or higher magnification), giving the lower leaf surface a slightly shiny to almost metallic appearance. {add Lyonia ferruginea and L. fruticosa in ERICACEAE; add Loropetalum in HAMAMELIDACEAE} | |
16 Flowers in spikes, axillary umbels (of shortened spikes), or the flowers solitary and leaf-opposing; leaves spaced, the bases oblique; stipules apparent, and clasping the stem; fruit rarely 3-angled (P. auritum), but not ridged; plants unarmed, the stems swollen at the nodes; branches somewhat zig-zagged, not arranged in conspicuous tiers | |
16 Flowers arranged in axillary spikes only; the fruit usually somewhat ridged; leaves clustered at branch tips (except T. arjuna), the bases typically cuneate; stipules reduced to glandular hairs at petiole base; plants armed or unarmed, the stem nodes not conspicuously swollen; branches arranged in tiers, the main branches erect, the lateral spreading horizontally | |
15 Flowers in other types of inflorescences, not spikes, if the inflorescence axillary only, then consisting of panicles or racemes (Cestrum) or subsessile to sessile fascicles (Myrsine and SAPOTACEAE); fruit various (acorns, berries, drupes, capsules, legumes). | |
17 Leaves 1-foliolate on the upper stems, sometimes 3-foliolate below, or all reduced to phyllodial spines; flowers papilionaceous, bright yellow; fruit a legume; stems bright green | |
17 Leaves simple throughout; flowers variously shaped but not papilionaceous; fruit not a legume, instead acorns, berries, capsules, or drupes. | |
18 Flowers apetalous; arranged in catkins (Quercus in FAGACEAE; Morella inodora in MYRICACEAE) OR thyrses (Dodonaea in SAPINDACEAE). | |
18 Flowers not apetalous, with a well-developed corolla, variously colored (white, cream, pink, greenish or reddish-orange), either urceolate OR tubular and with separate and spreading petals (rarely the perianth only consisting of green sepals), arranged in various terminal or axillary inflorescences, or sometimes solitary; fruit either a (3-) 5-valved capsule, or a spherical berry with (1-) 10+ seeds, OR a 1-8 seeded dry or fleshy drupe. | |
21 Flowers white to pink or reddish-orange, rotate, tubular, or urceolate (the petals also sometimes spreading apically, but united at least basally), in various terminal or axillary inflorescences or solitary; fruit either a 2-5 valved capsule (conspicuously linear-cylindric in CAPPARACEAE), a spherical berry with 10+ seeds OR a drupe bearing 4 bony nutlets (Bourreria). | |
23 Fruit a fleshy berry; inflorescences of axillary fascicles (SAPOTACEAE), axillary and paniculate (Cestrum) or in Solanum, leaf-opposed and variously arranged (terminal, axillary, and extra-axillary). | |
26 Leaves 1 per node or also paired (on one side of the stem) at some nodes (the leaves then uneven in size); inflorescences leaf-opposed; flowers campanulate, lacking a tubular corolla | |
23 Fruit not a fleshy berry, instead a valved capsule or a drupe bearing 4 bony nutlets (Bourreria); inflorescences terminal or axillary (or occasionally flowers solitary), never leaf-opposed. | |
27 Capsules pendulous and conspicuously elongate-cylindric, borne on a slender gynophore (a specialized stipe bearing the gynoecium); seeds white, grey, tan, or brownish in color and usually contrasting with the bright red interior of the capsules; plants shrubs or sometimes loosely scrambling over other plants; stamens usually much longer than the petals and conspicuously exerted from the flowers | |
27 Capsules (or drupes) erect, not linear nor long-cylindric, not borne on a gynophore; seeds variously colored, and sometimes surrounded by a fleshy aril (but the entire capsule interior not bright red); plants shrubs; stamens shorter than or only minimally longer than the corolla (not long exerted) | |
29 Fruit a capsule (the seeds with fleshy arils or attachments); flowers rotate, but not salverform (the corolla tube not lengthened), arranged solitary, few, or in cymes, terminal or axillary, the corolla lobes usually deltoid or somewhat triangular in shape; leaves glabrous or sometimes puberulent (Pittosporum), but the pubescence not rough. | |
30 Flowers few, not showy, green to greenish-white, inflorescence not densely arranged (flowers also sometimes solitary); capsules not beaked; leaves not revolute or undulate | |
30 Flowers numerous, showy, white; inflorescence densely arranged; capsules with short beak; leaf margins revolute or slightly undulate | |
21 Flowers white, rotate, the petals spreading, distinct (i.e., separate to the base; nearly so in Myrsine), not tubular; arranged in axillary fascicles or racemes or in subsessile to sessile axillary fascicles (Myrsine); fruit either a fleshy drupe with 4-8 pyrenes, or a dry to leathery single-seeded drupe. | |
33 Inflorescence an axillary fascicle or cluster, the fascicles short-pedicellate, subsessile, or sessile | |
34 Inflorescence of subsessile or short-pedicellate fascicles, the flowers 4-merous, white (lacking pink streaks or dots); fruit a fleshy drupe with 4-8 pyrenes | |
1 Leaves deciduous. | |
35 Inflorescence branched, spicate, a catkin, or consisting of a solitary flower or axillary clusters or whorls, not an involucrate head. | |
![]() Show caption*© Alan Cressler: Leitneria floridana (male catkin), St. Marks National Wildlife Refuge, St. Marks Unit, Wakulla County, Florida 1 by Alan Cressler | |
38 Inflorescences consisting of solitary, axillary flowers | |
38 Inflorescence of 2 or more flowers; perianth 3-5-merous; fresh plants not musky-fragrant; fruits various, not as above. | |
40 Flowers 3-merous; fruit fleshy, red or greenish-yellow at maturity; ovary superior; [Basal Angiosperms or Eudicots]. | |
42 Fruit either a drupe or berry (indehiscent, and variously fleshy or dry) or a dry 3-valved capsule with 1 seed; inflorescence axillary (solitary, clusters, fascicles, or racemes), or in a terminal raceme (Pyrularia in SANTALACEAE). | |
43 Leaves with various vestiture, but not as above. | |
44 Ovary inferior or half-inferior; inflorescence an axillary cluster or raceme, or a terminal raceme. | |
45 Fruit an elongate drupe (definitely longer than thick), with 1 seed. | |
47 Fruits spherical, < 10 mm long. | |
49 Inflorescence a narrowly cylindrical raceme, clustered several to many at the tip of the previous year’s wood and below the current season’s growth; fruit < 3 mm in diameter | |
50 Fruit fleshy, with 4-8 seeds; leaf pubescence simple or absent. | |
51 Fruit yellow to red, the pedicel 10-30 mm long; leaf venation pinnate, but irregular and reticulated | |
51 Fruit dark red to black, the pedicel < 10 mm long; leaf venation very neatly pinnate, with the secondary veins nearly straight and parallel to one another |
Key G6: trees with alternate, simple, unlobed, entire leaves
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2 Leaves tiny, scale-like, broadest at the base and more or less clasping the stem, < 10 mm long and < 1 mm wide | |
3 Fruit a hesperidium; petiole flanged or winged for most of its length (except C. reticulata, which sometimes lacks wings entirely), constricted at the base of the blade (except linear in C. medica) | |
3 Fruit various (but not a hesperidium); petiole linear (sometimes swollen, but not flanged nor winged with leafy tissue). | |
6 Leaves (when fresh) strongly odorous, glandular-punctate (appearing as translucent dots), with strongly parallel venation; bark on medium-aged to mature trees papery and peeling; fruit a sub-globse capsule (the hypanthium somewhat woody) | |
7 Plants exuding milky sap (when punctured); fruit a berry, these large, subglobose and roughened on the exterior (Manilkara) or smaller and ellipsoid to ovoid in shape (Chrysophyllum); plants (leaves or elsewhere) with rufous, 2-branched hairs (strongly rufous in Chrysophyllum; if leaves strongly glaucous and glabrous, as in Manilkara jamiqui, then leaf apices clearly retuse) | |
10 Flowers solitary, terminal, large (> 5 cm in diameter); pistils many, carpels separate; petals many (typically > 8); leaves mostly > 10 cm long (at least some on a branch longer than 10 cm); fruit an aggregate of follicles, each dehiscing along 1 suture; stipule scar circumferential at each node, encircling the twig | |
10 Flowers axillary or terminal, arranged variously in racemes, panicles, umbels, cymes, fascicles, or sometimes solitary (if so, < 5 cm in diameter); pistil 1, with 1-8 fused carpels; petals 3-8 (apetalous in Conocarpus); leaves < 30 cm long; fruit either a drupe, berry, or capsule; stipule scars either absent or linear or triangular, not circumferentially encircling the twig. | |
11 Inflorescence terminal, the flowers arranged in a corymb, umbel, compound cyme, or raceme (panicle in Pittosporum and sometimes Conocarpus); fruit either a capsule (dehiscing along 1 to 5 longitudinal sutures, elongate in CAPPARACEAE) or a 1-4-seeded drupe (Bourreria; Canella; Lumnitzera). | |
13 Plants never with pneumatophores; leaf blades without pit-domatia. | |
15 Fruit a drupe, the drupe bearing 4 bony nutlets with abaxial ridges; flowers white, rotate and salverform (the corolla tube evident), arranged in terminal cymes, the corolla lobes usually orbicular | |
15 Fruit a capsule (splitting along 1, 3, or 5 sutures); flowers white or pink, rotate but not salverform (lacking an evident lengthened corolla tube), arranged in terminal (or sometimes axillary) corymbs, umbels, or racemes (panicles in Pittosporum). | |
16 Capsules pendulous and conspicuously elongate-cylindric, borne on a slender gynophore; seeds few-many, contrasting sharply with the bright red capsule interior; plants shrubs or small trees, or sometimes loosely scrambling over other plants; stamens usually much longer than the petals and conspicuously exerted from the flowers | |
17 Ovaries 2-carpellate; capsules dehiscing along one major adaxial suture, appearing berry-like before dehiscence, the seeds often surrounded by a glutinous material | |
18 Capsules ovoid to globose or subglobose, about as long as broad, 5-8 mm long; leaves 5-12 cm long, 2-3× as long as wide | |
11 Inflorescence axillary, the flowers arranged in a raceme, panicle, umbel, fascicle, or sometimes flowers solitary; fruit drupaceous, fleshy to dry, but not regularly dehiscent along sutures. | |
19 Flowers bearing petals, the inflorescence variously shaped (flowers occasionally solitary); plants of various inland and coastal habitats. | |
22 Flowers arranged in axillary racemes; leaves clustered apically or spaced; stipules absent or shedding and linear-lanceolate; plants unarmed; branches not arranged in conspicuous tiers. | |
21 Inflorescence either an axillary umbel, cyme, or fascicle (or reduced to solitary) or an axillary compound inflorescence (panicle or compound cyme), with 2-3 orders of branching. | |
24 Fruit a fleshy and oily 1-seeded drupe; flowers 3-merous, with separate and undifferentiated perianth segments; fresh plants strongly aromatic; inflorescence compound, a panicle or compound cyme (with 2-3 orders of branching); [Basal Angiosperms] | |
24 Fruit a fleshy (but not oily) 1-8-seeded drupe, a berry, or a 2-4-locular capsule; flowers 4-8-merous, with differentiated sepals and petals, the petals usually basally fused; fresh plants not strongly aromatic; inflorescence an axillary umbel or fascicle (or reduced to solitary), a central axis absent or < 1 cm long; [Eudicots]. | |
25 Leaves somewhat 2-ranked (subdistichous), the bases of the blades usually oblique; flowers yellowish-green, occasionally present on the trunks (plants cauliflorous), the trunks often fluted (with irregular vertical ridges, at least on larger plants); [c. and s. FL only in our flora area] | |
25 Leaves usually not subdistichous, the bases of the blades not oblique; flowers usually white (Ilex) or white, cream, greenish, or yellowish (Sideroxylon; Tricerma), these never present on trunks, the trunks rounded (not fluted); [collectively widespread, including FL] | |
27 Plants armed with nodal thorns; stamens 5 and staminodia 5, epipetalous; fruit a berry or drupe with 1 seed;flowers 5-merous | |
1 Leaves deciduous. | |
29 Juncture of petiole and leaf blade eglandular, but the uppermost 1-3 mm of the petiole swollen into a prominent upper pulvinus; corolla bilaterally symmetrical, with 5 petals, pink to purple (rarely white in some cultivars); flowers bisexual; fruit an oblong, flat legume, 6-10 cm long; main palmate leaf veins 5-7 (-9) | |
28 Leaf base cuneate, rounded, truncate, subcordate, or auriculate (with 2 small “earlobe-like” lobes at the base of the leaf blade), with 1 (mid) vein from the base (3 veins from the base in Celtis in CANNABACEAE); leaf blade about as wide as long, or somewhat to much longer, 0.9-10× as long as wide. | |
30 Leaves 0.9-1.4× as long as wide (some taxa keyed in both leads). | |
31 Stipule scars circumferential, forming a line around the twig; flowers and aggregate fruits solitary, terminal; [Basal Angiosperms] | |
31 Stipule scars not circumferential (or not apparent); flowers and simple fruits in inflorescences of 1-many flowers, axillary or terminal, but not simultaneously solitary and terminal; [Eudicots]. | |
33 Petioles < 1 cm long; leaves various in shape, often acuminate at the apex and/or cuneate at the base, often with some tendency to toothing; hairs on foliage stellate (use at least 10× magnification), at least in part; fruit either a nut borne in a cup (acorn) or a dry, subglobose 3-valved capsule, with 1 seed. | |
35 Leaves glabrous, glabrescent or variously pubescent (including densely and silkily so, giving the leaf surface a shiny appearance), but not as above. | |
40 Pubescence of the foliage simple or absent (except stellate in STYRACACEAE); flowers bisexual, conspicuous, borne variously, but not in catkins (except in Leitneria); fruit various. | |
![]() Show caption*© Alan Cressler: Leitneria floridana (male catkin), St. Marks National Wildlife Refuge, St. Marks Unit, Wakulla County, Florida 1 by Alan Cressler | |
43 Flowers solitary; ovary superior; perianth either 3-merous and whorled or many-merous and spiraled; leaves mostly > 20 cm long and > 8 cm wide, distinctly broadest towards the apex (> 0.6× of the way from the leaf blade base to apex) (except Magnolia acuminata, which is sometimes both shorter, narrower, and broadest near the middle or towards the base); [Basal Angiosperms]. | |
44 Flowers terminal, > 4 cm across, white, pale yellow, or pink; perianth many-merous, spiraled; fresh foliage not noticeably aromatic; fruit an aggregate of follicles; leaves cuneate or auriculate at the base; twigs with circumferential stipule scars at each node | |
45 Leaves with prominently parallel-arcing secondary veins; inflorescence a terminal corymb; leaves clustered at the tips of the twigs, thus appearing pseudo-whorled; trichomes of the leaf undersurface predominantly 2-branched (some simple) (use at least 10× magnification); flowers 4-merous; fruit a blue drupe; small tree | |
45 Leaves with secondary veins more obscure and complexly branching into tertiary veins; inflorescence axillary (often on the previous year’s wood), solitary to variously fascicled, clustered, or in racemes; leaves arrayed distichously along horizontal or arching twigs, not prominently clustered or pseudo-whorled (except often in Cyrilla in CYRILLACEAE, Symplocos in SYMPLOCACEAE, and Nyssa in NYSSACEAE); trichomes of the leaf undersurface either simple or stellate (or absent); flowers 4-5-merous; fruit a green, blue, or black drupe, an orange berry, or a green to brownish indehiscent capsule; small to large tree. | |
46 Pubescence of foliage and other parts stellate (use at least 10× magnification); petals 4-5, white, 10-25 mm long; fruit dryish, indehiscent, either longitudinally 2-4-winged or not winged | |
46 Pubescence of foliage and other parts simple; petals either 0, or 4-5 and pink, white, or greenish-yellow, or 10 and greenish-yellow; fruit either a somewhat to very fleshy drupe or berry or a dry, brownish, spherical drupe, 2-2.5 mm in diameter. | |
47 Leaves < 2.5 cm wide, dark green above, somewhat thickened, and tardily deciduous or semi-evergreen; fruit a dry, brownish, spherical drupe, 2-2.5 mm in diameter; inflorescence a narrowly cylindrical raceme with > 40 flowers | |
47 Leaves > 2.5 cm wide, usually medium-green above, herbaceous in texture, promptly seasonally deciduous; fruit a somewhat to very fleshy drupe or berry, > 5 mm in diameter; inflorescence a solitary flower or cluster, head, or irregular raceme of < 15 flowers. | |
48 Fruit a drupe (green when ripe), cylindrical to barrel-shaped, 8-12 mm long; leaves rather thick and leathery in texture, persistent into the winter, dropping tardily or at latest the following spring; flowers bisexual; stamens 30-50, in 5 fascicles | |
49 Fruit a spherical berry, 15-50 mm long, orange when ripe, subtended by the enlarged and persistent woody or leathery calyx; vascular bundles 1 per leaf scar; leaves never toothed; leaves whitish-green beneath; leaf midrib and upper petiole with tiny glands on their upper surfaces (reddish initially, then darkening) (use at least 10× magnification); leaves glabrate to tomentose with curly hairs beneath; female and male flowers on separate trees (dioecious); stamens 16; widest point of the leaf usually at the middle or below, the apex acute to acuminate | |
49 Fruit an ovoid or ellipsoid drupe, 8-30 -40 mm long, blue-black, yellow, orange, or red when ripe; vascular bundles 3 per leaf scar; leaves sometimes bearing a few irregular teeth; leaves pale to medium green beneath; leaf midrib and upper petiole lacking reddish to dark glands on their upper surfaces; leaves glabrous or glabrate beneath; female and male flowers on the same tree (monoecious); stamens 5-12; widest point of the leaf usually beyond or at the middle, the apex obtuse to strikingly and abruptly acuminate |