Sceptridium lunarioides (Michaux) Holub. Section: Hiemobotrychium. Common name: Winter Grapefern. Phenology: Jan-Apr. Habitat: Old fields, pastures, young forests, granitic flatrocks, juniper-oak-blue ash woodlands over limestone, cemeteries. Distribution: Wc. NC, c. TN, and s. SC south to n. FL, and west to e. TX and se. OK.
ID notes: The species is hard to spot, and all the more difficult to find because of its phenology; the leaves appear in late fall and die by early spring.
Origin/Endemic status: Endemic
Taxonomy Comments: Wagner (1992) proposed that Botrychium lunarioides be treated in a new monotypic section, Hiemobotrychium, of Botrychium, subgenus Sceptridium. Zhang et al. (2020) advocated its placement in a monotypic genus, Holubiella. Its apparent involvement in the generation of the allopolyploid species Sceptridium jenmanii seems a compelling reason to retain the species in Sceptridium.
Synonymy ⓘ: = K4; = Botrychium lunarioides (Michx.) Sw. — Ar, ETx1, Fl1, FNA2, NS, POWO, RAB, TxFerns, WH3; = Botrypus lunarioides Michx. = Holubiella lunarioides (Michx.) Škoda — Cao & Hauk (2022), Zhang & Zhang (2022), Zhang et al (2020). Basionym: Botrypus lunarioides Michx. 1903
Links to other floras: = Botrychium lunarioides - FNA2
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Heliophily ⓘ: 7
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© Alan M. Cressler | Original Image ⭷
© Alan Cressler: Sceptridium lunarioides, Broxton Rocks Preserve, The Nature Conservancy, Coffee County, Georgia 3 by Alan Cressler source | Original Image ⭷
© Alan Cressler: Sceptridium lunarioides, Stone Mountain Park, DeKalb County, Georgia 3 by Alan Cressler source | Original Image ⭷
© Alan Cressler: Sceptridium lunarioides, Broxton Rocks Preserve, The Nature Conservancy, Coffee County, Georgia 1 by Alan Cressler source | Original Image ⭷
© Alan Cressler: Sceptridium lunarioides, Broxton Rocks Preserve, The Nature Conservancy, Coffee County, Georgia 2 by Alan Cressler source | Original Image ⭷
© Alan Cressler: Sceptridium lunarioides, Stone Mountain Park, DeKalb County, Georgia 3 by Alan Cressler source | Original Image ⭷
© Alan Cressler: Sceptridium lunarioides, Woodruff-Seminole Site, U.S. Army Corps of Engineers, Decatur County, Georgia 3 by Alan Cressler source | Original Image ⭷Feedback
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