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Key to Digitaria

Poaceae

Digitaria

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1 Inflorescence a simple open panicle, with well-developed primary and secondary branches, the entire panicle usually detaching at maturity; spikelets solitary (or in pairs or triplets), on short to long pedicels, the pedicels mostly >2x as long as the spikelets, mostly spreading (sometimes distally appressed on branches); [Leptoloma].
  2 Spikelets 3.5-4.6 mm long; upper glumes 5-7 veined; anthers > 1 mm long; [TX coastal dunes]
  2 Spikelets 2.2-3.3 mm long; upper glumes 3 (-5)-veined; anthers < 1 mm long; [collectively widespread, of various habitats].
    3 Lower lemmas 7-veined; outer veins not equally spaced; spikelets glabrous to short pubescent; [widespread]
    3 Lower lemmas 5-veined; veins equally spaced; spikelets densely pubescent; [e. OK and e. TX westwards].
      4 Plants with long rhizomes, loosely cespitose or loosely tufted
1 Inflorescences digitate or sub-digitate, or with primary branches alternately arranged along a central axis; with (1) 2-3+ spikelets per node on primary branches, arranged in two rows along one side of the branch (and usually appressed on branches); the entire inflorescence not detaching at maturity.
        5 Inflorescences of primary branches alternately arranged along a central axis; all the rachilla internodes more or less conspicuously elongated, hence upper florets stipitate; spikelets 3.0-6.6 mm long, the lower lemmas and upper glumes with conspicuously long marginal hairs, these generally spreading and at least ½ as long as the spikelet at maturity); upper lemmas dark brown at maturity; [Trichachne].
          6 Lower lemma with 5 equally spaced nerves on the back
          6 Lower lemma with 3 nerves on the back, evenly spaced or not.
             7 Spikelets (3.7-) 4-6.6 mm long, with ochraceous indumentum, narrowly ellipsoid, apex acuminate; lower lemma glabrous on both sides of midnerve and either pilose or alternately pilose and glabrous in the remaining zones
             7 Spikelets 3-4.5 mm long, with hairs whitish-silvery or whitish with purplish tints, narrowly ovate or ovate, apex acuminate or abruptly acuminate; lower lemma glabrous on both sides of the midnerve and pilose in the remaining zones.
               8 Spikelets 3-4.5 mm long, ovate, apex abruptly acuminate; panicle 5-19 cm long, with 4-10 appressed racemes, densely spiculate, on an axis 5-7 cm long
               8 Spikelets 3-3.2 mm long, ellipsoid or linear-lanceolate, apex acuminate; panicle 10-18 cm long, with 6-9 patent racemes, loosely spiculate, on an axis 7-10 cm long
        5 Inflorescences digitate, sub-digitate, or with alternately arranged primary branches; the rachilla internodes not conspicuously elongated, thus the upper florets not stipitate; lower lemma and upper glume without conspicuous marginal pubescence (or if with hairs, these not ca. ½ as long as the lower lemma and upper glume); spikelets 1.2-4.1 mm long; upper lemmas variously colored at maturity (including dark brown); [Digitaria].
                 9 Inflorescence branches (each "finger") not at all winged, the axes usually triquetrous only (with 3 distinct angles); upper glumes 3-veined.
                   10 Spikelets glabrous or nearly so (sometimes the upper glume apices sparsely pubescent); [FL only in our area].
                     11 Spikelets 2.3-2.4 mm long; leaf blades 2.0-4.5 mm wide; sheaths and leaf blades densely hairy with papillose-based hairs (but not densely gray-villous); plants erect, unbranched, and not rooting at lower nodes
                     11 Spikelets 2.7-3.2 mm long; leaf blades 2.0-2.2 mm wide; sheaths and leaf blades densely gray-villous on both surfaces; plants erect to somewhat decumbent, not rooting at lower nodes but often branching at aerial nodes
                   10 Spikelets pubescent (at least the lower lemmas), often with gland-tipped hairs; the upper glumes [collectively widespread, including FL].
                       12 Basal leaf sheaths variously pubescent; cauline leaf blades 1-6 mm wide; spikelets 1.3-1.9 (-2.2) mm long (D. filiformis) or 2.0-2.8 mm long.
                          13 Spikelets 1.3-1.9 (-2.2) mm long; panicle branches 3-13 cm long; culms 10-80 cm tall
                          13 Spikelets 2.0-2.8 mm long; panicle branches 10-25 cm long; culms 75-100 cm tall (to 150 cm tall in D. villosa).
                            14 Basal and lower leaf sheaths appressed pubescent, but the hairs not papillose-based; plants cespitose perennials; spikelets 2.2-2.5 mm long; ligules 2.0-3.0 mm long; culms to 100 cm tall; [endemic to Lake County, FL]
                            14 Basal and lower leaf sheaths with spreading, papillose-based hairs; spikelets 2.0-2.8 mm long; panicle branches 10-25 cm long; ligules 0.3-1.5 mm long; culms 75-150 cm tall; [widespread]
                 9 Inflorescence branches not merely triquetrous, instead each "finger" at least minimally winged (< ½ as wide as midribs) or widened (> ½ as wide as midribs); upper glumes 3-7 veined.
                              15 Basal and lower sheaths glabrous or nearly so (D. nuda sheaths mostly glabrous, sometimes with long hairs basally, but the leaf blades also primarily glabrous).
                                16 Spikelets 1.2-1.7 mm long; lower glumes absent (or reduced to veinless, membranous rim); wings of the rachis at least ½ was wide or wider than the midribs
                                  17 Plants conspicuously stoloniferous, the stolons long and branching; leaves few (3-4), clustered at the base of plants, the blades 1.5-4.0 cm long; panicles usually with 2 widely divergent primary spike-like branches (occasionally 3 or 4, these extra "fingers" often reduced in size)
                                  17 Plants lacking stolons; leaves few or numerous, the blades (1.5-) 4.0-9.0 cm long, if clustered at the base of plants, usually at least some blades > 4 cm long; panicles with 2-7 primary spike-like branches (often at least 3 per inflorescence).
                                    18 Culms decumbent and rooting at the nodes; spikelets lanceolate, 1.5-1.7 mm long; panicles with 2-4 spike-like primary branches, the inflorescence racemose (the "fingers" not in evident verticils); [endemic to dry sandy habitats in Hernando and Marion counties, FL]
                                    18 Culms erect, usually not branching from upper nodes; spikelets narrowly elliptic, 1.2-1.7 mm long; panicles with 2-7 spike-like branches, once or twice verticillate; [non-native of disturbed habitats, widespread in our flora area]
                                16 Spikelets 1.7-3.5 mm long; lower glumes present or absent; wings narrow or widened, relative to width of the midrib.
                                       19 Spikelets 2.5-3.5 mm long; plants perennial; culms 50-250 cm long, erect or decumbent, rooting or not at the lower nodes; [waif]
                                       19 Spikelets 1.7-2.8 mm long; plants primarily annuals; culms 20-70 cm long, decumbent, rooting and branching at the lower nodes.
                                         20 Plants always with axillary panicles in the lower leaf sheaths (these sometimes concealed by the sheaths); panicles bearing spikelets in groups of 3; lower portions of the pedicels adnate to the axes; leaf blades 3.0-5.0 mm wide; sheaths sometimes sparsely pubescent; [widespread non-native]
                                         20 Plants never with axillary panicles concealed in the lower leaf sheaths; panicles bearing spikelets in unequally pedicellate pairs (at least on lower and middle portions of the inflorescence); leaf blades 1.5-2.5 mm wide; sheaths mostly glabrous, sometimes with long scattered hairs basally only if not entirely glabrous; [waif]
                                             22 Spikelets glabrous, 1.7-2.3 mm long, the upper glumes < ½ as long as the spikelets; ligules fimbriate; blades scabrous adaxially and sometimes with sparse hairs basally, but otherwise mostly glabrous (abaxially); plants densely tufted perennials
                                             22 Spikelets variously pubescent, 2.0-3.6 mm long (only obscurely pubescent in D. simpsonii, but spikelets at least 3.0 mm long), the upper glumes ca. as long as the spikelets; ligules not fimbriate; blades pilose or hirsute (sometimes nearly glabrous in D. texana); plants usually decumbent and branching at lower nodes (D. simpsonii and D. texana) OR if the plants tufted and erect, then spikelets 2.4-2.8 mm long (D. subcalva).
                                               23 Spikelets 2.0-2.8 (-3.6) mm long, the upper glumes 3-5 veined (D. texana) or 5-7 veined (D. subcalva); plants perennial.
                                                 24 Spikelets 2.4-2.8 mm long; primary inflorescence branches 2-4, the inflorescence lax, diffuse, and gray to purplish-gray; pedicels on upper paired spikelets longer (2.5-3.0 mm); leaf sheaths strongly papillose-hispid; [endemic to wet, fringing calcareous prairies, peninsular FL]
                                                 24 Spikelets 2.0-3.6 mm long; primary inflorescence branches 5-12, the inflorescence dense and mostly greenish; pedicels on upper paired spikelets shorter (1.0-2.5 mm long); leaf sheaths weakly papillose-hispid; [endemic to coastal prairies of TX, adventive in coastal VA and SC]
                                                      26 Spikelets 1.5-1.8 mm long, villous with crinkled hairs; lower portions of the panicles with spikelets in triplets; leaf blades conspicuously pubescent on both surfaces, the longer hairs 1.5-2.5 mm long and papillose-based
                                                      26 Spikelets (1.7-) 2.1-3.4 long, variously pubescent; lower portions of the panicles with spikelets in unequally pedicellate pairs; leaf blades pilose or pubescent, the hairs usually < 1.5 mm long
                                                        27 Spikelets 2.1-2.4; sheaths with scattered papillose-based hairs, ligules erose; the inflorescences with the lower most branches whorled (the upper branches paired or solitary; on depauperate inflorescences merely 2-3 branches opposite or whorled); [primarily c. and s. FL in our area, a waif in MS, AL, and SC]
                                                        27 Spikelets (1.7-) 2.5-3.4 mm long, averaging 3.0 mm; sheaths keeled, ligules not erose; plants annual, decumbent and rooting at the nodes; leaf blades pilose on the upper surface; the inflorescence subdigitate, lacking whorled branches below; [widespread in the US]
                                                          28 Plants perennial and usually stoloniferous, sometimes also rhizomatous; culms erect or decumbent, not rooting at the lower nodes (except D. milanjiana which does or does not root); [uncommon non-natives].
                                                            29 Midveins of the lower lemmas smooth throughout; basal sheaths glabrous or often densely pubescent (the hairs 4-6 mm long); spikelets wooly pubescent, the hairs of variable length (sometimes > 1 mm long); rhizomes sometimes present, if so, these giving the plant bases a knotty appearance; [throughout FL]
                                                            29 Midveins of the lower lemmas scabrous (at least distally); basal sheaths glabrous, sometimes pubescent (if so, the hairs shorter); spikelets often at least somewhat pubescent, but the hairs never conspicuously long (< 1 mm long); [c. FL]
                                                          28 Plants annual or nearly so, lacking stolons and never rhizomatous; culms erect but also often long-decumbent and usually rooting at the lower nodes; [natives and non-natives, collectively widespread].
                                                              30 Spikelet pairs homomorphic, the lower lemmas of the upper and lower spikelets not conspicuously differing in their pubescence types; lower lemmas of the sessile spikelet with lateral veins crowded towards the margins (well separated from the midvein).
                                                                   32 Spikelets 2.7-4.1 mm long, the lower glumes 0.2-0.8 mm long; lateral veins of the lower lemmas usually glabrous; leaf blades usually with a few scattered papillose-based hairs at the base of the upper surfaces (occasionally throughout the upper surface); [weedy native]
                                                                   32 Spikelets 1.7-3.4 mm long; the lower glumes 0.2-0.4 mm long; lateral veins of the lower lemmas scabrous for the distal 2/3 of their length; leaf blades usually with papillose-based hairs on both surfaces (sometimes glabrous); [weedy non-native]