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1 Leaves either 3-foliolate or palmately 4-11-foliolate (all the leaflets attached at a common point, or the leaflets slightly pedate in Helleborus foetidus in RANUNCULACEAE). |
..2 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela |
..2 Inflorescence, flower, and fruit structure various, but not with the combination of features as above. {add under 2b [ Cynosciadium] APIACEAE} |
....3 Leaflets obviously and sharply serrate; pistils 1 or 5-many; fruit an achene, or an aggregate of achenes, drupelets, or follicles. |
......4 Leaflets 7-11, slightly pedate in their arrangement, evergreen |
......4 Leaflets 3-9, palmate, deciduous |
........5 Pistil 1; fruit an achene |
........5 Pistils 5-many; fruit an aggregate of achenes, drupelets, or follicles |
....3 Leaflets entire, finely denticulate, or very obscurely toothed (or irregularly serrate or lobed in Cardamine in BRASSICACEAE); pistil 1; fruit simple, a legume, capsule, silique, or berry. |
..........6 Corolla bilaterally symmetrical; petals 5; fruit a legume; carpel 1 |
..........6 Corolla radially symmetrical; petals 4 or 5; fruit either an elongate capsule or a berry; carpels 1, 2, or 5. |
............ 7 Leaflets radially arranged at the summit of the petiole, not differentiated in size or placement into a terminal leaflet and 2 lateral leaflets; leaflets prominently notched at the apex; petals 5, yellow; inflorescence axillary, cymose or umbelliform; carpels 5 |
............ 7 Leaflets differentiated in size and placement into a terminal leaflet and 2 or more lateral leaflets; leaflets not regularly notched at the apex (a few may be slightly retuse); petals 4, white, pink, or yellow; inflorescence terminal and racemose; carpels 1 or 2. |
............ ..8 Carpels 2; fruit a red berry; petals connate, purplish-blue |
............ ..8 Carpel 1; fruit a capsule; petals separate, white, pink, or yellow. |
............ ....9 Stem leaves 1-3, alternate [or whorled or opposite]; leaflets 3, irregularly serrate, lacerate, or additionally divided or lobed; fruit a silique; carpels 2 |
............ ....9 Stem leaves >3, alternate; leaflets (1-) 3-7, each entire or obscurely toothed; fruit a capsule; carpel 1 |
1 Leaves either 1-pinnately compound (all leaflets attached to a central rachis) or more complexly compound (with several orders of branching, some leaflets at least attached to second-order branches from the rachis). |
............ ......10 Inflorescence an involucrate head subtended by phyllaries, heads solitary or many, variously arrayed in secondary inflorescences; fruit a cypsela; ovary inferior |
............ ......10 Inflorescence various, but not as above; fruit various, not as above; ovary superior. |
............ ........11 Leaves 1-pinnately compound (all leaflets attached to a central rachis). |
............ ..........12 Flowers bilaterally symmetrical, papilionaceous; fruit a legume; leaves even-pinnately or odd-pinnately compound, the terminal leaflet sometimes replaced by a tendril; leaflets entire or at most minutely denticulate |
............ ..........12 Flowers radially symmetrical (or barely bilaterally symmetrical in Erodium in GERANIACEAE); fruit a capsule, capsular but of 5 mericarps, or an aggregate of achenes, nutlets, or follicles (in some cases the # of pistils from many down to 2 or even 1); leaves odd-pinnately compound, never with tendrils; leaflets serrate (or entire to shallowly lobed in Polemonium in POLEMONIACEAE, Cardamine in BRASSICACEAE, and Floerkea in LIMNANTHACEAE). |
............ ............ 13 Pistils many (only 1-2 in Agrimonia, Poteridium, Poterium, and Sanguisorba); fruit an aggregate of achenes, nutlets, or follicles; hypanthium present; stamens 5-many (only 4 in Poteridium and Sanguisorba) |
............ ............ 13 Pistil 1 (or deeply 2-3-lobed in Floerkea in LIMNANTHACEAE); fruit a silique, capsule, schizocarp of 2-3 mericarps, or a capsular schizocarp of 5 mericarps ( Erodium in GERANIACEAE); hypanthium absent; stamens 3-6. |
............ ............ ..14 Petals 3 |
............ ............ ..14 Petals 4-5. |
............ ............ ....15 Petals 4, distinct; stamens 6; inflorescence a terminal raceme; fruit a silique/silicle |
............ ............ ....15 Petals 5, fused (distinct in Erodium in GERANIACEAE); stamens 5; inflorescence axillary or terminal, cymose, consisting of subcapitate, umbel-like, or helicoid cymes; fruit either a capsule, or a capsular schizocarp of 5 mericarps ( Erodium in GERANIACEAE). |
............ ............ ......16 Flowers slightly bilaterally symmetrical (2 of the petals of different size than the other 3); fruit a capsular schizocarp of 5 mericarps; carpels 5 |
............ ............ ......16 Flowers radially symmetrical; fruit either a loculicidal capsule or a berry; carpels 2 or 3. |
............ ............ ........17 Fruit a berry; fresh plant rankly fragrant |
............ ............ ........17 Fruit a capsule; fresh plant not aromatic. |
............ ............ ..........18 Capsule 2-valvate; carpels 2; leaflets prominently serrate or with some tooth-like sublobes |
............ ............ ..........18 Capsule 3-valvate; carpels 3; leaflets with entire margins |
............ ........11 Leaves more complexly compound (with 2 or more orders of branching, some leaflets at least attached to second-order branches from the rachis). |
............ ............ ............ 19 Leaves 2× even-pinnate; flowers in spikes or spherical heads; fruit a legume or loment. |
............ ............ ............ 19 Leaves either 2× odd-pinnate or more complexly 2-4× ternately or ternately-pinnately compound; flowers in various inflorescences; fruits various (not legumes or loments). |
............ ............ ............ ..20 Leaf segments linear, less than 2 mm wide. |
............ ............ ............ ....21 Inflorescence an umbel; ovary inferior, of 2 fused carpels; fruit a schizocarp of 2 mericarps |
............ ............ ............ ....21 Inflorescence either a terminal solitary flower or terminal raceme or panicle; ovary superior, either of 2 fused carpels or of 1-5or many distinct 1-carpellate pistils; fruit either a capsule or an aggregate of follicles or achenes. |
............ ............ ............ ......22 Ovary of 2 fused carpels; fruit a capsule (1-seeded and indehiscent in Fumaria) |
............ ............ ............ ......22 Ovary of 1-5or many distinct 1-carpellate pistils; fruit an aggregate of follicles or achenes |
............ ............ ............ ..20 Leaf segments broader, lanceolate, ovate, or elliptic, > 5 mm wide. |
............ ............ ............ ........23 Herbaceous vine climbing by axillary tendrils; stamens 8 |
............ ............ ............ ........23 Erect or sprawling herb; stamens 5-6 or >15. |
............ ............ ............ ..........24 Leaflets sharply serrate, with usually many teeth on each leaflet side, the total number of “points” per leaflet > 10. |
............ ............ ............ ............ 25 Inflorescence an umbel; ovary inferior, of 2 fused carpels; fruit a schizocarp of 2 mericarps; inflorescence an umbel |
............ ............ ............ ............ 25 Inflorescence a panicle or raceme; ovary superior, of 1-8 carpels; fruit an aggregate of follicles, a single follicle, or an indehiscent berry-like fruit. |
............ ............ ............ ............ ..26 Flowers bisexual; carpels 1-8; fruit an aggregate of follicles, a single follicle, or an indehiscent berry-like fruit |
............ ............ ............ ............ ..26 Flowers unisexual; carpels (in pistillate flowers) of 3 (-5) carpels; fruit an aggregate of follicles |
............ ............ ............ ..........24 Leaflets entire, or with 1-several, broad, obtuse, rounded, or broadly acute “sublobes”, especially towards the tip of the leaflet, the total number of “points” per leaflet < 10. |
............ ............ ............ ............ ....27 Inflorescence an umbel; ovary inferior, of 2 fused carpels; fruit a schizocarp of 2 mericarps |
............ ............ ............ ............ ....27 Inflorescence a raceme, panicle, or cyme; ovary superior, of either 1-2 fused carpels or of many separate 1-carpellate pistils. |
............ ............ ............ ............ ......28 Perianth bilaterally symmetrical, the corolla of 4 fused petals; plant a scandent vine or erect or sprawling herb |
............ ............ ............ ............ ......28 Perianth radially symmetrical, of 1-5 whorls of separate perianth parts; plant an erect herb. |
............ ............ ............ ............ ........29 Perianth of 4-5 whorls of 3 parts each (some of the whorls modified into nectaries); pistil 1, 1-carpellate; fruit a drupelike, blue, naked seed; largest leaflets > 6 cm long, obviously longer than wide |
............ ............ ............ ............ ........29 Perianth of 1 whorl; of 4-5 parts; pistils many, each 1-carpellate; fruit an aggregate of achenes or an aggregate of follicles; largest leaflets either < 6 cm long, or if longer than 6 cm, also about as wide as long |
1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela |
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.). |
..2 Perianth uniseriate (represented only by undifferentiated tepals or sepals) or completely absent; flowers usually unisexual, less commonly bisexual). |
....3 Inflorescence a cyathium, consisting of a single pistillate flower (reduced to a single 3-carpellate pistil) and 2 or more staminate flowers (each reduced to 1 stamen), borne in a cup-like involucre, the involucre bearing pointed or rounded glands, these sometimes brightly colored and petaloid, mimicking an individual flower (the cyathia then secondarily arranged in terminal cymes, or solitary and axillary, etc.); fresh plants with milky juice; fruit a 3-lobed, 3-locular capsule |
....3 Inflorescence not a cyathium (and staminate or bisexual flowers with > 1 stamen; fresh plants lacking milky juice (except Stillingia in EUPHORBIACEAE); fruit various, not as above. |
......4 Leaf margins toothed in some manner (serrate, dentate, crenate, etc.) |
........5 Leaf teeth rounded to subacute, resembling shallow lobes, irregular, few (mostly < 6 per leaf side). |
..........6 Fruit a single-seeded achene or utricle; [plants of various disturbed or saline, usually sunny habitats] |
..........6 Fruit a 3-lobed, circumscissilely dehiscent capsule; [plants native of rich moist shaded forests or exotics in suburban woodlands] |
........5 Leaf teeth sharp to crenate, not lobe-like, regular, many (mostly > 10 per leaf side). |
............ 7 Leaf bases cuneate |
............ 7 Leaf bases cordate to rounded. |
............ ..8 Styles 3; fruit a 3-lobed, 3-carpellate capsule (1 carpel sometimes aborting); inflorescence either a terminal or leaf opposed raceme, or a dense axillary condensed cyme with conspicuous toothed bracts subtending the flowers |
............ ..8 Styles 1 or 2; fruit either an achene or a multiple of achenes; inflorescence either an axillary dense cyme (almost a head), or an axillary spike with glomerules, or a terminal or axillary panicle. |
............ ....9 Styles 2; inflorescence a dense axillary cyme (almost a head); fruit a multiple of achenes; plant lacking stinging hairs; [exotic plant of weedy situations] |
............ ....9 Style 1; inflorescence an axillary spikes with glomerules, or a terminal or axillary panicle; plant either with stinging hairs or not; [plant a rare exotic ( Boehmeria nivea) or a native of moist forests ( Boehmeria cylindrica, Laportea)] |
......4 Leaf margins entire. |
............ ......10 Ovary inferior or half-inferior. |
............ ........11 Leaf base cordate; calyx 3-lobed, fused into a bilaterally symmetrical, curved brown or yellowish tube; fruit a capsule |
............ ........11 Leaf base cuneate, rounded, or truncate; calyx of 3-4-5 distinct sepals, radially symmetrical, white or yellow; fruit a dry, nutlike drupe or an achene. |
............ ..........12 Leaves subsessile or very short petiolate, elliptic or narrowly elliptic, broadest near the middle; [native] |
............ ..........12 Leaves distinctly petiolate, rhombic, widest near the base; [rarely naturalized exotics]. |
............ ............ 13 Inflorescence of a single axillary flower |
............ ............ 13 Inflorescence terminal, spikelike |
............ ......10 Ovary superior. |
............ ............ ..14 Inflorescence a leaf-opposed (sometimes apparently terminal) spike or raceme; flowers visually white from white petaloid sepals, white bracts, or white stamens. |
............ ............ ....15 Sepals present, 4 or 5; petaloid, white; carpels 1 to many (-12); stamens 4 to many (-25); fruit a berry or an apically 2-lobed achene (as in Petiveria); leaf bases cuneate or rounded (but not cordate); [Eudicots]. |
............ ............ ......16 Sepals 4; carpels 1; fruit subglobose berries or apically lobed achenes |
............ ............ ......16 Sepals 5; carpels 6-12, distinct; fruit an oblate berry |
............ ............ ....15 Sepals absent; carpels 3-4; stamens 2-6 (-8); fruit a capsule, a 1-seeded drupe, or a schizocarp of 3-4 mericarps; leaf bases cordate or subcordate; [Basal Angiosperms]. |
............ ............ ........17 Fruit a 1-seeded drupe; stamens 2; plants terrestrial or epiphytic |
............ ............ ........17 Fruit a capsule or schizocarp with 3-4 mericarps; stamens 3 or 6-8; plants terrestrial |
............ ............ ..14 Inflorescence not leaf-opposed, either simpler (single axillary or glomerules of flowers) or more complexly branched (axillary or terminal panicles or complex cymes); flowers white, reddish, scarious, or greenish. |
............ ............ ..........18 Stipules tubular, sheathing (= ocreae); flowers subtended by tubular, sheathing bracteoles (= ocreolae); nodes usually prominently swollen; perianth usually of 5-6 white to pink tepals |
............ ............ ..........18 Stipules not tubular or sheathing; flowers not subtended by ocreolae; nodes not swollen; perianth absent or of 3-5 sepals. |
............ ............ ............ 19 Inflorescence an terminal involucrate cluster; flowers bisexual; stamens 9 |
............ ............ ............ 19 Inflorescence axillary, or a terminal panicle or raceme that is not involucrate; stamens 3-6. |
............ ............ ............ ..20 Ovary 3-locular; styles 3, each bifid; fruit a capsule, with 6 seeds |
............ ............ ............ ..20 Ovary 1-locular; styles 1-3, not bifid; fruit a utricle or achene (1-seeded). |
............ ............ ............ ....21 Styles 1, stigma 1; flowers unisexual |
............ ............ ............ ....21 Styles 1-3, if style 1, then stigmas 3; flowers bisexual or unisexual. |
............ ............ ............ ......22 Tepals acute, scarious |
............ ............ ............ ......22 Tepals usually blunt, herbaceous |
..2 Perianth biseriate (represented by differentiated whorls of sepals and petals, the sepals usually green or drab in color, the petals often brightly colored); flowers nearly always bisexual (there are exceptions). |
............ ............ ............ ........23 Ovary inferior or half-inferior. |
............ ............ ............ ..........24 Petals connate. |
............ ............ ............ ............ 25 Inflorescence leaf-opposed, a dense, cylindrical spike |
............ ............ ............ ............ 25 Inflorescence various, either a terminal head, or axillary and solitary, or variously axillary or terminal and more diffuse. |
............ ............ ............ ............ ..26 Leaves toothed; flowers blue to white |
............ ............ ............ ............ ..26 Leaves entire; flowers white |
............ ............ ............ ..........24 Petals distinct. |
............ ............ ............ ............ ....27 Petals 5; stamens 5; inflorescence a head; fruit a schizocarp of 2 mericarps |
............ ............ ............ ............ ....27 Petals 4 7; stamens 6 or more; inflorescence various, not a head; fruit a capsule. |
............ ............ ............ ............ ......28 Petals 4-7; stamens 1× or 2× as many as the petals, 4-7, 8, 10, 12, or14; leaves herbaceous in texture |
............ ............ ............ ............ ......28 Petals 5 (or sometimes doubled in horticultural forms); stamens 6-40 (or more); leaves fleshy in texture |
............ ............ ............ ........23 Ovary superior. |
............ ............ ............ ............ ........29 Corolla bilaterally symmetrical, petals connate (except distinct in VIOLACEAE); fruit a capsule or legume (except a 1-seeded indehiscent spinose pod in Krameria in KRAMERIACEAE). |
............ ............ ............ ............ ..........30 Petals distinct, 5; carpels 3; fruit a 3-loculed capsule |
............ ............ ............ ............ ..........30 Petals connate (at least basally), 4, 5, 6, 7, or 8; carpels 1, 2, 4, 5, or 6 (rarely 3 in Reseda in RESEDACEAE); fruit a legume or 1-, 2-, or 5-loculed capsule (except a 1-seeded indehiscent pod in Krameria in KRAMERIACEAE). |
............ ............ ............ ............ ............ 31 Stamens 6-10 (-25), more than the number (4 or 5) of petals and the number (4 or 5) of the sepals; fruit a legume or a 1-6-carpellate capsule. |
............ ............ ............ ............ ............ ..32 Stamens fused, monadelphous or diadelphous. |
............ ............ ............ ............ ............ ....33 Stamens 10, monadelphous or diadelphous; fruit a legume |
............ ............ ............ ............ ............ ....33 Stamens 6-8, monadelphous; fruit a capsule |
............ ............ ............ ............ ............ ..32 Stamens distinct. |
............ ............ ............ ............ ............ ......34 Carpel 1; fruit a legume |
............ ............ ............ ............ ............ ......34 Carpels 3-6; fruit a capsule |
............ ............ ............ ............ ............ 31 Stamens 4-5, less than the number (5) of the petals; fruit a 2-5-carpellate capsule. |
............ ............ ............ ............ ............ ........35 Pistil 5-carpellate; capsule 5-locular, explosively dehiscent; inflorescence axillary, small clusters of flowers |
............ ............ ............ ............ ............ ........35 Pistil 2-carpellate; capsule 2 locular, opening gradually or not at all; inflorescence a terminal spike, raceme or panicle (or solitary, axillary flowers in Chaenorrhinum in PLANTAGINACEAE and Krameria in KRAMERIACEAE). |
............ ............ ............ ............ ............ ..........36 Fruit 1-seeded, indehiscent; sepals petaloid, red-purple; petals dimorphic, the upper 3 long-clawed, the lower 2 small, thickened, and glandlike |
............ ............ ............ ............ ............ ..........36 Fruit > 2-seeded, dehiscent; sepals sepaloid, green; petals not dimorphic. |
............ ............ ............ ............ ............ ............ 37 Stamens 5; corolla not spurred; capsule septicidal; pubescence of the stem and leaves either gland-tipped or dendritically branched |
............ ............ ............ ............ ............ ............ 37 Stamens 4; corolla with a distinct spur or sac at the base between the the 2 lower calyx lobes (except not spurred in Digitalis and Schwalbea); capsule loculicidal (only at the summit in Antirrhinum and Chaenorrhinum, and septicidal in Schwalbea); pubescence of the stem and leaves neither gland-tipped (except in Antirrhinum and Chaenorrhinum) nor dendritically branched. |
............ ............ ............ ............ ............ ............ ..38 Capsule septicidal; corolla not spurred |
............ ............ ............ ............ ............ ............ ..38 Capsule loculicidal; corolla spurred (except Digitalis) |
............ ............ ............ ............ ........29 Corolla radially symmetrical, petals connate or distinct; fruit various (including a capsule). |
............ ............ ............ ............ ............ ............ ....39 Petals distinct; stamens 5-many. |
............ ............ ............ ............ ............ ............ ......40 Pistils 4-10 (each 1-carpellate) in a ring, these sometimes fused basally, each with its own style/stigma; fruit either an aggregate of achenes or follicles or a 5 (-7) locular capsule. |
............ ............ ............ ............ ............ ............ ........41 Pistils 5 (-7); inflorescence a compound terminal cyme. |
............ ............ ............ ............ ............ ............ ..........42 Fruit an aggregate of follicles; leaves fleshy in texture; inflorescence; leaves entire of sparsely and coarsely serrate, with < 12 points per leaf; [plants primarily of dry habitats] |
............ ............ ............ ............ ............ ............ ..........42 Fruit a 5 (-7) locular capsule; leaves membranaceous in texture; leaves serrate, with > 20 points per leaf; [plants of wet habitats] |
............ ............ ............ ............ ............ ............ ........41 Pistils many; inflorescence of solitary flowers, or diffuse. |
............ ............ ............ ............ ............ ............ ............ 43 Leaves cuneate at the base; flowers in a diffuse inflorescence |
............ ............ ............ ............ ............ ............ ............ 43 Leaves cordate at the base; flowers solitary, on long pedicels |
............ ............ ............ ............ ............ ............ ......40 Pistil 1, with 1-to many carpels (in many MALVACEAE, the carpels loosely united in a ring [of more than 5] around the single style/stigma); fruit either a 1-, 2-, 3-, 5-, 6-, or 10-locular capsule, or a silique/silicle, or a ring of mericarps. |
............ ............ ............ ............ ............ ............ ............ ..44 Petals 4; sepals 4; stamens 6; fruit a silique/silicle |
............ ............ ............ ............ ............ ............ ............ ..44 Petals 5 (rarely 4 or 6); sepals 5 (rarely 4 or 6); stamens 5 (or multiples of 5), 6, or 12; fruit a capsule or a ring of mericarps. |
............ ............ ............ ............ ............ ............ ............ ....45 Stamens many, connate into a staminal tube; carpels 5-many; fruit a capsule or ring of mericarps; leaves usually serrate |
............ ............ ............ ............ ............ ............ ............ ....45 Stamens 5-many, distinct; carpels 2-5; fruit a capsule; leaves entire (serrate in Croton in EUPHORBIACEAE). |
............ ............ ............ ............ ............ ............ ............ ......46 Flowers unisexual; leaf vestiture of peltate scales and/or stellate hairs |
............ ............ ............ ............ ............ ............ ............ ......46 Flowers bisexual; leaf vestiture simple or stellate. |
............ ............ ............ ............ ............ ............ ............ ........47 Flowers 6-merous (the petals and sepals 6, the stamens 6 or 12); corolla pink or purplish (rarely white); fruit a septicidal capsule |
............ ............ ............ ............ ............ ............ ............ ........47 Flowers 5-merous (the petals and sepals 5, stamens 5 or various multiples of 5); corolla yellow, reddish, or blue; fruit a loculicidal or septicidal capsule. |
............ ............ ............ ............ ............ ............ ............ ..........48 Stamens 5; corolla yellow or blue; capsule either 10-locular and septicidal or 1-locular (with 3 carpels) and loculicidal. |
............ ............ ............ ............ ............ ............ ............ ............ 49 Capsule 10-locular (with 5 carpels) and septicidal |
............ ............ ............ ............ ............ ............ ............ ............ 49 Capsule 1-locular (with 3 carpels) and loculicidal |
............ ............ ............ ............ ............ ............ ............ ..........48 Stamens (5-) 10, 15, 20, 30 (-many); corolla white, pink, yellow, or reddish; capsule 3-, 5- (-10)-locular, loculicidal. |
............ ............ ............ ............ ............ ............ ............ ............ ..50 Stamens (5-) 10, 15, 20, 30 (-many); corolla yellow or reddish; capsule 3 (-10)-locular, loculicidal |
............ ............ ............ ............ ............ ............ ............ ............ ..50 Stamens 10; corolla white or pink; capsule 5-locular |
............ ............ ............ ............ ............ ............ ....39 Petals fused; stamens (4-) 5 (-7). |
............ ............ ............ ............ ............ ............ ............ ............ ....51 Pistils 2, united only by the style and stigma; fruit a schizocarp of 2 follicles (often single by abortion); plant with milky juice when fresh; leaves entire; inflorescence an umbel |
............ ............ ............ ............ ............ ............ ............ ............ ....51 Pistil 1 (of 2 or 3 fused carpels); fruit a capsule; plant lacking milky juice; leaves entire or serrate; inflorescence various (but not an umbel). |
............ ............ ............ ............ ............ ............ ............ ............ ......52 Ovary slightly to deeply 2-4-lobed; fruit a schizocarp of 4 mericarps or a drupe. |
............ ............ ............ ............ ............ ............ ............ ............ ........53 Ovary deeply 4-parted; style gynobasic; fruit a schizocarp of 4 mericarps |
............ ............ ............ ............ ............ ............ ............ ............ ........53 Ovary slightly 2-4-lobed, or not at all lobed; style terminal or reduced to a sessile terminal stigma; fruit a schizocarp of 4 mericarps, or a drupe |
............ ............ ............ ............ ............ ............ ............ ............ ......52 Ovary not lobed; fruit a capsule or berry. |
............ ............ ............ ............ ............ ............ ............ ............ ..........54 Leaves scale-like, 1-4.5 mm long, appressed to the stem; petals 4; stamens 4 |
............ ............ ............ ............ ............ ............ ............ ............ ..........54 Leaves larger (or only 2-8 mm long in Pyxidanthera in DIAPENSIACEAE, but then spreading); petals 5-7; stamens 5-7. |
............ ............ ............ ............ ............ ............ ............ ............ ............ 55 Plant a creeping subshrub (keyed here as a failsafe); leaves either 0.2-0.8 cm long and acicular, or 2-10 cm long and broadly ovate or elliptic. |
............ ............ ............ ............ ............ ............ ............ ............ ............ ..56 Leaves 0.2-0.8 cm long and acicular |
............ ............ ............ ............ ............ ............ ............ ............ ............ ..56 Leaves 2-10 cm long and broadly ovate or elliptic |
............ ............ ............ ............ ............ ............ ............ ............ ............ 55 Plant an herb, erect or sprawling; leaves > 1.5 cm long. |
............ ............ ............ ............ ............ ............ ............ ............ ............ ....57 Leaves cordate at the base; plant a twining vine |
............ ............ ............ ............ ............ ............ ............ ............ ............ ....57 Leaves cuneate to rounded at the base; plant an erect, sprawling, or reclining herb (twining in Solanum dulcamara in SOLANACEAE). |
............ ............ ............ ............ ............ ............ ............ ............ ............ ......58 Inflorescences (solitary or of several flowers) terminal on the stem. |
............ ............ ............ ............ ............ ............ ............ ............ ............ ........59 Corolla lobes longer than the fused corolla cup, blue, pink, or white; styles 2; herbage lacking stipitate glands; fresh plants not aromatic. |
............ ............ ............ ............ ............ ............ ............ ............ ............ ..........60 Flowers in axillary cymes; capsule subglobose; leaves 2-12 cm long |
............ ............ ............ ............ ............ ............ ............ ............ ............ ..........60 Flowers solitary or paired in the leaf axils; capsule cylindrical; leaves 0.5-8 cm long |
............ ............ ............ ............ ............ ............ ............ ............ ............ ........59 Corolla lobes very short, much shorter than the corolla cup or tube, sometimes barely perceptible and represented only by teeth on the edge of the corolla limb, white or pink; style 1; herbage often with stipitate glands; fresh plants often rankly aromatic |
............ ............ ............ ............ ............ ............ ............ ............ ............ ......58 Inflorescences (of solitary or several flowers) axillary or lateral on the stem. |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ 61 Flowers sessile or very-short pedicelled, solitary in the leaf axils. |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ..62 Stamens alternate with the corolla lobes; flower ca. 10 mm in diameter |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ..62 Stamens opposite the corolla lobes; flower ca. 1 mm in diameter |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ 61 Flowers either solitary and obviously pedicelled, or several in an axillary or lateral inflorescence. |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ....63 Corolla lobes longer than the fused corolla cup, blue |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ....63 Corolla lobes very short, much shorter than the corolla cup or tube, sometimes barely perceptible and represented only by teeth on the edge of the corolla limb, white, yellow, pink, various other colors (rarely including blue). |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ......64 Fruit a capsule, 4-seeded |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ......64 Fruit a berry or capsule, many-seeded |
1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela |
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.). |
..2 Perianth uniseriate, with only undifferentiated tepals; flowers many and small, greenish or brownish, inconspicuous individually; inflorescence of glomerules that are usually further aggregated into racemes or panicles; fruit an achene or utricle |
..2 Perianth biseriate, both sepals and petals present and differentiated (except uniseriate and of 2 white to cream-colored sepals 5-10 mm long in Macleaya in PAPAVERACEAE); flowers larger, usually with the petals prominently colored; inflorescence various, but not as above; fruit a capsule, silique/silicle, or a schizocarp of 2 mericarps. |
....3 Corolla bilaterally symmetrical and the petals connate into a tube (or the corolla 2-lipped but the corolla lobes twisted so as to make the flower asymmetrical); stamens 4; fruit a 2-locular and loculicidal capsule opening by 2 valves |
....3 Corolla radially symmetrical and either connate into a tube or distinct (except Reseda in RESEDACEAE, with bilateral symmetry but separate petals); stamens 5 or more; fruit a silique/silicle, a schizocarp of 4 mericarps, or a 1-, 3-, or 4-locular capsule (2-locular in Ipomoea in CONVOLVULACEAE and Glaucium in PAPAVERACEAE), opening variously. |
......4 Ovary inferior; fruit either a schizocarp of 2 mericarps or a 4-locular capsule. |
........5 Flowers 5-merous, aggregated into a head; fruit a schizocarp of 2 mericarps |
........5 Flowers 4-merous, in a diffuse inflorescence; fruit a 4-locular capsule |
......4 Ovary superior; fruit either a silique/silicle, or a 1-, 2-, or 3-locular capsule, or a berry. |
..........6 Sepals and petals of different numbers, the sepals 2-3, the petals 0, 4, or 6; stamens many |
..........6 Sepals and petals the same number, 4-8 each; stamens 5 or 6 (10-25 in Reseda in RESEDACEAE). |
............ 7 Corolla bilaterally symmetrical; stamens 10-25 |
............ 7 Corolla radially symmetrical; stamens 5 or 6. |
............ ..8 Petals 4, distinct; stamens 6; fruit a silique/silicle |
............ ..8 Petals 5, connate into a tube; stamens 5; fruit either a capsule or a berry. |
............ ....9 Plant a twining vine. |
............ ......10 Corolla with a long tube, much longer than the lobes, scarlet, white, pink, or blue; leaves shallowly to deeply pinnately parted into 3-many lobes |
............ ......10 Corolla with a short tube, the lobes longer than the tube, purplish; leaves with a single large terminal l lobe, and 2 small basal lobes (these almost separate as leaflets) |
............ ....9 Plant an erect or sprawling herb. |
............ ........11 Fruit a capsule, 1-locular; corolla white, pink, lavender, or blue, the tube short (< 4 mm long), the lobes flaring, the corolla < 15 mm long or wide |
............ ........11 Fruit either a capsule, 2- or-3 locular, or a berry; corolla scarlet, blue, white, yellow, greenish-yellow, or purple, the tube long (>10 mm long) and cylindrical, the corolla > 10 mm long or wide. |
............ ..........12 Stigmas 3; fruit a capsule with 3 valves; leaf lobes 0.5-5 mm wide, often themselves lobed, the sinuses very deep, often nearly to the leaf midvein |
............ ..........12 Stigmas 2; fruit either a capsule with 2 valves or a berry; leaf lobes > 5 mm wide, not lobed themselves, the sinuses shallow, often < 1/2 way to the midvein |