X
Jump to key:

Click the number at the start of a key lead to highlight both that lead and its corresponding lead. Click again to show only the two highlighted leads. Click a third time to return to the full key with the selected leads still highlighted.

Key to Asteraceae

Asteraceae

Copy permalink to share

1 Plant a shrub or liana (woody vine), definitely with woody growth well above ground level
1 Plant an annual, biennial, or perennial herb, lacking woody growth above ground level (or suffrutescent at the base).
  2 Leaves opposite or whorled, at least on the lower stem nodes (the leaves higher on the stem sometimes alternate).
    3 Heads discoid or disciform, with disk florets only
    3 Heads radiate, with both ray and disk florets present.
  2 Leaves either alternate (not opposite even at lower nodes of the stem) or basal only (the heads on scapiform stems).
      4 Heads liguliflorous (composed of ligulate florets, the laminae flattened and strap-like); plant sap usually milky
      4 Heads not liguliflorous (the heads instead composed of disc florets (discoid, disciform) and/or ray florets (radiant, or radiate); plant sap usually clear.
        5 Heads borne solitary on scapes, or many in a raceme on stems with only scale-like bracts.
        5 Heads borne on stems with leaves, the leaves alternate, opposite, or whorled on the stem, and sometimes also well developed basal leaves or rosettes.
          6 Heads discoid or disciform (composed primarily of disc florets and peripheral filiform florets), OR heads radiant, essentially composed of ordinary disc florets and dilated periphereal florets (these peripheral florets dilated, perfect, pistllate, or netural, but lacking broadly expanded laminae of ordinary ray florets).
             7 Leaves spiny-margined, phyllaries usually spine-tipped; disk flowers pink (rarely blue or yellow)
             7 Leaves not spiny-margined; phyllaries spine-tipped or not; disk flowers variously colored (including pink)
               8 Pappus none or rudimentary [look at summit of achenes or ovary]
               8 Pappus well-developed, of bristles, awns, scales, or a combination of these.
                 9 Pappus of bristles only (the bristles smooth, barbellulate, barbellate, or plumose)
                 9 Pappus wholly or partially of scales (broad to narrow, sometimes almost bristle-like, but at least in part flattened) or awns (these often barbed)
          6 Heads radiate, with both central disc and peripheral ray florets, the ray florets with prominent laminae.
                   10 Rays yellow, cream, orange, or rarely red-tinged.
                     11 Receptacles paleate (with chaff) [best seen by removing flowers or achenes from the receptacle]
                     11 Receptacles naked (epaleate), rarely with imbedded bristles (or Gaillardia sometimes with short setae 1-6 mm long but these much shorter than the pappus scales of the cypselae)
                   10 Rays white, pink, purple, or bluish.
                       12 Pappus none or rudimentary [look at summit of achenes or ovary]
                       12 Pappus well-developed, of bristles, awns, scales, or a combination of these.
                          13 Pappus of bristles only (the bristles smooth, barbellulate, barbellate, or plumose)
                          13 Pappus wholly or partially of scales (broad to narrow, sometimes almost bristle-like, but at least in part flattened) or awns (these often barbed)

Key to Asteraceae, Key A: woody composites (shrubs and lianas)

Copy permalink to share | Check for keys that lead to this key

1 Leaves opposite, at least on the lower stem nodes (the leaves higher on the stem sometimes alternate); [tribe Heliantheae or tribe Eupatorieae].
  2 Heads radiate, the rays yellow to orange; disc flowers yellow to orange.
    3 Leaves linear; receptacle epaleate, but densely setose; [extreme s. TX]
      4 Leaves mostly entire, rarely distally 3-lobed, often somewhat glaucous; disc florets 30-100+; receptacle densely setose; pappus of scales
      4 Leaves usually shallowly to deeply 3-9-lobed, green; Disc florets 100+; receptacle with oblong-rectangular paleae; leaves linear or linear-lobed; pappus absent
    3 Leaves lanceolate or broader, 1-5× as long as wide; receptacle paleate; heads borne singly or many; [collectively widespread].
        5 Leaves ovate, rhombic, or pentagonal, unlobed or 3-5-(-7) lobed; leaf blade margins toothed; leaves in part alternate upwards; [of disturbed, suburban areas of FL peninsula]; [tribe Heliantheae; subtribe Helianthinae]
          6 Ray laminae 5-16 mm long; leaf blades 7-33 cm long, the larger deeply 3-7-lobed; petiole bases dilated at base, fused to form a disc of leaf tissue at the stem
          6 Ray laminae 45-70 mm long; leaf blades 2-8 cm long, unlobed or 3-lobed; petiole bases not dilated at base
        5 Leaves lanceolate to oblanceolate, unlobed or obscurely 3-lobed; leaf blade margins toothed or entire; leaves strictly opposite; [tribe Heliantheae; subtribe Ecliptinae].
             7 Leaves obovate to oblanceolate, unlobed, entire; heads (5-) 8-13 (-18+) mm in diameter; cypselas 3-4-smgled, not winged; [of tidally inundated salt and brackish marshes of the outer Coastal Plain, MD to s. FL, west to s. TX]
             7 Leaves lanceolate, sometimes 3-lobed, usually serrate but sometimes nearly entire; heads 4-8 mm in diameter; cypselas flattened and winged; [TX only]
  2 Heads discoid or disciform (ray flowers lacking); disc flowers pink, purple, whitish, yellow, or orange.
               8 Disc flowers yellow to orange
               8 Disc flowers pink, purple, or white.
                 9 Heads small (involucres 2-8 mm high), solitary, axillary in the axils of leaves or leafy bracts (similar to the leaves but smaller) or interpretable as arrayed in bracteate racemes; heads nodding, the involucre 2-7 mm high; [collectively widespread in coastal portions of our area, of maritime situations]; [tribe Heliantheae; subtribe Ambrosiinae]
 Iva
                 9 Heads small to larger (involucres 3-15+ mm high), many, terminal on the branches of corymbiform or paniculiform arrays; heads erect, the involucre 4-12 mm high; [of Coastal Plain of FL, s. GA, and TX].
                   10 Pappus of 4-10 scales, 0.3-7 mm long; [tribe Heliantheae; subtribe Chaenactidinae]
                   10 Pappus of 5-70 barbellate bristles, 3-12 mm long; [tribe Eupatoriae]
                     11 Leaf blades spatulate to obovate, cuneate to attenuate at the base, rounded or shallowly notched at the apex; leaf margins entire; leaves opposite below, alternate higher on the stems; [native of FL]
                     11 Leaf blades triangular or lanceolate, truncate, cordate, or cuneate at the base, acute to acuminate at the apex; leaf margins serrate; leaves opposite throughout.
                       12 Phyllaries broad, 2-3× as long as wide, the inner phyllaries translucent-scarious white to pinkish at their broadly rounded apices, falling as the head goes to fruit
                       12 Phyllaries narrow, 4-7× as long as wide, the inner phyllaries green at their acute to acuminate apices, persistent on the head in age.
                          13 Involucres 2-3 mm in diameter; phyllaries 7-16 in 1-2 series; disc flowers 3-13 per head; [s. FL only (in our region)]
                          13 Involucres 3-7 mm in diameter; phyllaries 25-35 in 2-3 series; disc flowers 10-60 per head; [TX only (in our region)].
                            14 Disc flowers white (to light pink); [c. TX]
                            14 Disc flowers blue to lavender (rarely white); [extreme s. TX]
1 Leaves strictly alternate.
                              15 Heads radiate (with ray and disc flowers) or pseudoradiate (with pseudoradiate flowers only.
                                16 Rays white, pink, or rose.
                                  17 Rays pink or rose, 30-50 (-70); sprawling viny shrub to 40 dm tall; leaves ovate, averaging ca. 3× as long as wide, with clasping base, with a midvein and lateral veins and tertiary reticulation
                                  17 Rays white, 5-34+; shrubs 2-30 dm tall; leaves either obovate to oblanceolate, averaging ca. 4× as long as wide, dentate, or linear, ca. 10× as long as wide, entire, cuneate.
                                    18 Rays white, 5-11 (-15); leaves linear, ca. 10× as long as wide, entire; [native, s. and se. TX]
                                    18 Rays white, 21-34+; leaves obovate to oblanceolate, averaging ca. 4× as long as wide, dentate; [non-native, known in our region only from NY]
                                16 Rays or pseudorays yellow, orange-yellow, or light creamy yellow.
                                       19 Heads pseudoradiate, with 11-25 pseudoray flowers and 0 disc flowers; leaves 8-70 mm wide
                                       19 Heads radiate, with (0-) 1-11 (-15) ray flowers and 1-15 (-20) disc flowers; leaves (or their segments) 0.5-10 mm wide.
                                         20 Phyllaries in 1-2 series and equal or subequal in length to one another, the phyllaries often coherent/fused (the involucre thus appearing as a cylinder with ribs extending from base to top; calyculus (of bracts subtending the phyllaries and differing from them in texture, color, or orientation) present; [tribe Senecioneae]
                                         20 Phyllaries in 3-17 series and unequal in length (imbricated); calyculus absent.
                                           21 Rays light creamy yellow, 5-11 (-15); involucre 3-5 (-7) mm in diameter
                                           21 Rays bright yellow, (0-) 1-9; involucre 1-2.5 (-3) mm in diameter.
                                             22 Pappus of 40-60 barbellate bristles; ray blades 4-6 mm long; leaf surfaces minutely pebbled; intricately branched shrubs to 1 m tall, with persistent sterile shoots with evergreen leaves and annual flowering shoots (the entire shoot and leaves deciduous); [Coastal Plain, se. NC south to Panhandle FL and s. MS]
                                             22 Pappus absent, a low coroniform structure, or of scales; ray blades 2-3.5 mm long; leaf surfaces planar/revolute; shrubs 1-15 dm tall, not differentiated into evergreen, sterile shoots bearing deciduous, fertile shoots; [OK and TX westwards, very rarely as a waif eastwards].
                                               23 Pappus of 5-10 elongate scales
                                               23 Pappus absent or a low (< 0.1 mm high) crown
                              15 Heads discoid (with disc flowers only).
                                                 24 Pappus absent; leaves 1-2× pinnately or 1× palmately lobed.
                                                    25 Leaves either palmately 3-lobed, or pinnately lobed and the few (<11) lobes > 2 mm long
                                                    25 Leaves 1-pinnately lobed, the many (>21) lobes mostly < 1.5 mm long and wide
                                                 24 Pappus of bristles; leaves unlobed (the margins entire or serrate).
                                                      26 Plants dioecious, either male or female; heads either staminate, with 10-50 flowers, or pistillate, with 20-150 flowers
                                                      26 Plants hermaphroditic; heads bisexual, with 5-25 flowers, either all perfect (bisexual), or a mixture of perfect (bisexual) and functionally staminate flowers.
                                                        27 Leaves strongly basally disposed, crowded in the lowermost, woody or semiwoody portion of the stem
                                                        27 Leaves on the stem.
                                                          28 Leaves lanceolate to ovate (broadest below the midpoint); leaf margins entire to coarsely toothed.
                                                            29 Heads in leafy, paniculate arrays; leaf blades coarsely dentate, 0.5-4 cm long; shrubs to 12 dm tall; [c. and e. TX]
                                                            29 Heads borne in leafless, corymbiform arrays; leaf blades entire to denticulate, 5-16 (-20) cm long; shrubs to 40 dm tall; [FL and s. TX]
                                                          28 Leaves obovate, spatulate, or orbiculate (with rounded apex) or elliptic (with acute to acuminate apex), broadest at or beyond the midpoint; leaf margins entire.
                                                              30 Leaf blades obovate, spatulate, or orbiculate, rounded or slightly notched at the apex; leaf blade surfaces gray-green on both surfaces, granular when young; disc flowers 5 per head; [FL]
                                                              30 Leaf blades elliptic, acute to acuminate at the apex; leaf blade strongly bicolored above and below, the upper surface dark green and glabrate, the lower surface densely and finely tomentose and strongly whitened, the upper surface dark green and glabrate or glabrous; disc flowers 4-25 per head; [s. TX]

Key to Asteraceae, Key B: herbaceous composites with opposite or whorled leaves
and discoid or disciform heads (lacking ray florets)

Copy permalink to share | Check for keys that lead to this key

1 Pappus present, of 5-60 barbellate bristles; receptacle naked (without paleae or well-developed bristles); [tribe Heliantheae; subtribe Eupatoriae].
  2 Plant a twining herb, phyllaries and disk florets 4 per head
  2 Plant stiffly erect to sprawling but never twining, phyllaries and disk florets usually > 4 per head.
    3 Leaves in whorls of 3-7, > 2 cm wide
    3 Leaves opposite, rarely alternate or whorled, if whorled, < 2 cm wide.
      4 Cypselae rostrate (with long beaks); the achene portion not bearing ribs
      4 Cypselae not beaked, the achene variously ribbed.
        5 Achenes (and ovaries) 8-10-ribbed; outer phyllaries longitudinally striate
        5 Achenes (and ovaries) (3-) 4-5-ribbed; outer phyllaries not noticeably longitudinally striate.
          6 Heads pink to blue.
             7 Phyllaries in 4-6+ series, deciduous
             7 Phyllaries in 2-4 series, persistent (or deciduous in Praxelis).
               8 Heads pink (rarely bluish); receptacles flat; florets 18-25 per head; leaves sometimes gland-dotted
               8 Heads blue; receptacles conic; florets 25-70 per head; leaves gland-dotted.
                 9 Bristles ca. 30; florets 35-70+; phyllaries ca. 25, subequal, persistent
                 9 Bristles ca. 40; florets 25-30; phyllaries 15-25, unequal, deciduous (absence apparent in fruit)
          6 Heads white to cream or, rarely, pale lilac.
                   10 Florets 3-7 per head
                   10 Florets at least 9 per head.
                     11 Phyllaries not strongly imbricate, with the principal ones subequal and sub-biseriate; petioles 0.5-10 cm
                     11 Phyllaries clearly imbricate, in 3+ series, the margins usually glandular; some species epetiolate
1 Pappus either absent, or of scales, setae, or awns; receptacle naked, paleate, or bearing bristles.
                       12 Leaves whorled, linear, < 2 mm wide; head solitary; [aquatic herb growing in shallow stagnant water]
                       12 Leaves opposite (or alternate in part), broader in shape and > 5 mm wide; heads typically not solitary; [terrestrial or wetland plants].
                          13 Receptacles naked (epaleate).
                            14 Pappus of 2-12 scales; heads white, pink, or blue (yellow in Schkuhria).
                              15 Heads with 20-125 florets; leaves serrate or ± dentate (at least distally); [rarely naturalized exotics or s. TX/FL natives]; [tribe Eupatorieae].
                                16 Leaves petiolate; corolla tube glabrous, typically blue or lavender colored (sometimes white tinged); heads with 20-125 florets; [naturalized exotics or s. FL native]
                                16 Leaves sessile; corolla tube glandular-pubescent, typically white colored (sometimes pink-purple tinged); heads with 75-125 florets; [s. TX native]
                              15 Heads with 10-30 florets (or fewer in Schkuhria); leaves entire; [natives, of FL, s. GA, MS, and TX in our area]; [tribe Heliantheae; subtribe Chaenactidinae].
                                  17 Leaves 3-5 lobed (or foliolate); cypselae with curled hairs (Florestina tripteris) or villous on angles (Schkuhria pinnata); [s. TX in our area or ne. waif].
                                    18 Heads discoid, corolla white; cypselae with curled hairs; [s. TX in our area]
                                    18 Heads discoid, sometimes with 1 (-2) diminuitive rays, corolla (of discs) yellow; cypselae villous, at least at bases or on angles; [waif of ne US, native in sw. US]
                                  17 Leaves simple or unlobed; cypselae with straight hairs; [variously distributed in s. GA and FL w. to MO and s. TX]
                            14 Pappus none, or of a few bristles or irregular coroniform lobes; heads green to yellow.
                                       19 Heads bright yellow, closely aggregated into corymbiform arrays of flat-topped to dome-shaped glomerules
                                       19 Heads greenish or greenish yellow, borne in spiciform, racemiform, or paniculiform arrays.
                                         20 Heads in paniculiform arrays, all or most of the heads not subtended by bracts
                                         20 Heads in spiciform or racemiform arrays, each 1 (-2) heads subtended by a bract
 Iva
                          13 Receptacles with paleae or well-developed bristles (the pales sometimes modified into specialized perigynia or burs surrounding the achene).
                                           21 Involucre of four decussate, deltoid bracts (the outer two large and longer than and often enclosing the flowering heads); pappus absent; paleae tightly enclosing the cypselae; plants rooting at the nodes, sometimes free-floating in water; [uncommon non-native, c. FL]
                                           21 Involucre not of four decussate, deltoid bracts; plants not with the above combination of characters; [widespread natives and non-natives]
                                             22 Heads small, less than 1 cm in diameter at anthesis (the female heads enlarging in Xanthium); disc florets dull white or suffused with green or purple; florets mainly unisexual (either in the same heads and then males central and females peripheral, or in separate female and male heads); female florets 0-8 per head; [tribe Heliantheae; subtribe Ambrosiinae].
                                               23 Heads unisexual; cypselas shed within an indurated bur or “nut” with hooked or straight spines developed from the phyllaries and/or paleae.
                                                 24 Involucre of the female heads with tubercles or straight spines developing from the phyllaries; burs 1-8 mm long
                                                 24 Involucre of the female heads with hooked spines developing from the phyllaries/paleae; burs 10-35 mm long
                                               23 Heads bisexual, with functionally male and female flowers in the same head; cypselas shed individually, not enclosed.
                                                    25 Heads in paniculiform arrays, all or most of the heads not subtended by bracts
                                                    25 Heads in spiciform or racemiform arrays, each 1 (-2) heads subtended by a bract
 Iva
                                             22 Heads larger, mostly > 1 cm in diameter at anthesis; disc florets conspicuously white, yellow, pale yellow, or purple; florets mainly bisexual; female florets > 12 per head (except 2-8 in Polymnia and 8-15 in Verbesina occidentalis).
                                                      26 Leaves strongly basally disposed, the basal-most leaves sitting almost flat on the ground and overlapping; disc flowers maroon-purple
                                                      26 Leaves mainly or strictly cauline (not basally disposed); disc flowers white, yellow, or reddish-brown.
                                                        27 Disc flowers bright yellow or reddish-brown.
                                                          28 Involucre of phyllaries subtended by a calyculus of bracts obviously different in color, texture, and shape than the phyllaries.
                                                            29 Phyllaries usually distinct, rarely connate proximally; disk florets bright yellow
                                                            29 Phyllaries connate (both proximally-distally connate); disk florets yellow to reddish-brown
                                                          28 Involucre of phyllaries not subtended by a calyculus (phyllaries sometimes in 2+ series, but without obvious calyculi below).
                                                              30 Plants erect; heads without ray florets; [wet pine savannas and bogs; FL panhandle]
                                                              30 Plants primarily prostrate or trailing, sometimes erect; heads with or without ray florets; [habitats various, widespread natives and non-natives]
                                                        27 Disc flowers bright white or pale yellow.
                                                                 31 Stems not winged; plants erect or spreading.
                                                                   32 Cypselae 3-4 angled; pappus falling (of 2-12 awns or bristles); leaf margins irregularly crenate to serrate, the surfaces sometimes roughened but lacking glands; [e. of TX]
                                                                   32 Cypselae 5-ribbed; pappus absent; leaf margins entire or serrulate, the surfaces gland-dotted; [s. TX]
                                                                 31 Stems winged; plants erect.
                                                                     33 Disc flowers bright white; leaves unlobed or with 2 basal lobes
                                                                     33 Disc flowers creamy or pale yellow; leaves prominently pinnately lobed, with 5-many lobes

Key to Asteraceae, Key C: herbaceous composites with opposite leaves and radiate heads

Copy permalink to share | Check for keys that lead to this key

1 Ray florets white, pink, or purple (rarely pale yellow or lavender).
  2 Ray floret (the lamina) persistent on the achene and becoming papery and bleached
  2 Ray floret (the lamina) articulate from the achene and falling, thus the mature heads not appearing papery.
    3 Pappus of a minute crown (coroniform); disc florets white or whitish.
    3 Pappus not only coroniform, either absent or variously composed of scales, awns, or plumose bristles (Galinsogeopsis has 2 antrorsely barbed bristles plus a crowns of laciniate scales); disk florets yellow, red-brown, or pinkish.
      4 Heads with an involucre not subtended by a calyculus.
        5 Rays pink, the laminae 5-14 mm long; [OK and TX westward; subtribe Cheaenactidinae]
        5 Rays white or whitish-yellow; the laminae shorter, typically < 5 mm long (occasionally longer in Polymnia).
          6 Plants larger, 50-150+ cm tall; perennials; leaves > 10 cm long, sometimes with winged petioles or clasping basal appendages; disc florets functionally staminate; [natives of higher-quality, limestone or novaculite habitats; subtribe Polymniinae]
          6 Plants smaller, 2-40 cm tall or long (if stems > 40 cm then plants usually procumbent or decumbent); annuals or perennials; leaves usually < 10 cm long, lacking winged petioles or clasping basal appendages; disc florets bisexual and fertile.
             7 Stems usually copiously glandular-pubescent; leaf blades cordate to subdeltate or usually strongly 3-lobed (but not pinnate), the margins singly or doubly crenate; disc florets 40-100+; plants erect to decumbent; [native, s. TX westward; subtribe Peritylinae]
             7 Stems not copiously glandular-pubescent; leaf blades lanceolate to broadly ovate or deltate, or pinnately to palmately lobed (Tridax); plants erect or ascending to procumbent (Tridax]; [non-natives of disturbed habitats, widespread; subtribe Galinsoginae].
               8 Plants with leaves simple, unlobed; pappus either absent or of 5-20 laciniate or fimbriate scales; plants annual, erect
               8 Plants usually with a mix of simple and 3-lobed leaves; pappus usually of ca. 20 plumose (or setiform) scales; plants perennial, usually at least somewhat procumbent
      4 Heads with an involucre subtended by a calyculus of bracts (these often but not always reflexed); the phyllaries often appearing somewhat translucent or of a distinctly different color, shape, or texture from the leafy colored bracts below; [tribe Heliantheae; subtribe Coreopsidinae].
                 9 Phyllaries connate for at least ¼ their length; heads with or without ray florets; [MS westwards in our area]
                 9 Phyllaries distinct; heads with ray florets; [collectively widespread in our area].
                   10 Cypselas beaked, not strongly flattened, 7-30 mm long, with 1 groove per face; leaves highly dissected
                   10 Cypselas beakless, more or less strongly flattened and also often winged, 1.2-16 mm long, with 0 or 2 grooves per face; leaves simple to highly dissected.
                     11 Cypselas 2.5-16 mm long, usually not winged; pappus awns (if present) usually retrorsely barbed; rays white
                     11 Cypselas 1.2-4 mm long, winged (C. nudata) or not (C. rosea); pappus awns (if present) barbless; rays pink, purple, or white
1 Rays predominantly yellow, orange, or red (sometimes with some brown, maroon, or purple coloration as well).
                       12 Receptacle naked, epaleate.
                          13 Leaves and phyllaries with large, scattered, embedded oil glands, making the plants strongly aromatic (the glands translucent in living plants, usually golden-brown or blackish in herbarium specimens); plants annual, decumbent and much branched from the base (except Tagetes, annual and generally erect and sparingly branched); [tribe Heliantheae; subtribe Pectidinae].
                            14 Leaves unlobed, entire or serrate.
                              15 Leaves coarsely toothed, mostly opposite (the distal alternate)
                              15 Leaves entire (though with marginal setae), opposite
                            14 Leaves pinnately lobed or pinnatisect (pinnately divided nearly to the midrib but the leaflets not separate), the margins of terminal segments usually serrate (sometimes entire).
                                16 Pinnate leaf segments lanceolate, 3-10 mm wide; calyculus absent
                                16 Pinnate leaf segments linear, 1-3 mm wide; calyculus present, of linear bracts and subtending the involucre.
                                  17 Phyllaries distinct to their bases, or nearly so; ray florets 5-8; heads borne singly or in pairs/triplets; leaf surfaces puberulent
                                  17 Phyllaries connate > ½ their length; ray florets 5-21; heads borne singly; leaf surfaces glabrous to variously pubescent/floccose
                          13 Leaves and phyllaries lacking embedded oil glands, though smaller punctate glands sometimes present; perennial or annual plants, upright and little or moderately branched below the inflorescence.
                                    18 Leaf surfaces densely scabrous and canescent; [s. TX westward]
                                    18 Leaf surfaces glabrous to hirsute but never canescent; [collectively widespread]
                                       19 Leaves mainly basal or basally disposed (a few pairs low on the stem), the blades elliptic to ovate, the surfaces hirsute and stipate-glandular
                                       19 Leaves mainly cauline, the blades primarily linear, lanceolate oblanceolate, or occasionally oblong (if ovate, then leaves mostly cauline), the surfaces glabrous to strigose but not stipitate-glandular
                                         20 Pappus absent or of 1-2 antrorsely-barbed or subplumose bristles; leaf blades usually lobed; [c. and TX westward; formerly Perityle]
                                         20 Pappus absent, of scales, or coroniform (if coroniform then with 6-8 barbellulate bristles as in Jamesianthus); leaf blades unlobed; collectively widespread, including c. TX]
                                           21 Heads smaller and many (10-300+), arranged into dense, flat-topped corymbs; disc florets 1-15; ray florets 0-2, the laminae inconspicuous; phyllaries 6 (-9), in 1 series; [collectively more widespread but absent from n. AL and wc. GA]
                                           21 Heads larger and fewer (< 9 per inflorescence), arranged singly or in loose corymbs; ray florets 6-14, the laminae conspicuous; phyllaries 12-18, broadly ovate, squarrose and in several imbricate series; [on calcareous substrates in n. AL and wc. GA]
                       12 Receptacle paleate, with paleae, bristles, or scales.
                                             22 Disc florets functionally staminate (“sterile”, not producing cypselae), the style undivided, their ovaries much smaller than ovaries of the ray florets (which are functionally pistillate).
                                               23 Inner phyllaries prickly with straight or uncinate prickles, and each enveloping a cypsela and swelling into a bur-like structure
                                               23 Inner phyllaries unarmed, not becoming bur-like (though those of Melampodium do invest the fruit).
                                                 24 Plants with tack-glands or pit-glands on stems, leaves, and/or phyllaries; [waif, e. TX; native further westward]
                                                 24 Plants without tack-glands or pit glands.
                                                    25 Outer phyllaries orbicular, connate 1/4 - 1/3 of their lengths; [exotic, rare]
                                                    25 Outer phyllaries orbicular to lance-linear, not connate; [native, collectively widespread in our area].
                                                      26 Phyllaries in 2 series; pappus coroniform (Chrysogonum) or absent (Lindheimera)
                                                        27 Plants perennial, often trailing; pappus persistent, forming a minute, half-cup-shaped crown; [e. LA eastward, e. of MS river]
                                                        27 Plants annual, erect; pappus absent (cypselae with 2 pappus-like processes originating from cypsela wings); [w. LA westward, w. of MS river]
                                                      26 Taller, robust plants, the stems usually 5-40 cm long at maturity, erect; pappus absent or of 2 awns.
                                                          28 Cypselae strongly flattened, borne in 2-3 series from the 2-3 series of ray florets
                                                          28 Cypselae thick, not flattened, borne in 1 series from the 1 series of ray florets
                                             22 Disc florets functionally bisexual (“fertile”, producing cypselae), the style divided, their ovaries as large as, or larger than, ovaries of the ray florets (which may be either functionally pistillate or completely neuter).
                                                            29 Ray corolla (the lamina) persistent on the achene and becoming papery and bleached.
                                                              30 Outer phyllaries with a stiff, maroon-colored subulate apex
                                                              30 Outer phyllaries lacking a stiff subulate apex.
                                                                 31 Leaf margins serrate or toothed; plant a perennial; cypselae subterete, 4-5 mm long; [natives]
                                                                 31 Leaf margins entire; plant an annual; cypselae 3-angled or flattened, 6-10 mm long; [non-native waifs]
                                                            29 Ray corolla articulate from the achene and falling after flowering.
                                                                   32 Pappus of 20 plumose bristles 4-5 (-7.5) mm long
                                                                   32 Pappus absent or of scales or awns (if with awnlike bristles as in Acmella, then these never plumose).
                                                                     33 Plants with tack-glands or pit-glands on stems, leaves, and/or phyllaries; [waif, VA and NC northward; subtribe Madiinae]
                                                                     33 Plants without tack-glands or pit-glands on stems, leaves, and/or phyllaries; [natives and non-natives, collectively widespread]
                                                                       34 Paleae not notably clasping the cypsela; cypselae notably flattened (or weakly compressed/angled in Guizotia); heads small, the receptacle 3-8 mm in diameter (ca. 10-15 mm in Guizotia).
                                                                         35 Pappus absent; ray laminae longer, 8-14 mm long
                                                                         35 Pappus persistent, of 2-3(-5) scales or awns; ray laminae shorter, 2-5 mm long.
                                                                           36 Cypselae of two types, strongly dimorphic: those of the rays smooth and flattened, the margins conspicuously lacerate and the pappus awns reduced in length; those of the discs ribbed and muricate, with 2 lengthened and divergent awns
                                                                           36 Cypselae of the rays and discs similar, monomorphic and not as above (the achenes weakly dimorphic in C. vialis, but the apical awns of similar length and the margins with only small, inconspicuous winglike projections)
                                                                       34 Paleae either entirely enveloping and falling with each cypsela or conduplicate (V-shaped in cross section), the 2 sides of the V partially clasping the cypsela; cypselae flattened, terete, or angled; heads mostly larger.
                                                                              37 Phyllaries apparently 4, the outer 4 foliaceous and forming a fused quadrangle which conceals the much smaller and narrower inner phyllaries (each inner phyllary subtending a ray floret); cypselae finely 32-40 ribbed
                                                                              37 Phyllaries not as above, instead 5 or more and not forming a conspicuously fused quandrangle; cypselae angled or smooth (sometimes angled, but lacking many fine ribs)
                                                                                38 Paleae completely surrounding the cypselae of the disc florets, forming a perigynium or sac-like appendage
                                                                                38 Paleae conduplicate or partially enclosing the cypselae, but not forming a sac-like appendage.
                                                                                  39 Disc florets with hairy staminal filaments; pappus of 4-6 aristate to lacerate scales; [TX westward]
                                                                                  39 Disc florets without hairy staminal filaments; pappus absent or of 2-3 scales or awns (sometimes accompanied by up to 8-12 additional shorter scales in Helianthus and Simsia, but these readily falling); [collectively widespread].
                                                                                    40 Rays primarily orange; [waif of FL, AL, and LA]
                                                                                    40 Rays primarily yellow; [natives and non-natives, collectively widespread]
                                                                                      41 Cypselae (of at least the disc florets) strongly flattened and generally also winged (thin-margined in Simsia).
                                                                                        42 Cypselae 3-7 mm long; erect herbs; stems often winged
                                                                                        42 Cypselae 1-2.5 mm (Acmella) or 3.5-6 mm long (Simsia); plants erect, decumbent, or prostrate; stems never winged.
                                                                                           43 Phyllaries 8-15; cypselae 1-2.5 mm long; involucre ca. 3-6 mm in diameter
                                                                                           43 Phyllaries 13-43; cypselae 3.5-6 mm long; involucre 5-16+ mm in diameter
                                                                                      41 Cypselae subterete, quadrangular, variously angled, or diamond-shaped in ×-section, not winged.
                                                                                             44 Ray florets pistillate (“fertile”).
                                                                                               45 Erect herb; leaves entire or with a few teeth; peduncles < 3 cm long
                                                                                               45 Creeping herb; leaves serrate and also mostly 3-lobed; peduncles 3-10 cm long
                                                                                             44 Ray florets completely neuter.
                                                                                                 46 Leaves linear; plants perennial with woody caudices; pappus absent; leaf surfaces and ray laminae abaxially gland-dotted; [SC waif; native to sw. US]
                                                                                                 46 Leaf blades usually broader (if linear, leaves either whorled or pappus present); plants annual or perennial, with or without woody caudices; pappus typically of scales or awns; [collectively widespread in our flora area].
                                                                                                   47 Leaves linear, lanceolate, or ovate, almost always some leaves on a plant > 7 mm wide; plants from crowns, some species with thickened vertical storage roots (only H. tuberosus producing horizontal tubers); [collectively widespread in our area]
                                                                                                   47 Leaves linear, 0.5-7 mm wide; plants from slender horizontal tubers; [c. FL north to s. AL]

Key to Asteraceae, Key D: Herbaceous composites with the leaves basal only (stems bearing heads with bracts only)

Copy permalink to share | Check for keys that lead to this key

1 Heads quasi-radiate (corollas of flowers bilaterally symmetrical); [natives, of the Coastal Plain]; [tribe Mutisieae]
  2 Lower leaf surfaces finely hirtellous and stipitate-glandular; [TX]
  2 Lower leaf surfaces densely covered with dense wool; [Coastal Plain, NC to FL, west to TX]
1 Heads either radiate (with ray and disc flowers), disciform (with outer pistillate flowers barely radiate, the subligulate portion ca. 0.5 mm long, and inner disc flowers, or discoid (with disc flowers only)
    3 Heads disciform; involucres 2-4 mm in diameter; [natives, of s. FL]; [tribe Plucheae]
    3 Heads discoid or radiate; involucres 10-20 mm in diameter; [non-natives, more common northwards and inland]; [tribe Senecioneae].
      4 Heads discoid, disc flowers purplish; heads borne in a raceme or thyrse on a leafless stem, before the leaves
      4 Heads radiate, rays yellow, disc flowers yellow; heads borne solitary on scapes, before the leaves

Key to Asteraceae, Key E: herbaceous composites with leaves alternate or basal,
liguliflorous heads (composed of ligulate florets), and sap usually milky

Copy permalink to share | Check for keys that lead to this key

1 Cypselae (at least of the inner florets of the head) beaked, the beak usually > ½ the length of the cypselae body.
  2 Heads solitary and terminal at the end of a stem unbranched to its base.
    3 Flower color pink to purple or white
    3 Flower color yellow to yellowish-white.
      4 Leaves basal and cauline, grasslike (untoothed and unlobed); stem leafy
      4 Leaves basal only, variously toothed to pinnately lobed; stem scapiform (leafless).
        5 Pappus of bristles
        5 Pappus either of outer scales and inner bristles or entirely of aristate scales.
          6 Pappus of outer scales and inner plumose bristles; leaves oblanceolate to oblong; plants annual or perennial
          6 Pappus of aristate scales; leaves linear to narrowly lanceolate; plants annual
  2 Heads several per stem, in various corymbiform, umbelliform, spiciform, or paniculiform arrays (rarely solitary and terminal in the smallest and most depauperate individuals in a population.
             7 Achenes distinctly flattened or compressed.
               8 Florets 5 per head
               8 Florets (6-)10-50+ per head
                 9 Plants taprooted annuals or biennials, without rhizomes
                 9 Plants rhizomatous perennials, the rhizomes deeply rooted in the substrate
             7 Achenes terete or prismatic.
                   10 Pappus of plumose bristles, at least the inner series; plant an annual or biennial.
                     11 Receptacles lacking paleae
                     11 Receptacles with paleae
                   10 Pappus of simple capillary bristles; plant an annual, biennial, or perennial.
                       12 Beak of the cypsela with a ring of soft white reflexed hairs at the summit (just below the pappus); phyllary apices usually with keel-like flap
                       12 Beak of the cypsela lacking a ring of hairs as described; phyllary apices lacking keel (phyllary bases can be keeled in Crepis)
                          13 Pappus of 80-150 barbellulate bristles; plant an annual or biennial
                          13 Pappus either of 40-50 (or more) smooth bristles or of 20-30 barbellulate bristles; plant a perennial.
                            14 Phyllaries tomentose; pappus of 40-50 (or more) smooth bristles
                            14 Phyllaries lacking pubescence; pappus of 20-30 barbellulate bristles
1 Cypselae beakless, the apex typically rounded or truncate (sometimes tapered but lacking a distinct beak and conspicuously < ½ the length of cypselae body).
                              15 Leaves basally disposed (stem leaves few or none, if present generally smaller in size than the basal leaves, which are persistent into flowering and fruiting); corollas yellow, orange, or red.
                                16 Pappus absent or of both scales and barbellulate bristles.
                                  17 Heads solitary or 2-3; pappus absent, the cypselae never bearing scales or bristles; cypselae with 8-10 ribs (3-5 prominent); [waif, ne. US]
                                  17 Heads solitary; pappus present (occasionally absent in Krigia); cypselae with 10-20 ribs; [widespread natives]
                                    18 Pappus of 40-80 smooth to barbellulate bristles and sometimes a few subulate to setiform scales (but pappus never absent); [Great Plains, midwest US]
                                    18 Pappus absent, of scales, or of 5-45 bristles and 5+ scales; [collectively widespread]
                                16 Pappus of bristles only (these barbellulate, plumose, or rarely smooth).
                                       19 Pappus of plumose bristles (these somewhat flattened at their bases)
                                       19 Pappus of barbellulate bristles (sometimes smooth-barbellulate).
                                         20 Leaves with entire margins; plants perennials, from short rhizomes, a short caudex with fibrous roots, or a thick taproot.
                                           21 Leaf margins not undulate, the blades elliptic, oblong, or spatulate
                                           21 Leaf margins white-ciliolate and usually undulate, the blades linear-lanceolate
                                         20 Leaves coarsely toothed or pinnately lobed; plants annuals, from a taproot.
                                             22 Flowers usually pink to lavender; [TX; tribe Mutisieae]
                                             22 Flowers usually yellow to orange; [widespread, including TX; tribe Cichorieae].
                                               23 Involucre 5-12 mm high; achenes usually > 2.5 mm long; pappus bristles distinct, 3-7 mm long
                                               23 Involucre 3-5 mm high; achenes 1.5-2.5 mm long; pappus bristles basally connate (unified basally into a ring-like structure at the junction of achene summit), 2.5-3.5 mm long
                              15 Leaves basal and cauline (plant often beginning with a basal rosette, but by flowering bearing well-developed stem leaves about as large as the basal leaves, the basal rosette often withering prior to flowering and fruiting); corollas yellow, orange, red, blue, pink, white, or lavender.
                                                 24 Pappus absent or of scales.
                                                    25 Corollas pale blue (rarely pink or white)
                                                    25 Corollas yellow (rarely orange).
                                                      26 Phyllaries enfolding the outer cypselae
                                                      26 Phyllaries not enfolding the outer cypselae.
                                                        27 Stem winged and spiny; leaves spiny-margined; receptacle with paleae
                                                        27 Stem not winged or spiny; leaves not spiny-margined; receptacle lacking paleae.
                                                          28 Cypselae 1.2-2.8 mm long; heads borne single at the ends of scapiform stems that are unbranched (rarely few-branched near the base); plants to 7 dm tall
                                                          28 Cypselae 3-5 mm long; heads borne in corymbiform or thyrsiform arrays; plants to 15 dm tall
                                                 24 Pappus of numerous smooth, barbellate, or plumose bristles.
                                                            29 Cypselae more or less strongly flattened.
                                                              30 Pappus of 30-45+ bristles, these at least partly plumose
                                                              30 Pappus of simple capillary bristles only
                                                            29 Cypselae terete or prismatic, slightly or not at all flattened.
                                                                 31 Corollas pink, purple, lavender, white, or creamy-yellow.
                                                                   32 Leaves linear, entire, < 3 mm wide, or reduced to scales; cypselae 11-14 mm long; [of the Coastal Plain of GA and se. AL south to s. FL]
                                                                   32 Leaves broader, of various shapes, usually hastate, irregularly lobed, and/or serrate; cypselae 3.5-10 mm long; [collectively widespread in our area, south to n. FL]
                                                                 31 Corollas bright yellow, orange, or red.
                                                                     33 Leaves finely prickly; pappus of frizzy outer bristles and coarse, straight inner bristles
                                                                     33 Leaves not finely prickly; pappus of barbellulate bristles (sometimes of uneven thickness but not of 2 distinct types)
                                                                       34 Plants taprooted annuals and biennials (rarely perennials); pappus bristles white and soft in texture
                                                                       34 Plants fibrous-rooted perennials; pappus bristles white, light to medium tan, or sordid, stiff.
                                                                         35 Cypselae (2-) 2.5-7 mm long; pappus of (30-) 40-80 white, tan, or sordid bristles, in 1-2+ series; plants cespitose; corollas yellow
                                                                         35 Cypselae 1-2.5 mm long; pappus of 25-40+ white to sordid bristles, in 1 series; plants stoloniferous (cespitose in a few species); corollas yellow or orange

Key to Asteraceae, Key F: herbaceous composites with leaves spiny, leaves alternate or basal, and heads discoid

Copy permalink to share | Check for keys that lead to this key

1 Heads appearing spherical (the ‘visual head’ actually a secondary head aggregated of numerous 1-flowered heads); disk flowers blue (rarely almost white)
1 Heads not spherical; disk flowers pink (rarely yellow).
  2 Stem winged, the wings armed with spines.
    3 Pappus of plumose bristles
    3 Pappus of barbellulate bristles.
      4 Cypselas borne on the surface of the receptacle, intermixed with setiform scales; foliage glabrous or more loosely and sparsely pubescent
      4 Cypselas borne in the pits of the naked honeycombed receptacle; foliage densely white-tomentose
  2 Stem not winged.
        5 Leaves green with white mottles
        5 Leaves green (or relatively uniformly whitened by hairs, especially on the lower surface).
          6 Disc flowers pink (rarely white)
          6 Disc flowers yellow to maroon .
             7 Pappus absent or of scales or barbellulate bristles.
               8 Inner phyllaries tipped with small, simple spines; cypselas 4-angled; receptacles epaleate but bearing subulate scales; pappus absent or of narrow scales 1-10 mm long
               8 Inner phyllaries tipped by pinnately divided spines > 5 mm long; cypselas terete, 20-ribbed; receptacles epaleate but bearing flattened bristles; pappus of 2 series of smooth or roughened awns, the outer series 9-10 mm long, the inner 2-5 mm long
             7 Pappus of numerous, basally connate, plumose bristles
                 9 Heads radiate, the laminae broad and yellow-colored; leaves toothed with small spines present; [tribe Astereae].
                   10 Phyllaries spine-tipped with accompanying white margins (if lacking spine tips as in X. texanum, then inner phyllaries abruptly enlarged into ovate or sometimes elliptic blades 2-5 mm wide)
                   10 Phyllaries not spine-tipped and lacking conspicuous white margins, 0.9-1.7 mm wide.
                 9 Heads discoid (Carlina with inner phyllary tips stramineous, and reminiscent of ray laminae); leaves with prominent, sharp spines; [tribe Cardueae].
                     11 Receptacle epaleate but scaly, each flower surrounded by connatemembranous scales, dissected upwards into linear lobes
                     11 Receptacle epaleate but covered with tawny to white bristles or setiform scales

Key to Asteraceae, Key G: Herbaceous composites with the leaves alternate, the heads lacking rays, and with 0 pappus

Copy permalink to share | Check for keys that lead to this key

1 Receptacle paleate (bearing receptacular bracts which individually subtend at least some of the flowers of the head).
  2 Involucres absent or vestigial, usually with 0 phyllaries; heads very small, < 6 mm high, < 4 mm wide; annuals, < 4 dm tall; disc corollas hidden or dingy in color; [tribe Gnaphalieae]
    3 Bisexual flowers (1-) 2-10 (-11), pappus of (11-) 13-28+ bristles visible in heads; functionally staminate flowers 0.
      4 Receptacles cylindric to clavate (5-15× as tall as the diameter); most pistillate paleae open to ± folded (at most each enfolding, not enclosing a floret; apices acuminate to aristate); innermost paleae erect to ascending in fruit; cypselae mono- morphic (the outer ± equaling the inner)
      4 Receptacles fungiform to obovoid (0.4-1.6× as tall as the diameter); most pistillate paleae ± saccate, each ± enclosing a floret, apices blunt; innermost paleae spreading in fruit; cypselae dimorphic (the outer longer than the inner)
    3 Bisexual flowers 0 or 2-7, pappi 0; functionally staminate flowers 0 or 2-5, pappi 0 or of 1-10 (-13) bristles hidden in heads
        5 Paleae of the bisexual or staminate flowers flat to concave, not enclosing florets, apices entire; cypselae glabrous; [mainly west of the Mississippi River, rarely east (as in MS, AL, GA, and SC]
        5 Paleae of the bisexual flowers saccate, each enclosing a floret, apices 2-fid or 3-fid; cypselae (at least the outer) strigose; [se. Texas]
  2 Involucres present, conspicuous, of many phyllaries; heads larger, 2-20+ mm wide; perennials or coarse annuals, 2-25 dm tall; disc corollas apparent, yellow, white, or pink to purplish.
          6 Disc flowers yellow or greenish yellow; [either native of s. TX southwards, or a waif]
             7 Phyllary margins and tips scarious; fresh foliage aromatic
             7 Phyllary margins and tips herbaceous; fresh foliage not aromatic
          6 Disc flowers white or whitish to pink or purple; [collectively widespread].
               8 Heads in corymbiform arrays; heads bisexual; involucres not burlike or nutlike, lacking tubercles or hooked or straight prickles or spines; leaves lanceolate, unlobed, cuneate at base
                 9 Heads lacking ray flowers; disc flowers pink (rarely nearly white); [Coastal Plain; FL and GA]
                 9 Heads with 5 white (fading to cream), scoop-shaped, inconspicuous ray flowers (keyed here as a 'failsafe'); disc flowers white; [widespread in our region]
               8 Heads in spiciform or racemiform arrays; heads unisexual (each head either with only female or only male flowers); involucres burlike, with tubercles or straight or hooked spines or prickles; leaves ovate or broader (in outline), often pinnatifid or bipinnatifid, truncate, cordate, or cuneate at base; [collectively widespread in our region].
                   10 Involucres of pistillate heads (burs) 1-4 mm long, with (1-) 5-12+ straight spines; larger leaves bipinnatifid
                   10 Involucres of pistillate heads (burs) 10-40 mm long, with 30-75+ hooked prickles; phyllaries larger leaves pinnatifid or unlobed
1 Receptacle epaleate (lacking receptacular bracts individually subtending flowers, but the receptacle sometimes bearing bristles, setae, hairs, or pits with laciniate margins.
                     11 Flowers white, pink, purple, or bluish.
                       12 Annuals, (1-) 2-10 (-30+) cm tall; corollas whitish, inconspicuous, absent on most of the (pistillate) flowers; leaves 2-3× pinnati-palmately lobed into linear segments; cypselae with an apical spine
                       12 Perennials, 10-150 cm tall; corollas at least light pink and often darker pink, purple, or blue, conspicuous, present on all flowers; leaves pinnately lobed or entire, the segments not linear; cypselae not spined.
                          13 Heads ovoid, 10-21 mm long; flowers 15-40 mm long, with long, narrow lobes; phyllaries >30, in 6+ series, and the apices often with a fringe of appendages and/or a spine tip; basal and lower stem leaves usually pinnately lobed (to entire and unlobed); [widespread in our region, especially northwards]
                          13 Heads cylindrical, 2-6 mm long; flowers ca. 3 mm long, with deltate lobes; phyllaries 12-15 in 2-3 series, herbaceous and unadorned; basal and lower stem leaves entire and unlobed; [Coastal Plain of FL and GA]
                     11 Flowers yellow, cream, or rarely red-tinged.
                            14 Leaf blades unlobed.
                              15 Heads 3-4 (-6) mm in diameter, solitary; plants 0.3-3 dm tall; [ballast waif, especially in wet, saline places]
                              15 Heads (3-) 5-20+ mm in diameter, solitary or borne in corymbiform or paniculiform arrays; plants 1.5-15 dm tall.
                                16 Involucres 8-20+ mm in diameter; cypselae 1.5-4.5 mm long
                                16 Involucres (3-) 5-8 (-10) mm in diameter; cypselae 1.5-2 mm long
                            14 Leaf blades 1-3× pinnately or palmately lobed into linear or spatulate segments.
                                  17 Annual 0.1-5 dm tall.
                                    18 Disc corollas cream to pale yellow, 0.5-0.8 mm long; fresh plants malodorous; [waif]
                                    18 Disc corollas greenish yellow, 1.1-1.3 mm long; fresh plants pineapple-scented; [common weed, especially northwards in our region]
                                  17 Perennial, biennial or annual 4-15 dm tall.
                                       19 Heads borne in paniculiform, racemiform, or spiciform arrays, never flat-topped; flowers < 50 per head
                                       19 Heads borne in corymbiform arrays; flowers 60-300+ per head

Key to Asteraceae, Key H: Herbaceous composites with the leaves alternate, the heads lacking rays, and with a pappus of bristles only

Copy permalink to share | Check for keys that lead to this key

1 Receptacle paleate (bearing receptacular bracts which individually subtend at least some of the flowers of the head).
  2 Involucres absent or vestigial, usually with 0 phyllaries; heads very small, < 6 mm high, < 4 mm wide; annuals, < 4 dm tall; disc corollas hidden or dingy in color; [tribe Gnaphalieae].
    3 Receptacles cylindric to clavate (5-15× as tall as the diameter); most paleae of pistillate flowers open to ± folded (at most each enfolding, not enclosing a floret; apices acuminate to aristate); innermost paleae erect to ascending in fruit; cypselae monomorphic (the outer ± equaling the inner)
    3 Receptacles obovoid (0.4-1.6× as tall as the diameter); most paleae of pistillate flowers ± saccate, each ± enclosing a floret, apices blunt; innermost paleae spreading in fruit; cypselae dimorphic (the outer longer than the inner)
  2 Involucres present, conspicuous, of many phyllaries; heads larger, 2-20+ mm wide; perennials or coarse annuals, 2-25 dm tall; disc corollas apparent, white, or pink to purplish or reddish.
      4 Phyllaries tipped by a hooked spine
      4 Phyllaries not armed, herbaceous to the tip.
        5 Cypselae with 5 angles or 5 grooves; involucre 2-3 mm tall and in diameter; pappus of 1 (-5+) usually glandular setae (interpretable as bristles, hence keyed here); [Coastal Plain; FL and GA]
        5 Cypselae with 10 ribs; involucre 3.5-12 (-15) mm tall and in diameter; pappus of 35-40 barbellulate bristles; [mainly Coastal Plain; VA south to s. FL, west to e. LA]
          6 Heads larger, the involucre 6-15 mm high, with 15-40 phyllaries; leaves with conspicuous (at least at 10× magnification) resin dots.
          6 Heads smaller, the involucre 3.5-6 mm high, with 5-12 phyllaries; leaves without shining punctate glands (except punctate-glandular in Litrisa carnosa, of the FL peninsula).
             7 Stem with punctate glands; peduncles punctate-glandular and also hirsutulous; pappus bristles in 2 series; [of FL peninsula]
             7 Stem eglandular, glabrous or spreading-hirsute; peduncles glabrous or stipitate-glandular; pappus bristles in 1 series; [widespread in the Coastal Plain].
1 Receptacle epaleate (lacking receptacular bracts individually subtending flowers, but the receptacle sometimes bearing bristles, setae, hairs, or pits with laciniate margins; some taxa lacking paleae on many or most flowers of the head are keyed here as well, as a failsafe).
               8 Phyllaries in 1-2 series and equal or subequal in length to one another, the phyllaries often coherent/fused (the involucre thus appearing as a cylinder with ribs extending from base to top, except in species in which the phyllaries are radially winged); calyculus (of bracts subtending the phyllaries and differing from them in texture, color, or orientation) present or not; [tribe Senecioneae]
                 9 Annuals, 2-10 dm tall (or perennial, vining or sprawling and to 30 dm long in Gynura); leaves cauline and alternate; disc flowers orangey-brown, brick-red, purplish, yellow.
                   10 Plants perennial, vining or sprawling, velutinous or villous with purple hairs
                   10 Plants annual, upright or ascending, not purple-hairy.
                     11 Involucre subtended by several leafy bracts; cypselae 8- or 10-ribbed
                     11 Involucre not subtended by bracts; cypselae 5-ribbed
                 9 Perennials (robust annuals in Erechtites), 2-30 dm tall; leaves basally disposed and alternate on the stem (except only alternate in Erechtites); disc flowers cream or yellow (rarely pale lavender in Arnoglossum).
                       12 Plants annual, with leaves primarily on the stem; leaves sharply and raggedly toothed or even pinnately lobed; heads disciform (the outer flowers female, the middle bisexual, and the inner functionally male)
                       12 Plants perennial, with leaves basally disposed (larger basal leaves, decreasing in size upwards, the basal leaves sometimes withering late in the year); leaves with generally regular toothing, stem leaves sometimes pinnately lobed; heads discoid (all flowers bisexual)
                          13 Disc flowers yellow.
                            14 Perennials; leaves basally disposed, the basal or lower stem leaves unlobed (though toothed)
                            14 Annuals; leaves not basally disposed, evenly distributed on the stem, the lower stem leaves more-or-less pinnatifid
                          13 Disc flowers cream, white, or greenish white (to pale lavender in some Arnoglossum).
                              15 Phyllaries 5, radially wing-keeled or not; disc flowers 5; calyculus absent
                              15 Phyllaries 7-21, not radially winged; disc flowers 10-55 (-80+); calyculus absent or of 1-9+ bracts.
                                16 Plants (5-) 6-24 dm tall; calyculus of 4-9+ bracts; [collectively relatively widespread in our region]
                                16 Plants 1-7 dm tall; calyculus absent or of 1-5+ bracts; [Mountains of w. NC and e. TN]
               8 Phyllaries in 3-17 series and unequal in length (imbricated); calyculus absent.
                                  17 Stem prominently winged, the wings > 1 mm wide
                                  17 Stem not winged (sometimes the leaves decurrent < 2 cm down the stem from the node).
                                    18 Phyllaries usually 12-30+ in 3-10+ series, scarious (at least the margins).
                                       19 Plants perennial, mat-forming by stolons, with basal rosettes of spatulate leaves, these usually with obvious appressed hairiness on (at least) the lower and (sometimes also) the upper blade surfaces; erect stems (with scattered alternate leaves) 4-45 cm tall; plants usually dioecious (pistillate and staminate heads on separate plants)
                                       19 Plants annual, biennial, or perennial, not mat-forming or stoloniferous, erect, the leaves usually lanceolate or linear (spatulate in some species), leaf surface hairiness variable; erect stems (with alternate leaves) 5-80 cm tall; plants hermaphroditic (with heads containing both pistillate and functionally staminate flowers) or dioecious or subdioecious (in Anaphalis).
                                         20 Plants perennials; heads discoid (unisexual or nearly so); plants usually unisexual and dioecious
                                         20 Plants annuals (perennials in Omalotheca sylvatica); heads disciform (each bisexual); plants hermaphroditic.
                                           21 Leaves oblanceolate, 1.5-4 mm wide, notched at the tip or with a minute apiculus in the notch; pappus bristles persistent, plumose
                                           21 Leaves various in shape, the lager > 4 mm wide, acuminate, acute, obtuse or rounded at the tip; pappus readily falling, the bristles barbellate to barbellulate.
                                             22 Heads in capitate clusters arrayed in corymbiform or paniculiform arrays; phyllaries white or silvery; pappus bristles distinct, falling individually
                                               23 Pappus bristles connate basally into a ring, falling together
                                               23 Pappus bristles distinct, falling individually
                                             22 Heads in spicate arrays, or in glomerules axillary to leaves or well-developed bracts; phyllaries white to red-tinged or brown; pappus connate at the base into a ring, falling as a unit.
                                                 24 Perennial; phyllaries in 2-3 series; cypselas strigillose; involucre light to dark brown; [waif, in PA]
                                                 24 Annual; phyllaries in 3-7 series; cypselas papillate or glabrous; involucre white to red-tinged or brown; [widespread in our region]
                                                    25 Pappus of 12-28 bristles; cypsela faces papillate
                                                    25 pappus of 8-12 bristles; cypsela faces glabrous
                                    18 Phyllaries in 3-5 series, herbaceous to thickened, generally green in color.
                                                      26 Disc corollas yellow, pale yellow, orange, reddish, or brown.
                                                        27 Phyllaries 5, in one series; phyllaries streaked with elongate glands filled with strong-scented oil
                                                        27 Phyllaries 10-60+ in 2-9 series; phyllaries eglandular or with small punctate glands.
                                                          28 Phyllaries strongly armed with long spines
                                                          28 Phyllaries unarmed
                                                            29 Decumbent annual, to 2 dm tall; heads solitary and axillary
                                                            29 Erect or ascending annual or perennial, 1-15 dm tall; heads in corymbiform, paniculiform, or racemiform arrays.
                                                              30 Plants dioecious, either male or female; heads either staminate, with 10-50 flowers, or pistillate, with 20-150 flowers; heads pale yellow to white
                                                              30 Plants hermaphroditic; heads bisexual, with 2-35 flowers, either all perfect (bisexual), or a mixture of perfect (bisexual) and functionally staminate flowers.; heads yellow to pale yellow.
                                                                 31 Leaves entire; flowers 2-6 per head; heads arrayed in a neatly flat-topped corymb
                                                                 31 Leaves (at least the larger) toothed or lobed; flowers 5-35 per head; heads in racemiform, paniculiform, or corymbiform arrays, but if corymbiform, irregular.
                                                                   32 Larger leaves 2.5-10 cm long, 1-4 cm wide; involucres 6-12 mm in diameter; [widespread in our region]
                                                                   32 Larger leaves 1-5 cm long, < 1 cm wide; involucres 2-7 mm in diameter; [s. and se. TX].
                                                                     33 Inflorescences in rounded, corymbiform arrays; leaves glabrous
                                                                     33 Inflorescences in elongate, racemiform to paniculiform arrays; leaves moderately to densely hairy with both glandular and non-glandular hairs
                                                      26 Disc corollas pink, purple, lavender, blue, or white.
                                                                       34 Phyllaries either spine-tipped, or appendaged with a marginal zone that is strikingly different in color and texture than the phyllary body (except Leuzea); receptacle epaleate, but densely beset with bristles or subulate scales (flattened bristles).
                                                                         35 Phyllaries scarious-margined, neither spine-tipped nor appendaged with dentate or fringed appendages
                                                                         35 Plyllaries either spine-tipped or appendaged.
                                                                           36 Phyllaries tipped by a hooked spine
                                                                           36 Phyllaries appendaged, the appendage black, brown, tan, or scarious, and toothed or fimbriate.
                                                                              37 Phyllary bodies linear, their margins entire (phyllary appendages fimbriate; however); leaf margins entire, denticulate or serrulate
                                                                              37 Phyllary bodies oblong, ovate, or obovate, both their margins and appendages fimbriate; leaf margins entire, toothed, or pinnately lobed.
                                                                                38 Flower heads pink, lavender, or purplish; phyllary appendages scarious and spineless, decurrent along the phyllary margin nearly to the phyllary base; perennial; flowers pink to purple, flowering Jun-Oct
                                                                                38 Flowers heads blue; phyllary appendages not or only slightly decurrent along the phyllary margins; annual; flowers pale to medium blue, flowering Apr-Jun
                                                                       34 Phyllaries not armed, also not appendaged with a marginal zone that is strikingly different in color and texture than the phyllary body.
                                                                                  39 Plants dioecious, either male or female; heads either staminate, with 10-50 flowers, or pistillate, with 20-150 flowers; heads pale yellow to white
                                                                                  39 Plants hermaphroditic; heads bisexual, with 3-100+ flowers, either all perfect (bisexual), or a mixture of perfect (bisexual) and functionally staminate flowers.; heads purple, pink, lavender, or whitish.
                                                                                    40 Leaf blades orbicular; leaf margin scalloped or crenate; [rare waif in our region]
                                                                                    40 Leaf blades linear, lanceolate, or ovate; leaf margin entire, serrulate, or serrate; [collectively common and widespread].
                                                                                      41 Style branch appendages absent, the style branch stigmatic along the upper surface nearly to the tip; [tribe Vernonieae].
                                                                                        42 Pappus bristles in 1 series; heads disciform (the outer flowers of the head pistillate, the inner flowers of the head functionally staminate); leaf margins toothed; corollas white to cream or pink (rarely purple)
                                                                                        42 Pappus bristles in 2 series, the outer much shorter; heads discoid (all flowers bisexual and fertile); leaf margins toothed or entire; corollas purple or deep pink (rarely pale pink or white)
                                                                                      41 Style branch appendages deltate, lanceolate, or terete to clavate, the style branch various.
                                                                                           43 Style branch appendages deltate or lanceolate; style branches stigmatic along 2 lines from near the base to the attachment of the appendage; [tribe Astereae].
                                                                                             44 Disc flowers 8-20; pappus bristles in 2 series; [GA and Panhandle FL west to AL, GA, and LA]
                                                                                             44 Disc flowers 75-95+; pappus bristles in 1 series; [waif]
 Symphyotrichum ciliatum
                                                                                           43 Style branch appendages terete to clavate; style branches "eupatorious", linear to slightly clavate-thickened at the tip, the stigmatic surface usually papillate; [tribe Eupatorieae]
                                                                                               45 Lower stem leaves with at least a few divergent teeth; phyllaries elongate-striate nerved
                                                                                               45 All leaves entire (or at most with a few obscure crenations); phyllaries lacking elongate striations, the nerving not apparent.
                                                                                                 46 Heads borne in spiciform or racemiform arrays (all heads directly attached to a central access of the capitulescence)
                                                                                                 46 Heads borne in corymbiform, paniculiform, or thyrsiform arrays (at least some heads borne on secondary branches of the capitulescence.
                                                                                                   47 Involucres 2-3 mm tall; cypselas 5-ribbed
                                                                                                   47 Heads 3-20 mm tall; cypselas 10-ribbed.
                                                                                                     48 Heads 3.5-6 mm tall; phyllaries 5-12 in 1-2 (-3) series; [Coastal Plain, NC to FL to e. LA].
                                                                                                        49 Stem with punctate glands; peduncles punctate-glandular and also hirsutulous; pappus bristles in 2 series; [of FL peninsula]
                                                                                                        49 Stem eglandular, glabrous or spreading-hirsute; peduncles glabrous or stipitate-glandular; pappus bristles in 1 series; [widespread in the Coastal Plain].
                                                                                                     48 Heads 3-20 mm tall; phyllaries 15-50+ in 3-7 series; [collectively widespread].
                                                                                                          50 Heads borne in corymbiform arrays; involucres 6-12 (15) mm tall; receptacles paleate, at least towards the periphery of the receptacle; pappus of 35-40 barbellulate to barbellate bristles; [nearly restricted to the Coastal Plain]
                                                                                                          50 Heads borne in paniculiform or thyrsiform arrays; involucres 3-20 mm tall; receptacles epaleate; pappus of 12-40 coarsely barbellate, subplumose, or plumose bristles; [widespread in our region]

Key to Asteraceae, Key I: Herbaceous composites with the leaves alternate, the heads lacking rays, and with a pappus wholly or partly of scales or awns

Copy permalink to share | Check for keys that lead to this key

1 Disc flowers yellow, orange, red, or brownish.
  2 Leaves 1-2× pinnately lobed into linear or filiform segments.
    3 Pappus of 16-20 scales
    3 Pappus of 6-10 scales; [historical waif]
  2 Leaves unlobed, or if pinnately lobed, the segments broad (> 3 mm wide).
      4 Receptacle paleate
      4 Receptacle epaleate.
        5 Pappus scales rudimentary, crown-like
        5 Pappus scales conspicuous.
          6 Phyllaries 25-125 in 3-9 series; receptacles flat or slightly convex
          6 Phyllaries 6-25 (-60) in 2-3 series; receptacles globose.
             7 Stems not winged
             7 Stems winged from node to node
1 Disc flowers pink, purple, or white
               8 Receptacle paleate
               8 Receptacle epaleate.
                 9 Disc flowers 1-5 (rarely more) per head.
                   10 Disc flowers white (rarely cream), 3; heads not aggregated into a secondary cluster subtended by bracts; [se. TX]
                   10 Disc flowers lavender to purple, sometimes so lightly so as to be white, 1-4 (-5+); heads aggregated into clusters of 1-40 heads, subtended by 1-3 bracts; [collectively widespread in our region]
                     11 Heads (1-) 10-40 per secondary cluster, borne in corymbiform arrays; bracts subtending clusters (2-) 3, deltate; pappus of 5 (-6) scales, each aristate
                     11 Heads 1-5+ per secondary cluster, borne in spiciform arrays; bracts subtending clusters 1-2, linear, lanceolate, or spatulate; pappus of 6-10 scales, laciniate or aristate
                 9 Disc flowers (2-) 10-100+ per head (at least most heads with >10 flowers).
                       12 Phyllaries 5-15 (-21) in 1-2 (-3) series.
                          13 Involucres 2-3 mm in diameter; flowers 7-10 per head; [Coastal Plain, FL and GA]
                          13 Involucres 4-15+ mm in diameter; flowers 20-80 per head; [SC, GA, and FL westwards]
                       12 Phyllaries 18-many in 5+ series.
                            14 Phyllary tips modified into a spine or into an enlarged, lacerate or pectinate network or fringe; receptacle epaleate, but densely bristly; [tribe Cynareae].
                              15 Heads discoid, all flowers similar
                                16 Pappus normally of plumose bristles (and keyed elsewhere), but sometimes reduced to scales or awns
                                16 Pappus of an outer setiform scales (appearing as "flattened bristles"
                              15 Heads disciform or radiant, the outer flowers of the head sterile, with long lobes, appearing like false rays
                                  17 Plant a perennial; flowers pink to purple, flowering Jun-Oct
                                  17 Plant an annual; flowers pale to medium blue, flowering Apr-Jun
                            14 Phyllary tips not so modified, unarmed and unelaborate; receptacle epaleate and naked; [tribe Vernonieae].
                                    18 Peripheral flowers of the heads enlarged and bilaterally symmetrical, appearing like "false rays"
                                    18 Peripheral flowers of the heads similar to the interior flowers, all disc flowers.
                                       19 Annuals; cypselas not ribbed
                                       19 Perennials; cypselas 8-10-ribbed.
                                         20 Heads each subtended by 3-8 leafy bracts; pappus of narrow scales only, quickly falling
                                         20 Heads not subtended by leafy bracts; pappus of an inner series of scales or bristles and an outer series of bristles, persistent

Key to Asteraceae, Key J: Herbaceous composites with the leaves alternate or basal and the heads radiate, the rays yellow, orange, red

Copy permalink to share | Check for keys that lead to this key

1 Disc florets functionally staminate (sterile), with style undivided.
  2 Phyllaries 10 or less, in 2 series; plants annual; [primarily OK and TX]
  2 Phyllaries 12 or more, in (2-) 3-4 series; plants perennial; [collectively widespread, including OK and TX in our area]
    3 Cypselae with broad wings with 2 confluent awns
    3 Cypselae not winged, pappus absent or of 2-4 ciliate scales.
      4 Disc florets green, red, or maroon; pappus absent; leaves lanceolate, lyrate, or lyrate-pinnatifid (but not deeply pinnatifid); [nc. TX westward]
      4 Disc florets primarily yellow; pappus of 2-4 ciliate scales; leaves deeply pinnatifid [sw. AR and e. TX westward, waif in e. LA]
1 Disc florets bisexual (fertile), with style bifurcate.
        5 Leaves decurrent down the stem (the stem appearing winged); cypselae strongly flattened and often winged
        5 Leaves not decurrent; cypselae moderately compressed to 4-angled, not winged (or if winged, the cypselae obcompressed or obconic).
          6 Receptacular bracts connate to form a honeycomb-like structure in which the flowers are set; pappus of 7-12 scales
          6 Receptacular bracts not fused and not forming a honeycomb-like structure; pappus absent, of awns/scales, or a fused crown (Phoebanthus sometimes with up to 4 additional scales shorter than 2 primary lacerate scales).
             7 Heads subtended by a calyculus (bracts distinct from phyllaries)
             7 Heads without a calyculus.
               8 Leaves conspicuously clasping the stem
               8 Leaves not noticeably clasping the stem.
                 9 Leaves pinnately lobed, the primary lobes usually pectinately divided; [non-native]
                 9 Leaves unlobed or if pinnatifid to pinnately lobed, the primary lobes not pectinate; [natives].
                   10 Pappus of 2 caducous, or readily-falling scales; receptacles flat
                   10 Pappus absent, or of 1-2 lacerate scales (on shoulders of cypselae) plus additional lacerate (bristlelike) scales; receptacles subspheric, conical, columnar, or otherwise convex.
                     11 Receptacle low conic to convex; phyllaries (14-) 20-40+, in 2-3 series.
                       12 Leaf blades linear; plants from slender, horizontal tubers; pappus absent or of 1-2 lacerate scales (on shoulders of cypselae) plus additional lacerate (bristlelike) scales; [natives of AL and FL; subtribe Ecliptinae]
                       12 Leaf blades variously shaped, not all linear (often with some ovate or rhombic leaves); plants from woody caudices; pappus absent; [waif in SC, native in arid southwest; subtribe Helianthinae]
                     11 Receptacle strongly conic or columnar; phyllaries 3-21, in 1-2 series; [subtribe Rudbeckiinae].
                          13 Phyllary series unequal (outer longer than inner); cypselae strongly compressed; ray florets subtended by receptacular bracts
                          13 Phyllary series subequal; cypselae not strongly compressed; ray florets not subtended by receptacular bracts (only the disc flowers with bracts)

Key to Asteraceae, Key K: Herbaceous composites with the leaves alternate or basal and the heads radiate, the rays yellow, orange, red

Copy permalink to share | Check for keys that lead to this key

1 Pappus absent, or of crowns (coroniform) or scales (the scales entire, erose, aristate, or occasionally lacerate, but ultimately with broadened bases compared to bristles).
  2 Stems usually winged from decurrent leaf bases (except H. amarum, which bares stems with copious, mostly entire, linear leaves)
  2 Stems not winged.
    3 Pappus absent.
      4 Phyllaries in (3-) 4-9+ series, usually with apices hooked or looped; pappus of setiform awns or scales but falling (thus sometimes appearing absent post maturation)
      4 Phyllaries in 2-4 series, usually with apices not notably hooked or looped; pappus absent entirely (not just falling).
        5 Receptacles pitted and hairy (the hairs swollen and apically hooked); [primarily native in western portion of our area]
        5 Receptacles not pitted, nor hairy; [waifs].
          6 Leaves usually 1-3 pinnately lobed; receptacles epaleate
          6 Leaves entire or sometimes toothed, but not pinnately lobed; receptacle epaleate or sometimes with short bristles or setae
    3 Pappus coroniform or of scales.
             7 Heads smaller, the involucres (excluding phyllary apices) 0.8-7.5 mm in diameter.
               8 Ray floret laminae fan-shaped and apically 3 or 5 lobed (the laminae sometimes very short and not surpassing phyllary length in Hymenoxys).
                 9 Cauline leaves pronounced, simple or with 3-19+ lobes (the lobes linear-filiform); cypselae villous or pubescent (rarely glabrous); heads radiant, sometimes appearing nearly discoid due to diminutive or inconspicuous ray florets
                 9 Leaves strongly basally-disposed (except T. linearifolia, with basal and cauline leaves, the cauline leaves not with linear lobes), entire or with 2-6 teeth (or sometimes lobes); cypselae moderately to densely pubescent; heads clearly radiate
               8 Ray floret laminae not apically 3-5 lobed.
                   10 Leaves pinnately compound with linear leaf segments 1-3 mm wide (if not pinnate as in T. tephroleuca, then plants ashy white subshrubs); calyculus present (occasionally absent), of linear bracts subtending the involucre; plants with translucent oil glands evident throughout most parts; [subtribe Pectidinae]
                   10 Leaves simple (if also sometimes subshrubs, these not ashy white as in Gutierrezia); calyculus absent; plants sometimes gland-dotted on leaves (or sometimes also with resinous phyllaries, but not with oil glands throughout).
                     11 Plants annual, with a strong terpenoid smell when crushed, the leaves and phyllaries glutinous; phyllaries 12-15 in 1-2(-3) series
                     11 Plants annuals, biennials, or perennials, if annuals then not bearing a strong terpenoid smell when crushed, the leaves sometimes glutinous but not also the phyllaries; phyllaries 4-40 in 2-4 series (Gutierrezia) or 5-12 in 1-2 series (Psilostrophe).
                       12 Stems glabrous, or papillate-scabrous (but not clearly gray-green in appearance); involucre 1.5-5.5 mm in diameter; plants either annuals or subshrubs; phyllaries 4-40 in 2-4 series
                       12 Stems arachno-villous, appearing gray-green; involucre 4-7 mm in diameter; plants biennials or perennials; phyllaries 5-12 in 1-2 series
             7 Heads larger, the involucres (8-) 10-20+ mm in diameter.
                          13 Phyllaries in (3-) 4-9+ series; cypselae glabrous; leaves often toothed (and also often ovate to oblanceolate).
                            14 Pappus in 2 series, the outer of 25-40 barbellate bristles, the inner of 8-15+ setiform awns or subulate scales 7-10 mm long
                            14 Pappus in 1 series, of mostly 2-3 smooth to setiform awns OR subulate scales 2.5-8 mm long (sometimes with up to 6-8 total barbellulate/setiform awns in G. squarrosa, G. lanceolata, and G. texana)
                          13 Phyllaries in 2-3 series (2-3+ in Gaillardia); cypselae villous or otherwise sparsely to densely pubescent; leaves entire, toothed, or sometimes pinnately lobed (the lobes linear, as in Hymenoxys).
                              15 Receptacles entirely epaleate, not bearing any minute setae; ray florets (apical portion of the laminae) 3-5 lobed.
                                16 Cauline leaves pronounced, simple or with 3-19+ lobes (the lobes linear-filiform); cypselae villous or pubescent (rarely glabrous); heads radiant, sometimes appearing nearly discoid due to diminutive or inconspicuous ray florets
                                16 Leaves strongly basally-disposed (except T. linearifolia, with basal and cauline leaves, the cauline leaves not with linear lobes), entire or with 2-6 teeth (or sometimes lobes); cypselae moderately to densely pubescent; heads clearly radiate
                              15 Receptacles typically with spine-like setae or enations (except Gaillardia aestivalis) or very short bristles (i.e. not appearing "chaffy" overall, but these setae apparent upon dissection of the heads); ray florets (apical portion of the laminae) either unlobed (Amblyolepis) or 3-5 lobed (Gaillardia).
                                  17 Plants sweet-scented; cypselae prominently 10-ribbed; pappus scale apices rounded to acuminate but never aristate; heads borne singly or in paniculiform arrays; ray florets (the laminae) yellow; phyllaries usually not strongly reflexed in fruit
                                  17 Plants not sweet-scented; cypselae ca. 4-angled but lacking numerous prominent ribs; pappus scale apices often aristate; heads borne singly; ray florets (the laminae) often bicolored; phyllaries usually strongly reflexed in fruit
1 Pappus of barbellate to barbellulate bristles (sometimes also with shorter outer scales).
                                    18 Phyllaries in one series.
                                       19 Plants herbaceous vines; flowers conspicuously orange; calyculus of lanceolate-filiform bractlets; [non-native, FL and TX]
                                       19 Plants not herbaceous vines; flowers primarily yellow (sometimes yellow-orange); calyculus absent (or 1-2 indistinct bractlets); [natives or non-natives, collectively widespread]
                                         20 Pappus bristles red, brown, or purple colored; flowers yellow to yellow-orange; [waif, MD]
                                         20 Pappus bristles white; flowers primarily yellow; [native and non-native, widespread]
                                           21 Disc florets sterile; stems bracteate but lacking leaves, the leaves simple; [non-native of disturbed habitats, ne US]
                                           21 Disc florets fertile; stems with leaves, the leaves simple to pinnately lobed (or otherwise compound); [natives and non-natives, widespread and of both intact and disturbed habitats].
                                             22 Stem leaves deeply toothed, pinnately lobed, or otherwise compound, not clasping or weakly so; plants perennial or rarely annuals (P. glabella)
                                             22 Stem leaves shallowly toothed to subentire, conspicuously clasping the stem; plants annuals
                                    18 Phyllaries in 2-8+ series.
                                               23 Plants herbaceous vines; flowers conspicuously orange
                                               23 Plants not herbaceous vines; flowers primarily yellow (sometimes yellow-orange)
                                                 24 Pappus double, of both an inner (more apparent) series of longer bristles and an outer series consisting of short triangular scales or significantly shorter (and often coarser) bristles, these sitting more or less at the base of the inner pappus bristles (in Prionopsisthe outer bristles subtend the inner awns or scales).
                                                    25 Plants usually gland-dotted; phyllaries often looped/hooked; outer bristles subtending inner scales; involucres globose/hemispheric
                                                    25 Plants glabrous, sericeous, or glandular; phyllaries often ascending, sometimes squarrose; pappus scales subtending inner bristles; involucres variously shaped.
                                                      26 Outer pappus of scales connate, usually forming cups; ray laminae often reduced, 1.5-2+ mm long; [waifs; tribe Inuleae]
                                                      26 Outer pappus not connate; ray laminae prominent, usually > 3 mm longer; [native, sometimes weedy; tribe Astereae].
                                                        27 Ray florets without pappus
                                                        27 Ray florets with pappus (similar to that of disc florets).
                                                          28 Leaves parallel-veined, linear and grasslike
                                                          28 Leaves pinnately veined, usually broader and not grass-like.
                                                            29 Plants biennials or perennials, with taproots or fibrous-rooted; phyllaries linear-lanceolate to oblanceolate; inner pappus whitish to stramineous, outer pappus linear to narrowly-triangular
                                                            29 Plants annuals with taproots; phyllaries linear to linear-lanceolate; inner pappus yellow to rust-colored proximally, outer pappus scale-like; disc florets bisexual and fertile (B. pilosa) or functionally staminate (B. hirtella)
                                                 24 Pappus simple, of only bristles and not with an additional series of reduced bristles or scales (sometimes pappus in 2-4 unequal series, or outer bristles progressively shortened in Xanthisma).
                                                              30 Leaves toothed and tipped with spines (spinulose).
                                                                 31 Phyllaries spine-tipped with accompanying white margins (if lacking spine tips as in X. texanum, then inner phyllaries abruptly enlarged into ovate or sometimes elliptic blades 2-5 mm wide)
                                                                 31 Phyllaries not spine-tipped and lacking conspicuous white margins, 0.9-1.7 mm wide.
                                                              30 Leaves entire or if toothed, not tipped with conspicuous spines (as in Grindelia).
                                                                   32 Ray florets longer, 1.5-3+ cm long; anthers tailed; leaf bases conspicuously cordate-clasping
                                                                   32 Ray florets shorter, < 1.5 cm long (to 1.6 cm in Grindelia lanceolata); anthers not tailed (instead obtuse to sagittate), or if sagittate-tailed, then plants foul-smelling (as in Dittrichia, a waif in ne. US).
                                                                     33 Plants viscid, foul-smelling; anthers sagittate-tailed at the base [waif, ne. US]
                                                                     33 Plants scentless, or if bearing an odor, not foul-smelling; anthers cuneate to sagittate at the base, but not tailed; [widespread natives, rarely e. US waifs otherwise native to w. US].
                                                                       34 Pappus bristles > 60 (30-80+ in Senecio), white-colored.
                                                                         35 Stem leaves deeply toothed, pinnately lobed, or otherwise compound, not clasping or weakly so; plants perennial or rarely annuals (P. glabella)
                                                                         35 Stem leaves shallowly toothed to subentire, conspicuously clasping the stem; plants annuals
                                                                       34 Pappus bristles 10-45 (30-80+ in Senecio), white, tawny, or reddish-brown colored.
                                                                           36 Phyllaries in 1-2 equal/subequal series (outer minute bracts sometimes present); stem leaves shallowly toothed to subentire, conspicuously clasping the stem; plants annuals; [tribe Senecioneae)
                                                                           36 Phyllaries in 3+ series, unequal (or rarely subequal); [tribe Astereae].
                                                                              37 Heads small and cylindrical, taller than wide; rays inconspicuous or the laminae often shorter.
                                                                                38 Inflorescences flat-topped corymbs; leaves resinous-punctate, narrow, entire, sessile or subsessile
                                                                                38 Inflorescences rarely flat-topped; leaves not resinous-punctate, usually not linear, often toothed and petiolate
                                                                              37 Heads larger, globose, hemispheric, or broadly conic (2 cm+ broad in Grindelia), usually wider than long; rays conspicuous.
                                                                                  39 Leaves entire, sessile throughout; faces of cypselae strigose to sericeous; pappus persistent, of 30-35 tawny to reddish-brown barbellate bristles
                                                                                  39 Leaves toothed, proximal leaves petiolate; faces of cypselae variously textured (smooth, striate, furrowed, or rugose); pappus readily falling, of 2-3 smooth to setiform awns OR subulate scales 2.5-8 mm long (sometimes with up to 6-8 total barbellulate/setiform awns in G. squarrosa, G. lanceolata, and G. texana)

Key to Asteraceae, Key L: Herbaceous composites with the leaves alternate or basal and the heads radiate, the rays white, pink, purple, and the 0 pappus

Copy permalink to share | Check for keys that lead to this key

1 Rays primarily pink or purple, not white-tinged (flowers viewed from above, adaxial surface of the rays).
  2 Receptacles epaleate; leaves pinnatisect; [waif, ne. US]
  2 Receptacles paleate; leaves simple or bladeless; [widespread natives]
    3 Leaves without blades (the petioles terete, appearing Juncus-like); ray florets pink or purplish; disc florets fewer (60-120+), the corollas usually yellowish; [wet pinelands and seepages]
    3 Leaves broad and conspicuous; ray florets purple; disc florets many (200+), the corollas pink, green, red, purple, or yellow colored; [plants of prairies and similar habitats]
1 Rays primarily white to white-tinged or yellow throughout (lamina of the ray with at least some white basally if not uniformly yellow).
      4 Disc florets functionally staminate (thus cypselae only forming from ray florets)
      4 Disc florets bisexual, fertile.
        5 Leaves entire or toothed, but not deeply pinnatisect; sometimes with smaller rounded lobes (Leucanthemum basal leaves can have deeper, round lobes); plants usually not aromatic.
          6 Stem internodes usually winged (at least proximally)
          6 Stems not winged.
             7 Leaves primarily basal, the cauline leaves absent or fewer and much reduced in size from basal leaves; abaxial surface of the ray laminae white or sometimes pink-tinged (on live plants) but lacking a prominent colored midstripe.
               8 Leaves strigose, the margins crenate-serrate; roots not red-tipped; abaxial surface of ray laminae often pink or purple tinged, flowers closing at night
               8 Leaves glabrous (or faces sparsely hairy), the margins entire or toothed; roots usually red-tipped; rays drying pinkish but abaxial surface not conspicuously pink tinged on live plants
             7 Plants with well-developed cauline leaves, the margins entire or sometimes toothed (usually entire in Aphanostephus, occasionally deeply lobed); abaxial surface of the ray lamina sometimes white, but often with a prominent purple or blue midstripe (except Leucanthemum, which merely dries pink)
                 9 Flowering heads smaller, arranged in corymbiform or paniculiform arrays; ray florets with diminuitive laminae (0.3-5.0 mm long), heads thus sometimes superficially appearing disciform (rays sometimes double in horticultural forms as in Achillea ptarmica)
                   10 Plants aromatic; leaves linear to narrowly lanceolate, the surfaces glabrous or sparsely hairy; pales more or less folded but lacking distal papillae; cypselae not shed along with subtending phyllary and disc florets; [tribe Anthemideae]
                   10 Plants not aromatic; leaves variously shaped, at least some usually lyrate or oblanceolate (not all linear), the surfaces hairy and gland-dotted (at least abaxially); pales distally pappilate or fimbriate; cypselae shed along with subtending phyllary and 2 disc florets each invested in a pale; [tribe Heliantheae]
                 9 Flowering heads larger, singular on scapes (if in 2's or 3's, these large and not in broad corymbiform arrays), rays prominent, the laminae usually > 10 mm long
                     11 Rays white abaxially (drying pinkish); receptacles broadly convex, not pitted; plants rhizomatous perennials; pappus absent; [tribe Anthemideae]
                     11 Rays white or with prominent blue or purple midstripe (sometimes present after drying); receptacles conic, pitted; plants tap-rooted annuals (except Astranthium riddellii); pappus absent, coroniform, or of scales; [tribe Astereae]
                       12 Phyllaries scarious margined; cypselae 4-angled and with 4-12 thick ribs, pappus absent, of scales or coroniform; ray florets closing distally at night
                       12 Phyllaries sometimes hyaline but not scarious margined; cypselae compressed, lacking prominent ribs (though sometimes glochidiate), pappus absent or short coroniform; ray florets not closing at night
        5 Leaves deeply lobed or pinnatisect, the ultimate segments linear or if rounded, the sinuses of the lobes nearly reaching the midrib (pinnatifid); plants typically aromatic (except Tripleurospermum and Parthenium); [tribe Anthemideae].
                          13 Receptacles paleate (chaffy or with apparent bracts), the pales sometimes distally fimbriate or papillate (Parthenium).
                            14 Stem internodes usually winged
                            14 Stems not winged.
                              15 Rays (the laminae) white with conspicuous yellow coloration at the bases
                              15 Rays primarily white to white-tinged (without strong yellow color at the base of the laminae) or yellow throughout (lamina of the ray with at least some white basally; if rays are white).
                                16 Flowering heads smaller, arranged in corymbiform or paniculiform arrays; ray florets with diminuitive laminae (0.3-5.0 mm long); ray and disc florets both white or white-ish.
                                  17 Plants usually aromatic; pales more or less folded but not distally fimbriate or papillate; cypselae not shed along with phyllary and disc florets; [tribe Anthemideae]
                                  17 Plants not aromatic (at least not conspicuously so); pales distally fimbriate or papillate; cypselae shed along with subtending phyllary and 2 disc florets each invested in a pale; [tribe Heliantheae]
                                16 Flowering heads larger, arranged mostly singular at the ends of scapes, not in broad corymbiform arrays (except Tripleurospermum, which has larger heads and prominent rays), ray laminae prominent; ray and disc florets differing in color (rays typically white; discs typically yellow).
                                    18 Plants annual, erect or decumbent, ill-scented or not; ultimate leaf margins dentate to lobed; cypsela ribs usually 9-10 (sometimes lacking); [widespread non-natives]
                                    18 Plants perennial and prostrate (mat-forming), aromatic; ultimate leaf margins entire; cypsela ribs weak; [waif, ne. US]
                          13 Receptacles epaleate (lacking chaff).
                                       19 Ultimate leaf lobes broad, ovate-shaped (not linear)
                                       19 Ultimate leaf lobes linear.
                                         20 Plants usually aromatic; maturing flowering heads with deflexed ray laminae; cypsela 5-ribbed, lacking apical resin sacs
                                         20 Plants not aromatic; ray laminae not deflexed; cypsela with 3 ribs and 1-5 abaxial-apical resin sacs

Key to Asteraceae, Key M: Herbaceous composites with the leaves alternate or basal and the heads radiate, the rays white, pink, purple, and with a pappus of bristles only

Copy permalink to share | Check for keys that lead to this key

1 Head 1 per plant (individual rosette); stems 0.1-0.3 dm tall; [TX and OK westwards]
1 Heads 5-300+ per (well-developed) plant; stems 0.3-35 dm tall; [collectively widespread].
  2 Stems with numerous thorns (1-2 cm long, modified axillary branches), and few or no leaves
  2 Stems leafy, not armed with thorns.
    3 Leaves pinnatifid to bipinnatifid; plant annual
    3 Leaves unlobed; plant perennial from caudices, crowns, or rhizomes.
      4 Annuals from taproots; ray florets 0.3-7 mm long.
        5 Ray laminae 0.3-1.0 mm long; cypselas 1.0-1.5 mm long, compressed, 1-nerved on each face
        5 Ray laminae 1.3-7 mm long; cypselas (1.2-) 1.5-2.7 (-3) mm long, compressed or not, 5-18-nerved.
          6 Disc flowers 25-35 per head; ray flowers in 1 series
          6 Disc flowers 4-10 (-13) per head; ray flowers in 1-3 series
      4 Perennials from caudices, crowns, or rhizomes; ray florets > 3 mm long.
             7 Basal and lower stem leaves petiolate, with blades cordate to rounded at the base
               8 Heads borne in corymbiform arrays, the branches often subtended by large and leaf-like bracts; phyllaries < 2.5 (-3)× as long as wide, often > 1 mm wide; plants (in most species) colonial by rhizomes; rays (in most species) white; phyllaries (in many species) glandular
               8 Heads borne in paniculiform arrays, the branches bearing small and narrow bracts; phyllaries > 3× as long as wide, often < 1 mm wide; plants (in most species) solitary; rays (in most species) blue, violet, or pink; phyllaries eglandular
             7 Basal and lower stem leaves sessile (sometimes cordate-clasping), or if petiolate then with cuneate to attenuate bases.
                 9 Rays pink, purple, lavender, blue, or pale pink [plants with pale pink rays may need to be keyed both ways].
                   10 Heads borne in paniculiform or in diffuse and irregular arrays
                   10 Heads borne either in flat-topped or rounded, corymbose arrays, or in narrow racemiform or spiciform panicles, or solitary.
                     11 Leaves 1.2-4.0 cm long, 1-4 mm wide, stiff, scabrous margined and weakly spine-tipped; plants 1-7 dm tall; pappus in 2 series, the inner much longer than the outer (ca. 1 mm long) bristles
                     11 Leaves either longer or broader or both, herbaceous; Leaves (5-) 13-45 mm wide, herbaceous, neither scabrous-margined nor spine-tipped; plants 4-30 dm tall; pappus not divided into distinct inner and outer series.
                       12 Plants 10-35 dm tall; larger leaves (basal or low on the stem) with cuneate or attenuate bases, the blades 30-50 cm long; [non-native, rarely persistent or spreading from horticultural use]
                       12 Plants 1-12 dm tall; larger leaves with blades 2-20 cm long; [natives, collectively widespread and common in our region]
                 9 Rays white (or very slightly pink-flushed).
                          13 Heads borne in paniculiform or in diffuse and irregular arrays
                          13 Heads borne either in flat-topped or rounded, corymbose arrays, or in narrow racemiform/spiciform panicles, or solitary.
                            14 Heads arrayed in 1 (or more) cylindrical thyrses
                            14 Heads borne in either narrow racemiform/spiciform panicles, or in corymbose arrays.
                              15 Heads borne in narrow racemiform/spiciform panicles
                              15 Heads borne in corymbose arrays.
                                16 Disc flowers white
                                16 Disc flowers yellow (and often fading to pink or purplish).
                                  17 Rays 2-7, the ray ligules 2-11 mm long, often twisted or contorted; phyllaries whitish with a distinct green tip, the green area about as wide as long
                                  17 Rays (8-) 9-40, the ray ligules 4-20 mm long, generally planar (or neatly coiling at post-maturity); phyllaries various.
                                    18 Pappus double, with an outer set of bristles < 1 mm long; leaves entire
                                    18 Pappus not divided into distinctively different outer and inner series; leaves toothed or entire.
                                       19 Achenes not glandular, either glabrous or strigillose
                                       19 Achenes densely gland-dotted

Key to Asteraceae, Key N: Herbaceous composites with the leaves alternate or basal and the heads radiate, the rays white, pink, purple, and the pappus absent or essentially so

Copy permalink to share | Check for keys that lead to this key

1 Receptacles paleate (with chaff); [tribe Heliantheae].
  2 Stems not winged; leaves simple; disc florets and ray florets differing in color
  2 Stems usually winged; leaves simple or pinnately to palmately lobed; disc florets concolorous (of similar color) with ray florets
1 Receptacles epaleate (lacking chaff).
    3 Leaves 1-3 pinnately-lobed or pinnatifid.
      4 Plants arachnose or wooly pubescent throughout; [waif, SC; tribe Arctotideae]
      4 Plants glabrous or densely viscid but not wooly tomentose throughout (if hirsuto-pilose, this usually concentrated distally).
        5 Receptacles flat or broadly conic, not conspicuously pitted; heads in corymbiform arrays; pappus a short crown; rays white, pink, or purple adaxially; [widespread non-natives, tribe Anthemideae]
        5 Receptacles conic, pitted; heads borne singly (Aphanostephus) or in loose corymbiform arrays (Egletes); rays white adaxially; [natives primarily w. of MS river; tribe Astereae].
          6 Plants aromatic, but not densely viscid, instead sometimes hairy distally; ray laminae not shorter than involucre; phyllary margins broadly scarious; pappus coroniform or a mix of variously shaped scales; [widespread w. of MS river]
          6 Plants densely viscid with stipitate-glandular trichomes; ray laminae sometimes shorter than involucre; phyllary margins scarious or not; pappus of uneven rings; [TX]
    3 Leaves entire or toothed, but neither deeply lobed nor pinnatifid.
             7 Plants 1-3 cm tall, densely pin-cushioned (pulvinate); flowering heads sessile, borne singly, conspicuously nestled within the rosette leaves and proportionately of similar size; pappus persistent, of 12-35+ setiform scales in 1 series; phyllaries in 4-6+ series
             7 Plants of various size, but not densely pin-cushioned; flowering heads borne on stems, if peduncles short the heads not conspicuously nestled within similar-sized leaf rosettes; pappus various, phyllaries in 2-6 series.
               8 Pappus of 5-40+ longer inner bristles plus shorter outer scales or setae (these sometimes falling); stems and leaves often (but not always) stipitate-glandular.
                 9 Inner barbellate bristles numerous (5-40+); stems and leaves usually glandular; plants tap-rooted, fibrous-rooted, or stoloniferous; annuals, biennials, or perennials
                 9 Inner bristles barbellate or thicker, fewer (ca. 5); stems and leaves glandular (C. asteroides) or eglandular (C. asteroides; C. belliloides); plants tap-rooted annuals
               8 Pappus primarily of awns, scales, crowns, or thickened rings (these minute, 0.1-0.8 mm long), sometimes also accompanied by shorter inner scales or bristles; stems and leaves eglandular (sometimes minutely glandular in Chaetopappa bellidifolia).
                   10 Cypsela margins narrowly winged or winglike.
                     11 Receptacles obscurely pitted; pappus of 2-3 sclerified awns plus often with 7-12 shorter bristles or scales; [collectively widespread]
                     11 Receptacles smooth; pappus of 2 thin awns plus an elliptic ring of shorter awns; [s. TX]
                   10 Cypsela margins not narrowly winged (4-angled in Aphanostephus but lacking marginal wings).
                       12 Receptacles conic, pitted; cypselae 4-angled; pappus of setiform to awn-tipped scales or short ciliate crowns (0.1-0.2 mm long, minute, thus requiring magnification)
                       12 Receptacles flat or slightly convex, smooth (not pitted); cypselae variously shaped but not conspicuously 4-angled; pappus of minute, thickened rings (C. bellidifolia) or erose cuplike crowns (C. imberbis)

Key to Asteraceae, old Key J: Herbaceous composites with the leaves alternate or basal and the heads radiate, the rays white, pink, purple

Copy permalink to share | Check for keys that lead to this key

1 Receptacles chaffy (paleate).
  2 Phyllaries dry, scarious-margined.
    3 Ray florets 1-5 mm long; heads small in corymbiform arrays
    3 Ray florets > 5 mm long; heads large, terminating the branches
  2 Phyllaries herbaceous, not scarious on margins.
      4 Ray florets < 2.5 mm long; disk florets sterile, with an undivided style
      4 Ray florets > 5 mm long; disk florets fertile, with a divided style.
        5 Ray florets pink or light purple, > 15 mm long; heads single
        5 Ray florets white, 5-10 mm long; heads 20-100 in a compound corymb
1 Receptacles naked (epaleate).
          6 Pappus absent or coroniform.
             7 Achene faces minutely glandular-glochidiate (bearing small gland-like bristles)
             7 Achene faces glabrous or short-strigose but not glandular-glochidiate.
               8 Leaves basal; cypselae marginally 2-ribbed; mature plants < 2 dm tall
               8 Leaves cauline (and sometimes also basal); cypselae with ± 10 ribs; mature plants > 2 dm tall
                 9 Heads solitary or 2-3 in corymbiform arrays
                 9 Heads 5-60+ in corymbiform arrays
          6 Pappus present, of bristles or scales.
                   10 Heads primarily discoid with reduced ray florets
                   10 Heads radiate with conspicuous ray florets.
                     11 Taprooted annuals; ray florets 1-7 mm long.
                       12 Leaves and stems not fleshy, rarely glabrous; cypselas < 1.4 mm long
                       12 Leaves and stems fleshy, mostly glabrous; cypselas > 1.4 mm long
                     11 Not taprooted and mostly perennials; ray florets > 3 mm long.
                          13 Ray florets usually > 60; blooming Apr-Oct
                          13 Ray florets usually < 60; blooming late May-Nov.
                            14 Receptacles hemispheric to conic; pappus often with 2-4 awns (Boltonia) or lacking awns and coroniform (Astranthium).
                              15 Achenes not narrowly winged, the surfaces minutely glandular-glochidiate (bearing small gland-like bristles)
                              15 Achenes narrowly winged, the surfaces sometimes puberulent but not glandular-glochidiate
                            14 Receptacles flat to slightly convex; pappus lacking awns.
                                16 At least the basal and lower leaves both petiolate and cordate/subcordate at base.
                                  17 Often colonial; inflorescence corymbiform, flat-topped or rounded; outer phyllaries > 1 mm broad
                                  17 Not colonial; inflorescence paniculiform, often elongate; outer phyllaries < 1 mm broad
                                16 Basal and lower leaves not both petiolate and cordate/subcordate at base.
                                    18 Leaves sessile and auriculate or cordate-clasping
                                    18 Leaves petiolate or epetiolate but not auriculate or cordate-clasping.
                                       19 Cypselas glandular; pappus double
                                       19 Cypselas not glandular; pappus single or double or in four series.
                                         20 Leaves silvery-silky on both sides (at least when young), entire
                                         20 Leaves not silvery-silky, entire or toothed.
                                           21 Pappus double, with inner bristles distinctly longer than outer bristles.
                                             22 Leaves not rigid, veiny, lanceolate to elliptic or ovate, > 6 mm wide
                                             22 Leaves rigid, 1-nerved, linear to linear-spatulate, < 5 mm wide
                                           21 Pappus simple with all bristles often about the same length.
                                               23 Ray florets white, few (usually 3-8); cypselas densely silky
                                               23 Ray florets white to pink or blue or purple, more numerous (usually 8-30); cypselas glabrous to pubescent but not densely silky.
                                                 24 Ray florets white; involucres < 6 mm long; phyllaries < 1 mm wide
                                                 24 Ray florets white or pink to blue or purple; involucres 7-12 mm long; phyllaries usually > 1 mm wide.
                                                    25 Phyllaries glandular
                                                    25 Phyllaries not glandular.
                                                      26 Phyllaries long-attenuate or loose and spreading
                                                      26 Phyllaries appressed, not long-attenuate.
                                                        27 Plants coarse-hairy; lowest leaves > 5 cm wide
                                                        27 Plants glabrous to hairy; lowest leaves < 3 cm wide