Dryopteris carthusiana (Villars) H.P. Fuchs. Common name: Spinulose Woodfern, Toothed Woodfern. Phenology: Jun-Oct. Habitat: Acidic, organic-rich bogs, swamps, less frequently in moist rocky ravines, rich forests, and sloping rock outcrops. Distribution: Circumboreal, in North America ranging from n. QC west to YT, south to NC, SC, ne. GA, TN, AR, NE, w. MT, and WA.
Origin/Endemic status: Native
Taxonomy Comments: This species is a fertile allotetraploid derived from hybridization of Dryopteris intermedia and "Dryopteris semicristata", a hypothetical species which may now be extinct. Its chromosome complement is symbolized IISS. The triploid backcross Dryopteris ×triploidea (Dryopteris carthusiana × intermedia, with chromosome symbolized IIS) is frequent in areas with both parents present.
Synonymy ⓘ: = Ar, C, Can, FNA2, Il, K4, Mo1, NE, NS, NY, Pa, Tn, Va, W, Montgomery & Paulton (1981); Polypodium carthusianam Vill.; = Dryopteris austriaca (Jacq.) Woyn. ex Schinz & Thell. var. spinulosa (O.F.Müll.) Fiori — G; = Dryopteris spinulosa (O.F.Müll.) Watt — GrPl, RAB, S13, Sf, Tat, WV; = Dryopteris spinulosa (O.F.Müll.) Watt var. spinulosa — F; > Dryopteris carthusiana var. carthusiana — POWO. Basionym: Polypodium carthusianam Vill. 1786
Links to other floras: = Dryopteris carthusiana - FNA2
Show parent genus | Show parent in key(s)
Wetland Indicator Status:
- Atlantic and Gulf Coastal Plain: FACW
- Eastern Mountains and Piedmont: FAC
- Great Plains: FACW
- Midwest: FACW
- Northcentral & Northeast: FACW
Heliophily ⓘ: 3
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© Erik Danielson source | Original Image ⭷
© Gary P. Fleming | Original Image ⭷
© Erik Danielson source | Original Image ⭷
© Bruce A. Sorrie | Original Image ⭷
© Bruce A. Sorrie | Original Image ⭷
© Erik Danielson source | Original Image ⭷
© Erik Danielson source | Original Image ⭷
© Erik Danielson source | Original Image ⭷
© Alan Cressler: Dryopteris carthusiana, Bird Hills Nature Area, Huron River Valley, Ann Arbor, Washtenaw County, Michigan 1 by Alan Cressler source | Original Image ⭷
© Erik Danielson source | Original Image ⭷
© Erik Danielson source | Original Image ⭷
© Erik Danielson source | Original Image ⭷
© Erik Danielson source | Original Image ⭷
© Alan Cressler: Dryopteris carthusiana (bottom), and Dryopteris intermedia (top), Southern Appalachian Mountain Bog, North Carolina 1 by Alan Cressler source
© Alan Cressler: Dryopteris carthusiana (right), and Dryopteris intermedia (left), Southern Appalachian Mountain Bog, North Carolina 1 by Alan Cressler source | Original Image ⭷Feedback
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