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Key to Dichanthelium, Key E: Spikelets 2.1-3.2 mm long, larger culm blades < 13 mm wide, culm nodes not bearded,
the lowermost sometimes puberulent or sparsely pilose

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1 Ligule 1.6-4 mm long  Dichanthelium oligosanthes

1 Ligule < 1.5 mm long.

..2 Leaf blades, at least the lower, cordate or subcordate at the base, 4-25 mm wide.

....3 Culm blades 4-12 mm wide; spikelets obpyriform when viewed dorsally, strongly plano-convex when viewed laterally, usually markedly reddish-purple basally; fertile lemma papillose  Dichanthelium webberianum

....3 Culm blades 5-25 mm wide; spikelets elliptic to elliptic-obovoid when viewed dorsally or laterally, greenish to faintly purple-tinged basally; fertile lemma not papillose.

......4 Ligule ciliate, 0.2-0.8 mm long; culm blades 5-14 mm wide; spikelets 1.8-2.6 mm long; first glumes 0.5-1 mm long  Dichanthelium boreale

......4 Ligule an eciliate membrane 0-0.3 mm long; culm blades 5-25 cm wide; spikelets 2.0-3.7 mm long; first glumes 0.5-2.6 mm long.

........5 Internodes crisp-puberulent; larger culm blades 4-8 (-11) cm long, 5-10 (-12) mm wide, broadest near the base; spikelets 2.1-2.7 mm long; first glumes 0.7-0.9 mm long  Dichanthelium commutatum ssp. ashei

........5 Internodes glabrous to sparsely pubescent, larger culm blades 5-18 cm long, 5-25 mm wide, broadest near the middle or broadly linear; spikelets 2.0-3.7 mm long; first glumes 0.5-2.6 mm long.

..........6 Cauline blades 5-18 cm long, 6-14 mm wide, averaging about 10 × as long as wide, the margins nearly parallel; spikelets 2.9-3.7 mm long; first glumes 11-26 mm long, 2/5 or more as long as the spikelets  Dichanthelium commutatum ssp. equilaterale

..........6 Cauline blades 7-12 cm long, (6-) 12-25 mm wide, ovate-lanceolate, averaging about 6 × as long as wide, the margins curved most of their length; spikelets 2.0-3.2 mm long; first glumes 5-10 mm long, about 1/4 as long as spikelets.

............ 7 Culms erect; blades more nearly symmetrical; spikelets 2.0-3.2 mm long; sterile lemma apex usually blunt to rounded  Dichanthelium commutatum ssp. commutatum

............ 7 Culms basally decumbent; blades strongly asymmetrical-falcate; spikelets 3.0-3.2 mm long; sterile lemma apex usually pointed  Dichanthelium commutatum ssp. joorii

..2 Leaf blades tapering to the base, 2-12 mm wide.

............ ..8 Ligule a stramineous to light brown membrane, with or without terminal cilia.

............ ....9 Ligule a ciliate membrane; spikelets (2.6) 2.8-3.4 (3.6) mm long; first glume (0.9) 1.1-1.6 mm long  Dichanthelium aculeatum

............ ....9 Ligule an eciliate membrane; spikelets 1.6-2.8 (3.0) mm long; first glume 0.3-1.1 mm long.

............ ......10 Panicle rachis smooth, pellucid-punctate; first glume 0.3-0.6 (-0.8 ) mm long, as broad as or broader than long, truncate to obtuse; larger leaves 10-25 cm long, 8-15 mm wide; ligule 0.5-1.3 mm long; lowest elongate culm internode > 2 mm in diameter; lowest nodes usually glabrous or pubescent  Dichanthelium scabriusculum

............ ......10 Panicle rachis scabrous or smooth, not pellucid-punctate; first glume 0.5-1.1 mm long, longer than wide, rounded to acute; larger leaves 3.5-12 cm long, 3-9 mm wide; ligule 0.1-0.6 mm long; lowest elongate culm internode < 2 mm in diameter; lowest nodes retrorsely bearded or glabrous.

............ ........11 Lowest nodes usually retrorsely bearded; ligules (0.1-) 0.3-0.6 mm long; largest vernal blades 7-12 cm long, (4.5-) 6-9 mm wide; panicle peduncle scabrous; spikelets ovate-lanceolate, acute, 2.0-2.4 mm long; first glume lanceolate, blunt to acute; fertile lemma smooth  Dichanthelium cryptanthum

............ ........11 Lowest nodes usually glabrous; ligules 0.1-0.2 (-0.3) mm long; largest vernal blades 3.5-7 cm long, 3-6 mm wide; panicle peduncle smooth; spikelets elliptic, blunt to acute, 1.6-2.2 mm long; first glume ovate to rotund, rounded to acute; fertile lemma papillose  Dichanthelium lucidum

............ ..8 Ligule of short white hairs or absent.

............ ..........12 Leaves basally disposed, usually matted or cushion-forming, larger than the mid and upper culm leaves; blade margins uniformly papillose-ciliate; culms branching only at the base, 0.5-3.5 dm tall; internodes glabrous or sparsely pubescent  Dichanthelium strigosum var. leucoblepharis

............ ..........12 Basal leaves usually rosette-forming, not matted or cushion-forming, usually much smaller than culm leaves; blade margins glabrous, or ciliate only below the middle (or sometimes papillose-ciliate throughout in D. portoricense ssp. patulum, which has densely puberulent internodes); culms branching at the nodes in age, 1.5-7.5 dm tall.

............ ............ 13 Blades progressively longer upwards on culm, the larger 9-20 cm long and 3-8 mm wide, (15-) 20 × or more as long as wide, firm but not stiff, not longitudinally grooved or wrinkled; blade adaxial surface (weakly-) strongly scabrous; sheaths glabrous; second glume and sterile lemma exceeding fertile lemma  Dichanthelium bicknellii

............ ............ 13 Blades similar or more often decreasing in size upwards on culm, the larger 3-16 cm long and 2-25 mm wide; if 15 × or more as long as wide, then blades stiff and longitudinally grooved or wrinkled (sect. Angustifolia); blade adaxial surface smooth to moderately scabrous; sheaths glabrous or pilose; second glume and sterile lemma shorter than to exceeding fertile lemma.

............ ............ ..14 Blades of mid-culm leaves typically long and stiff, acuminate, linear or narrowly lanceolate, usually > 10× as long as wide, only 2-6 mm wide when < 8 cm long.

............ ............ ....15 Vegetative parts glabrous (spikelets pubescent, lowest internodes and sheaths sometimes sparsely pubescent, blades and sheaths sometimes ciliate); mature panicles less than ¼ as wide as long, the branches erect-ascending, the spikelets often subsecund; autumnal blades 4-10 cm long, involute, < 2 mm wide; spikelets 1.8-2.2 mm long.

............ ............ ......16 Larger vernal blades 5-8 cm long and 3-5 mm wide; vernal panicles usually less than ¼ as wide as long, spikelets often subsecund along the strongly ascending to erect branches; spikelets 1.8-2.2 mm long, second glume and sterile lemma pubescent, summits blunt, about equaling fertile lemma; [of coastal wetlands and Piedmont prairies from NC and AR south to the Caribbean and Central America]  Dichanthelium neuranthum

............ ............ ......16 Larger vernal blades 5-9 cm long and 2-3 mm wide; vernal panicles usually ¼ to ½ as wide as long, the branches ascending; spikelets 2.3-3.0 mm long, second glume and sterile lemma sparsely pubescent to glabrous, summits pointed to blunt, exceeding fertile lemma; [of c. and s. FL]  Dichanthelium pinetorum

............ ............ ....15 Vegetative parts pubescent, at least in the lower portion of plant; mature panicles usually more than half as wide as long, the branches spreading-ascending, the spikelets not noticeably subsecund; autumnal blades 1-6 cm long, involute and < 2 mm wide in D. aciculare and D. arenicoloides, or to 9 cm long, flat and 2-4 mm wide in D. angustifolium (autumnal blade dimensions not yet known for D. wilcoxianum); spikelets 1.5-3.2 mm long.

............ ............ ........17 Culms 15-35 cm long; all culm leaf blades long-hirsute on both surfaces, their sheaths densely papillose-hirsute; leaf blades of the primary branches often exceeding the immature vernal panicles  Dichanthelium wilcoxianum

............ ............ ........17 Culms 15-75 cm long; culm leaf blades glabrous to sparsely pubescent, or the lower softly villous abaxially in D. arenicoloides; lower sheaths usually villous, mid-culm and upper sheaths glabrous (though often ciliate basally); leaf blades of primary branches not exceeding immature vernal panicles.

............ ............ ..........18 Spikelets (3.0-) 3.2-3.8 mm long, fusiform, pointed at summit, attenuate at base, with both glumes attached 0.3-0.5 mm below sterile lemma, the autumnal spikelets 3.5-3.8 mm long; larger vernal leaf blades 3-6 mm wide, the lower and mid-culm leaf blades of similar width; autumnal leaf blades involute  Dichanthelium fusiforme

............ ............ ..........18 Spikelets 1.8-3.1 mm long, obovate, blunt, and the base not attenuate (except in D. arenicoloides with autumnal spikelets 2.3-3.1 mm long); larger vernal leaf blades 2-8 mm wide, the lower usually wider and often shorter than mid-culm leaf blades; autumnal leaf blades involute or flat.

............ ............ ............ 19 Longest vernal blades to 16 cm, widest vernal leaf blades 4-8 mm, usually longitudinally wrinkled; vernal and autumnal spikelets 2.3-3.1 mm long; autumnal leaf blades flat, the larger to 9 cm by 2-4 mm  Dichanthelium angustifolium

............ ............ ............ 19 Longest vernal leaf blades to 6 cm (D. aciculare ) or 12 cm (D. arenicoloides) long, 2-5 mm wide, not noticeably wrinkled; vernal spikelets 1.5-2.8 mm long, autumnal spikelets either 1.5-2.3 (D. aciculare) or 2.3-3.1 (D. arenicoloides); autumnal leaf blades involute, the larger to 6 cm by 1.5 mm.

Show caption*© Scott Ward ............ ............ ............ ..20 Longest vernal leaf blades to 12 cm, widest vernal leaf blades 3-4 (-5) mm; vernal spikelets 2.1-2.8 mm long; autumnal spikelets 2.3-3.1 mm long, pointed, attenuate, the glumes attached 0.3-0.5 mm below sterile lemma; first glumes 0.7-1.5 mm long; larger autumnal leaf blades to 6 cm by 1.5 mm  Dichanthelium arenicoloides

Show caption*© Janet Wright, some rights reserved (CC BY-NC), uploaded by Janet Wright ............ ............ ............ ..20 Longest vernal leaf blades to 6 (-8) cm, widest vernal leaf blades 2-5 mm; vernal and autumnal spikelets 1.7-2.3 mm long, blunt, not attenuate, the glumes attached < 0.2 mm below sterile lemma; first glumes 0.6-0.9 mm long; larger autumnal leaf blades to 3 cm by 1 mm  Dichanthelium aciculare

............ ............ ..14 Blades of mid-culm leaves lanceolate, thin or firm but not stiff, usually < 10× as long as wide, usually 7 mm or more wide when as much as 8 cm long.

............ ............ ............ ....21 Spikelet bases strongly attenuate, apices blunt to beaked; first glume (2/5-) ½-2/3 (-4/5) as long as spikelet, attached 0.3-0.5 (-0.7) mm below expansion of second glume and sterile lemma  Dichanthelium malacon

............ ............ ............ ....21 Spikelet bases not noticeably attenuate, apices rounded to acute; first glume < ½ as long as spikelet, attached < 0.3 mm below expansion of second glume and sterile lemma.

............ ............ ............ ......22 Spikelets 2.9-3.8 mm long, broadly elliptic, rounded at the summit, with broad and thick nerves  Dichanthelium scribnerianum

............ ............ ............ ......22 Spikelets 2.1-2.9 mm long, elliptic or obovate, rounded or pointed at the summit, the nerves often raised, but not broad and thick.

............ ............ ............ ........23 Culm internodes and sheaths glabrous, sparsely pilose, or villous.

............ ............ ............ ..........24 Culms 50-150 cm long; at least lowest sheaths papillose-pilose.

............ ............ ............ ............ 25 Lowest internodes usually purple-villous; nodes retrorsely (-spreading) bearded; spikelets 1.8-2.2 mm long, moderately densely pubescent; first glume broadly ovate, subacute, 0.4-0.7 mm long, usually < 1.5 × as long as wide; second glume about as long as sterile and fertile lemmas; fertile lemma 1.7-1.8 mm long  Dichanthelium mundum

............ ............ ............ ............ 25 Lowest internodes glabrous or pubescent or spreading villous, usually not purplish; nodes glabrous, sparsely pubescent to sparsely bearded; spikelets 2.2-2.8 (-3.0) mm long, sparsely pubescent; first glume 0.7-1.1 mm long, usually > 1.5 × as long as wide; second glume shorter than sterile and fertile lemmas; fertile lemma 2.1-2.2 mm long  Dichanthelium recognitum

............ ............ ............ ..........24 Culms 18-100 cm long; sheaths glabrous or the lowest softly pubescent but not papillose-pilose.

............ ............ ............ ............ ..26 Spikelets strongly plano-convex when viewed laterally, obpyriform when viewed dorsally, (2.1-) 2.2-2.6 mm long; fertile lemma and palea papillose or smooth; first glume usually broadened above base, rounded or broadly tapered distally; glume bases often reddish-purple.

............ ............ ............ ............ ....27 Culm leaves 2-5 mm wide; fertile lemma and palea smooth, shiny  Dichanthelium patentifolium

............ ............ ............ ............ ....27 Culm leaves 4-12 mm wide; fertile lemma and palea papillose-roughened  Dichanthelium webberianum

............ ............ ............ ............ ..26 Spikelets biconvex to elliptic when viewed laterally, variously shaped but not obpyriform when viewed dorsally; fertile lemma and palea smooth or reticulate (or papillose in D. lucidum with weak and sprawling culms and spikelets 1.7-2.3 mm long); first glume usually tapered from near base to apex; glume bases various  Key to Dichanthelium, Key to the Dichanthelium dichotomum group

............ ............ ............ ........23 Culm internodes crisp-puberulent (sparsely so in D. webberianum); sheaths puberulent or glabrous.

............ ............ ............ ............ ......28 Spikelets elliptic, sub-acute to pointed, greenish or faintly purple-tinged basally  Dichanthelium commutatum ssp. ashei

............ ............ ............ ............ ......28 Spikelets strongly plano-convex when viewed laterally, obpyriform when viewed dorsally, broadly rounded, usually markedly reddish-purple basally.

............ ............ ............ ............ ........29 Fertile lemma and palea papillose; spikelets 2.2-2.6 mm long; lower culm blades 6-12 mm wide, glabrous  Dichanthelium webberianum

............ ............ ............ ............ ........29 Fertile lemma and palea smooth (minutely reticulate but not papillose); spikelets (1.8) 1.9-2.2 (-2.5) mm long; lower culm blades 2-8 mm wide, glabrous, glabrate, or puberulent (especially abaxially)  Dichanthelium portoricense ssp. patulum

Key to Dichanthelium, Key G: Spikelets 0.8-2.0 mm long, lower culm internodes glabrous

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