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Harperellanodosa Rose. Harperella. Phen: Jun-Aug. Hab: Rocky or gravelly riverbeds, upland depression ponds, seepage on granite flatrocks. Dist: Disjunct and fragmented in distribution: w. MD, e. WV, VA, and c. NC; SC and c. GA; N. AL; AR and e. OK.
Origin/Endemic status: Endemic
Taxonomy Comments: Three taxa are sometimes recognized, but recent studies (molecular and morphological) show insufficient reliable bases for separating them (Feist et al. 2012; M.A. Feist, 2012, pers. comm.). Earlier work is suggestive, and additional study is needed. The three taxa formerly recognized are H. vivipara (leaves 4-12 (-15) cm long; plants 1-5 (-8) dm tall, adventitiously perennial by proliferating from the nodes; rays 1-9 mm long, 2-5 (-9) per inflorescence; pedicels 0.5-2.0 (-2.5) mm long; umbellets with 2-8 flowers) of rocky or gravelly riverbeds in w. MD, e. WV, VA, and c. NC; H. fluviatilis (leaves 4-12 (-15) cm long; plants 1-5 (-8) dm tall, adventitiously perennial by proliferating from the nodes; rays 1-9 mm long, 2-5 (-9) per inflorescence; pedicels 0.5-2.0 (-2.5) mm long; umbellets with 6-15 flowers) of rocky or gravelly riverbeds in n. AL, AR, and e. OK; and H. nodosa (leaves 8-30 cm long; plants 4-10.5 dm tall, strictly annual, not proliferating from the nodes; rays 10-25 mm long, 6-15 per inflorescence; pedicels 3.0-6.0 mm long) of upland depression ponds in SC and seepage on granite outcrops in GA. See references for additional details and discussion.
Other Comments: Belden et al. (2004) provided details on the Virginia occurrence in Aquia Creek, Stafford County.
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Horticultural Information
Intro:Erect, semi-aquatic annual with quill-like leaves found in rocky river beds, upland depression ponds and seepages on granite flatrocks.
Leaves:Leaves alternate, on petioles with a sheathing base, round in cross-section and tapering to a point, to 12 in. long but reduced upward.
Inforescence:
Flowers:Flowers in flat-topped, compound umbel at top of stem and from leaf axils; white; tiny; consisting of 5 petals that curve up and strongly inward, 5 stamens with dark pink anthers, and a 2-part ovary.
Fruits:Fruit oval, consisting of a joined pair of elliptic, ridged seeds.
Comments:
Height:6-36 in.
plant sale text:
bloom table text:
description:Erect, semi-aquatic annual with quill-like leaves found in rocky river beds, upland depression ponds and seepages on granite flatrocks.
leaves:Leaves alternate, on petioles with a sheathing base, round in cross-section and tapering to a point, to 12 in. long but reduced upward.
inflorescence:
flowers:Flowers in flat-topped, compound umbel at top of stem and from leaf axils; white; tiny; consisting of 5 petals that curve up and strongly inward, 5 stamens with dark pink anthers, and a 2-part ovary.
fruits:Fruit oval, consisting of a joined pair of elliptic, ridged seeds.
