Humulus americanus Nuttall. Northeastern Hops. Phen: Jul-Aug; Sep-Oct. Hab: Disturbed areas, particularly in rich, alluvial soils. Dist: NS and NL (Newfoundland) south to VA and NC, west to NE, MT, and AB. While native in North America, the native distribution is obscure.

Origin/Endemic status: Native

Taxonomy Comments: The three taxa (two native and one introduced) of the H. lupulus complex in our area are subtly different, the differences apparently sometimes further obscured by introgressive hybridization.

Synonymy: = Humulus lupuloides (E.Small) Tembrock – K4, Tembrock, McAleer, & Gilligan (2016), superfluous name; = Humulus lupulus L. ssp. americanus (Nutt.) Á.Löve & D.Löve – NE; = Humulus lupulus L. var. lupuloides E.Small – C, FNA3, GrPl, Il, K1, NY, Pa, Small (1978); < Humulus lupulus L. – F, G, K3, RAB, S, S13, Tat, Va, W, WV

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Wetland Indicator Status:

• Atlantic and Gulf Coastal Plain: FACU (taxonomic split from wetland indicator species)
• Eastern Mountains and Piedmont: FACU (taxonomic split from wetland indicator species)
• Great Plains: FACU (taxonomic split from wetland indicator species)
• Midwest: FACU (taxonomic split from wetland indicator species)
• Northcentral & Northeast: FACU (taxonomic split from wetland indicator species)

Heliophily ?: 6