Dryopteris goldieana (Hooker ex Goldie) A. Gray. Phen: Jun-Sep. Hab: Boulderfield forests, rich cove forests, seepage swamps, especially over calcareous sedimentary or mafic metamorphic or igneous rocks. Dist: NB west to s. ON and MN, south to nw. SC, n. GA, TN, KY, AR (Peck 2011), IL, and IA.
Origin/Endemic status: Native
Taxonomy Comments: This species is one of the diploid ‘parent species’ of the e. North American reticulately-evolved Dryopteris complex. Its genome (symbolized GG) forms half of the genome of the tetraploid D. celsa, and one third of the hexaploid D. clintoniana. The epithet is correctly spelled "goldieana", following the ICNafp.
Synonymy: = K4, S13; = Dryopteris goldiana (Hook. ex Goldie) A.Gray – Ar, C, F, FNA2, GrPl, Il, K1, K3, Mo1, NE, NY, Pa, RAB, Sf, Tat, Tn, Va, W, WV, Montgomery & Paulton (1981); = Dryopteris goldiana (Hook. ex Goldie) A.Gray ssp. goldiana – G