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1 Leaves strongly bicolored (pale green toward the base of each of the 4 leaflets, darker green toward the tip); aquatic forms with a swollen air bladder just below the leaf
2 Roots present (1-3) between the nodes, as well as at the nodes.
3Distaltooth 0.3-0.8 mm long; sporocarps 3.5-5.0 mm long; sporocarps 1 or 2(-4) per fertile frond, attached to stipe up to 5 mm above stipebase; common peduncle joining 2 peduncles to stipe, if present then only 1 mm
3Distaltooth absent or < 0.2 mm long; sporocarps 4.5-6.0 mm long; sporocarps (1-) 2-5 (-7) per fertile frond, attached to stipe up to 25 mm above stipebase; common peduncle joining 2 or more peduncles to stipe 2-8 mm