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1 Leaves either 2-3-foliolate or palmately 4-11-foliolate (all the leaflets attached at a common point). |
..2 Leaves 2-foliolate; fruit a capsule, opening by a circumscissile lid |
..2 Leaves either 3-foliolate or palmately or pedately 4-11-foliolate. |
....3 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela |
....3 Inflorescence, flower, and fruit structure various, but not with the combination of features as above. |
......4 Inflorescence an umbel; ovary inferior; fruit a mericarp of 2 schizocarps |
......4 Inflorescence various, usually not an umbel (sometimes an umbel in Oxalis in OXALIDACEAE); ovary superior; fruit an aggregate, legume, berry, or 2-valved capsule. |
........5 Leaflets either entire or barely and very shallowly crenulate or notched at the tip (but otherwise entire). |
..........6 Inflorescence a spadix, surrounded by a spathe; fruit a berry; [Monocots {illogically keyed here because of the likelihood of being mistaken for a dicot}] |
..........6 Inflorescence a raceme or umbel, not surrounded by a spathe; fruit a capsule or legume; [Eudicots]. |
............ 7 Flowers bilaterally symmetrical; fruit a legume; [plant of uplands] |
............ 7 Flowers radially symmetrical; fruit a 2-valved or 5-valved capsule; [plant of uplands or wetlands] |
............ ..8 Leaflets not notched at the tip; flowers white; [plants of saturated saturated or ponded wetlands] |
............ ..8 Leaflets notched at the tip; flowers pink, white, or yellow; [plants of uplands or temporarily flooded wetlands] |
........5 Leaflets serrate, serrulate, or cleft. |
............ ....9 Petals 4; stamens 6; fruit a silique |
............ ....9 Petals 5 or more; stamens 10 or more; fruit either a legume or an aggregate of achenes or follicles |
............ ......10 Stamens many, fused into a staminal tube; carpels 10-20, in a ring; pubescence stellate (sometimes mixed with simple hairs) |
............ ......10 Stamens 10-many, separate, or fused but not all into a staminal tube; carpel either 1 (FABACEAE), or 3-7 in a ring (RANUNCULACEAE), or many and spirally arranged on a conical receptacle (RANUNCULACEAE or ROSACEAE) |
............ ........11 Leaflets serrulate; flowers bilaterally symmetrical; fruit a legume; corolla variously colored, including white |
............ ........11 Leaflets serrate; flowers radially symmetrical; fruit an aggregate of achenes or of follicles; corolla white or yellowish or greenish. |
............ ..........12 Fruit an aggregate of follicles |
............ ..........12 Fruit an aggregate of achenes (borne on a fleshy, expanded receptacle in Fragaria and some Potentilla) |
............ ............ 13 Flowers lacking a hypanthium |
............ ............ 13 Flowers with a hypanthium |
1 Leaves 1-pinnately compound (all leaflets attached to a central rachis) or more complexly compound (with several orders of branching, some leaflets at least attached to second-order branches from the rachis). |
............ ............ ..14 Leaves 1-pinnately compound (all leaflets attached to a central rachis). |
............ ............ ....15 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela |
............ ............ ....15 Inflorescence, flower, and fruit structure various, but not with the combination of features as above. |
............ ............ ......16 Flowers bilaterally symmetrical; fruit a legume |
............ ............ ......16 Flowers radially symmetrical; fruit a silique/silicle, or a schizocarp of mericarps, or an achene. |
............ ............ ........17 Petals 4; stamens 6; fruit a silique/silicle |
............ ............ ........17 Petals 0 or 5 (if 0, the sepals petaloid); stamens 2, 4, 5, or many. |
............ ............ ..........18 Stamens 5; fruit a schizocarp of 2 mericarps |
............ ............ ..........18 Stamens 2, 4, or many; fruit an achene |
............ ............ ..14 Leaves more complexly compound (with 2 or more orders of branching, some leaflets at least attached to second-order branches from the rachis). |
............ ............ ............ 19 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela |
............ ............ ............ 19 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head subtended by bracts, but then with other features differing, such as stamens 4, or green calyx present, or petals separate, or fruit a schizocarp of mericarps, etc.). |
............ ............ ............ ..20 Leaf segments or ultimate lobes linear or lanceolate, > 2× as long as wide, < 4 mm wide. |
............ ............ ............ ....21 Inflorescence an umbel; ovary inferior; fruit a mericarp of 2 schizocarps |
............ ............ ............ ....21 Inflorescence various, but not an umbel; ovary superior; fruit an aggregate of follicles or an elongate capsule. |
............ ............ ............ ......22 Carpels 2, fused; fruit an elongate capsule; flowers bilaterally symmetrical |
............ ............ ............ ......22 Carpels 5-10 or many, separate; fruit an aggregate; flower radially symmetrical |
............ ............ ............ ..20 Leaf segments or ultimate lobes ovate or elliptic, < 3× as long as wide, > 5 mm wide. |
............ ............ ............ ........23 Inflorescence an umbel; ovary inferior; fruit a mericarp of 2 schizocarps or a 5-seeded drupe. |
............ ............ ............ ..........24 Fruit a schizocarp of 2 mericarps |
............ ............ ............ ..........24 Fruit a 5-seeded drupe |
............ ............ ............ ........23 Inflorescence various, but not an umbel; fruit an aggregate of follicles or achenes, an elongate capsule, or a naked seed resembling a drupe. |
............ ............ ............ ............ 25 Leaflets with < 10 ultimate ‘points’ (lobe or tooth terminations), these rounded to broadly acute, often large in comparison to the leaflet and appearing as “sublobes”; pistil 1 or 4-many. |
............ ............ ............ ............ ..26 Corolla bilaterally symmetrical; fruit an elongate capsule; [cultivated exotic, rarely persistent near gardens] |
............ ............ ............ ............ ..26 Corolla radially symmetrical; fruit an aggregate of follicles or achenes, or a naked seed resembling a drupe; [native plants of moist to dry forests and rock outcrops]. |
............ ............ ............ ............ ....27 Leaflets 5-8 cm long, obviously longer than broad; pistil 1; fruit a naked blue seed resembling a drupe; flowers mainly 3-merous |
............ ............ ............ ............ ....27 Leaflets 1-6 cm long, about as long as broad if > 4 cm long; pistils 4-many; fruit an aggregate of follicles or achenes; flowers mainly 4-5-merous |
............ ............ ............ ............ 25 Leaflets with >11 ultimate ‘points’ (lobe or tooth terminations), these acuminate to acute; pistils 1-8. |
............ ............ ............ ............ ......28 Pubescence of the stem and lower leaf surface glandular; flowers unisexual, on the same plant (monoecious); stamens 10; pistils 2, partly fused; fruit an aggregate of follicles |
............ ............ ............ ............ ......28 Pubescence of the stem and lower leaf surface non-glandular (or absent); flowers either bisexual (the plants hermaphroditic), or unisexual and the male and female flowers on separate plants (the plants dioecious); stamens 15 or more; pistils 1-8, separate; fruit an aggregate of follicles, a follicle, or a red or white berry. |
............ ............ ............ ............ ........29 Flowers bisexual (plants hermaphroditic); carpels 1-8 per flower; inflorescence a raceme, or a panicle of racemes with just a few branches; fruit an aggregate of follicles, a follicle, or a red or white berry |
............ ............ ............ ............ ........29 Flowers unisexual (plants dioecious); carpels 3-4 per pistillate flower; inflorescence a panicle of racemes, with numerous branches; fruit an aggregate of follicles |
1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela |
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head subtended by bracts, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or petals separate, or fruit a schizocarp of mericarps, etc.). |
..2 Basal leaves 2-lobed, pinnately lobed, or palmately lobed (not considering cordate, hastate, or auriculate leaf bases as “lobed”). |
....3 Basal leaves 2-lobed, hinged between the lobes, each lobe with stiff, marginal, eyelash-like bristles; [Coastal Plain of NC and SC, rarely planted and weakly naturalized elsewhere] |
....3 Basal leaves 3-many-lobed, palmately or pinnately; [collectively widespread]. |
......4 Leaf lobing pinnate. |
........5 Gynoecium of separate pistils (each with a single carpel); fruit an aggregate |
........5 Gynoecium of a single pistil (with 2, rarely more, carpels); fruit simple. |
..........6 Stamens many; sepals 2, petals 4; fresh plants with yellow, orange, or white milky juice |
..........6 Stamens 4, 5, or 6; sepals 4 or 5; petals 4 or 5. |
............ 7 Petals 4, distinct; stamens 6 |
............ 7 Petals 5, fused; stamens 2, 4, or 5. |
............ ..8 Corolla radially symmetrical; stamens 5 |
............ ..8 Corolla 2-lipped, bilaterally symmetrical or asymmetrical; stamens 2 or 4. |
............ ....9 Corolla lobes not twisted, the flower bilaterally symmetrical; stamens 2 |
............ ....9 Corolla lobes twisted so as to make the flower asymmetrical; stamens 4 |
......4 Leaf lobing palmate. |
............ ......10 Petiole attachment peltate. |
............ ........11 Leaves < 10 cm in diameter |
............ ........11 Leaves > 15 cm in diameter |
............ ......10 Petiole attachment marginal. |
............ ..........12 Ovary inferior. |
............ ............ 13 Petals 4; stamens 8; fruit a capsule |
............ ............ 13 Petals 5; stamens 5; fruit a schizocarp of 2 mericarps. |
............ ............ ..14 Fruit tuberculate; leaves 3-lobed |
............ ............ ..14 Fruit smooth; leaves with 5 or more lobes |
............ ..........12 Ovary superior, or half-inferior by fusion of a hypanthium a part of the way up the ovary wall. |
............ ............ ....15 Gynoecium of separate pistils (each with a single carpel); fruit an aggregate. |
............ ............ ......16 Perianth of 5 green sepals and 5 colored petals. |
............ ............ ........17 Carpels partly fused, arrayed in a ring of 10-20 |
............ ............ ........17 Carpels separate, spiral, many |
............ ............ ......16 Perianth of a single whorl of 3-12 petaloid sepals (the petals absent or small and rudimentary). |
............ ............ ..........18 Leaves 2, the single flower terminal and associated with the upper leaf; fruit an aggregate of berries |
............ ............ ..........18 Leaves normally > 2, flowers not as above; fruit an aggregate of achenes, utricles, or follicles |
............ ............ ....15 Gynoecium of a single pistil (with 1-5 carpels); fruit simple. |
............ ............ ............ 19 Hypanthium present, partially fused or not fused to the pistil; ovary partially inferior to superior |
............ ............ ............ 19 Hypanthium absent; ovary superior. |
............ ............ ............ ..20 Petals connate at base; leaves sharply toothed |
............ ............ ............ ..20 Petals distinct; leaves with rounded lobes or teeth. |
............ ............ ............ ....21 Corolla radially symmetrical; petals 8-12; capsule fusiform, narrowed to both ends, > 5× as long as wide |
............ ............ ............ ....21 Corolla bilaterally symmetrical; petals 5; capsule ovoid, < 2× as long as wide |
..2 Basal leaves not lobed, at most serrate or crenate (and sometimes also cordate, hastate, auriculate, or peltate at the base). |
............ ............ ............ ......22 Basal leaves petiolate, the blade with a cordate, hastate, auriculate, or peltate base. |
............ ............ ............ ........23 Leaf margins entire. |
............ ............ ............ ..........24 Inflorescence a terminal and/or axillary raceme, panicle, or cyme of many small flowers; fruit an achene; perianth uniseriate, of 0, 4-5, or 6 tepals. |
............ ............ ............ ............ 25 Flowers unisexual; staminate flowers 4-5 tepals, pistillate flowers lacking a perianth |
............ ............ ............ ............ 25 Flowers bisexual; flowers with 6 tepals |
............ ............ ............ ..........24 Inflorescence either a terminal spike, or a 1-7-flowered terminal cyme, or of a solitary axillary or terminal flower; fruit various; perianth biseriate (of differentiated sepals and petals (except uniseriate, of 3 fused sepals in ARISTOLOCHIACEAE). |
............ ............ ............ ............ ..26 Flowers bilaterally symmetrical; inflorescence a terminal spike (with > 20 flowers); petals 4, usually scarious, transparent; sepals 4, green; stamens 4 |
............ ............ ............ ............ ..26 Flowers radially symmetrical; inflorescence either of a solitary flower or of a 1-7-flowered terminal cyme; petals 5, 8-12, or 0; sepals 5 (green), 3 (brown), or 5-9 (yellow); stamens 5, 12, or many. |
............ ............ ............ ............ ....27 Gynoecium of separate pistils (each with a single carpel); fruit an aggregate of achenes or follicles; flowers bright yellow, either of 5-9 distinct petaloid sepals, or of 8-12 distinct petals subtended by 3-4 green distinct sepals |
............ ............ ............ ............ ....27 Gynoecium either of a single pistil with 6 carpels or of a single pistil with 4 carpels or of 2 nearly separate carpels; fruit a simple capsule (or deeply 2-lobed); flowers white, brown, or greenish, either of 5 fused or distinct white petals and 5 fused or distinct green sepals, or of 3 fused brown or greenish petaloid sepals. |
............ ............ ............ ............ ......28 Flowers brown or green, of 3 fused brown or greenish petaloid sepals (and 0 petals); carpels 6; stamens 12; leaves 4-10 cm wide |
............ ............ ............ ............ ......28 Flowers white, of 5 white or whitish petals and 5 green sepals; carpels 2; stamens 5; leaves 1-12 (-15) cm wide |
............ ............ ............ ............ ........29 Petals separate; sepals separate; plant glabrous |
............ ............ ............ ............ ........29 Petals fused; sepals fused; carpels 2; plant pubescent. |
............ ............ ............ ............ ..........30 {XXXX}; [common, widespread in our area] |
............ ............ ............ ............ ..........30 {YYYY}; [rare exotic] |
............ ............ ............ ........23 Leaf margins crenate, serrate, or incised. |
............ ............ ............ ............ ............ 31 Gynoecium of separate pistils (each with a single carpel); fruit an aggregate; perianth of 5 green sepals and 5 colored petals. |
............ ............ ............ ............ ............ ..32 Carpels 10-20, partly fused, arrayed in a ring; petals white, pink, red, or purplish |
............ ............ ............ ............ ............ ..32 Carpels many, separate, spiral; petals yellow or white. |
............ ............ ............ ............ ............ ....33 Flowers lacking a hypanthium; fruit an aggregate of achenes or aggregate of follicles |
............ ............ ............ ............ ............ ....33 Flowers with a hypanthium; fruit an aggregate of drupelets or aggregate of achenes |
............ ............ ............ ............ ............ 31 Gynoecium of a single pistil (with 1-5 carpels); fruit simple. |
............ ............ ............ ............ ............ ......34 Flowers bilaterally symmetrical; inflorescence of a solitary flower; fruit a 3-locular capsule |
............ ............ ............ ............ ............ ......34 Flowers radially symmetrical; inflorescence an umbel (or composite of umbelliform units, or a terminal panicle. |
............ ............ ............ ............ ............ ........35 Ovary superior; inflorescence a terminal panicle or terminal raceme. |
............ ............ ............ ............ ............ ..........36 Inflorescence a terminal raceme; perianth of 4 green sepals and 4 white petals; fruit a silique/silicle; fresh foliage in spring and summer with a strong garlic odor; larger leaves < 10 cm in diameter |
............ ............ ............ ............ ............ ..........36 Inflorescence a terminal panicle; perianth of 6 cream-colored petaloid sepals; fruit a winged achene; fresh foliage lacking a garlic odor; larger leaves typically > 30 cm in diameter |
............ ............ ............ ............ ............ ........35 Ovary inferior; inflorescence an umbel (or a composite of umbellate units); fruit a schizocarp of mericarps. |
............ ............ ............ ............ ............ ............ 37 Petiole attachment peltate |
............ ............ ............ ............ ............ ............ 37 Petiole attachment marginal (the blade cleft to the petiole. |
............ ............ ............ ............ ............ ............ ..38 Leaf blades longer than wide, sharply V-cleft at the base and otherwise shallowly denticulate |
............ ............ ............ ............ ............ ............ ..38 Leaf blades as broad or broader than long, cleft at the base and also irregularly serrate or crenate around the margin |
............ ............ ............ ......22 Basal leaves petiolate or not, with a truncate, rounded, or cuneate leaf base. |
............ ............ ............ ............ ............ ............ ....39 Leaves tubular, with a sutured ventral flange, erect or reclining, adapted as a pitfall for insects (flat, phyllodial leaves sometimes present as well, common in the winter in some species, such as S. oreophila) |
............ ............ ............ ............ ............ ............ ....39 Leaves flat, not sutured into a tubular shape. |
............ ............ ............ ............ ............ ............ ......40 Stem leaves opposite; perianth 5-merous, at least the corolla bilaterally symmetrical (barely so in VALERIANACEAE), or the parts curved so as to be asymmetrical ( Pedicularis in OROBANCHACEAE); stamens 2, 3, or 4. |
............ ............ ............ ............ ............ ............ ........41 Ovary inferior; stamens 3 |
............ ............ ............ ............ ............ ............ ........41 Ovary superior; stamens 2 or 4. |
............ ............ ............ ............ ............ ............ ..........42 Corolla narrowly tubular, the five lobes flaring at nearly 90 degrees and nearly radially symmetrical |
............ ............ ............ ............ ............ ............ ..........42 Corolla distinctly 2-lipped (with prominently large upper and lower corolla lobes) or hooded (the upper lip hood-like), distinctly bilaterally symmetrical, or the lobes twisted so as to make the corolla asymmetrical. |
............ ............ ............ ............ ............ ............ ............ 43 Corolla yellow, the upper lip often slightly to strongly maroon, hooded but the corolla lobes twisted so as to make the flower asymmetrical |
............ ............ ............ ............ ............ ............ ............ 43 Corolla white, lavender, or blue, 2-lipped and bilaterally symmetrical. |
............ ............ ............ ............ ............ ............ ............ ..44 Sepals separate to the base or nearly so, not forming a tube |
............ ............ ............ ............ ............ ............ ............ ..44 Sepals connate for at least 0.3× their length to form a tube |
............ ............ ............ ............ ............ ............ ......40 Stem leaves alternate; perianth radially symmetrical (less commonly bilaterally symmetrical); stamens 5, 6-8, 9, 10 (rarely 4). |
............ ............ ............ ............ ............ ............ ............ ....45 Ovary inferior (or half-inferior in Samolus). |
............ ............ ............ ............ ............ ............ ............ ......46 Perianth 4-merous; stamens 8 |
............ ............ ............ ............ ............ ............ ............ ......46 Perianth 5-merous; stamens 5. |
............ ............ ............ ............ ............ ............ ............ ........47 Inflorescence an umbel; fruit a schizocarp of 2 mericarps |
............ ............ ............ ............ ............ ............ ............ ........47 Inflorescence an axillary or terminal raceme; fruit a capsule |
............ ............ ............ ............ ............ ............ ............ ....45 Ovary superior. |
............ ............ ............ ............ ............ ............ ............ ..........48 Pistils many, each with a single carpel; fruit an aggregate of achenes |
............ ............ ............ ............ ............ ............ ............ ..........48 Pistil 1, with 1-5 carpels; fruit simple (a legume, silique/silicle, capsule, utricle, or schizocarp of 4 nutlets). |
............ ............ ............ ............ ............ ............ ............ ............ 49 Corolla bilaterally symmetrical (barely so in Limosella in SCROPHULARIACEAE); stamens 2, 4, 6, 8, or 10. |
............ ............ ............ ............ ............ ............ ............ ............ ..50 Stamens 6-8 or 10. |
............ ............ ............ ............ ............ ............ ............ ............ ....51 Petals separate; stamens 10 |
............ ............ ............ ............ ............ ............ ............ ............ ....51 Petals fused; stamens 10 or 6-8. |
............ ............ ............ ............ ............ ............ ............ ............ ......52 Stamens 10, monadelphous |
............ ............ ............ ............ ............ ............ ............ ............ ......52 Stamens 6-8, epipetalous |
............ ............ ............ ............ ............ ............ ............ ............ ..50 Stamens 2 or 4. |
............ ............ ............ ............ ............ ............ ............ ............ ........53 Stamens 2 |
............ ............ ............ ............ ............ ............ ............ ............ ........53 Stamens 4. |
............ ............ ............ ............ ............ ............ ............ ............ ..........54 [plants of coastal wetlands] |
............ ............ ............ ............ ............ ............ ............ ............ ..........54 [plants of uplands or inland seeps or fens] |
............ ............ ............ ............ ............ ............ ............ ............ ............ 55 Flowers (and subtending bracts) red or yellow |
............ ............ ............ ............ ............ ............ ............ ............ ............ 55 Flowers purple, blue, or lavender |
............ ............ ............ ............ ............ ............ ............ ............ 49 Corolla radially symmetrical; stamens 5, 10, 4-6, or 9. |
............ ............ ............ ............ ............ ............ ............ ............ ............ ..56 Perianth of 6 tepals; stamens 4-6 or 9; carpels 3 |
............ ............ ............ ............ ............ ............ ............ ............ ............ ..56 Perianth of green sepals and more brightly colored petals; stamens 5 or 10; carpels 2, 3, 4, or 5. |
............ ............ ............ ............ ............ ............ ............ ............ ............ ....57 Leaves covered with sticky, gland-tipped hairs (often red), as flypaper traps for insects |
............ ............ ............ ............ ............ ............ ............ ............ ............ ....57 Leaves lacking sticky gland-tipped hairs. |
............ ............ ............ ............ ............ ............ ............ ............ ............ ......58 Fruit a schizocarp of 4 nutlets (ovary obviously 4-lobed in flower) |
............ ............ ............ ............ ............ ............ ............ ............ ............ ......58 Fruit a capsule or silique/silicle (or utricle in Limonium in PLUMBAGINACEAE). |
............ ............ ............ ............ ............ ............ ............ ............ ............ ........59 Inflorescence of a solitary, terminal flower; carpels 2-3 (-4). |
............ ............ ............ ............ ............ ............ ............ ............ ............ ..........60 Corolla with a long tube and flaring corolla lobes (united > ½ its length) |
............ ............ ............ ............ ............ ............ ............ ............ ............ ..........60 Corolla of separate petals or united only for a short length (< ¼ its length). |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ 61 Leaves serrate |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ 61 Leaves entire |
............ ............ ............ ............ ............ ............ ............ ............ ............ ........59 Inflorescence of several to many flowers; carpels 5 (3 in Galax in DIAPENSIACEAE). |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ..62 Inflorescence an umbel; petals recurved, pink to almost white |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ..62 Inflorescence a raceme or panicle. |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ....63 Fruit a silique/silicle; petals 4; stamens 6 |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ....63 Fruit either a capsule or a utricle; petals 5; stamens 5 or 10. |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ......64 Inflorescence a somewhat to very diffuse panicle, with 3 or more orders of branching, not giving at all the impression that the overall inflorescence is made of racemose units. |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ........65 Leaves entire; stamens 5; [plants of tidal marshes] |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ........65 Leaves serrate or crenate; stamens 10; [plants of various habitats, especially rock outcrops and bottomland forests and streambanks, never in tidal marshes] |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ......64 Inflorescence either a single terminal raceme (sometimes spike-like), or of 1 to several terminal and axillary racemes (these sometimes combined into a diffuse panicle, but one whose structure is clearly made up of many racemes). |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ..........66 Inflorescence of 1-several terminal and axillary racemes, the plant typically well-branched, especially from the base; stamens 5 |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ..........66 Inflorescence of a single, terminal raceme, the plant unbranched; stamens 10 (or 5, with 5 staminodes) |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ............ 67 Inflorescence spike-like, the flowers many (> 40), barely pedicelled; capsule 3-locular |
............ ............ ............ ............ ............ ............ ............ ............ ............ ............ ............ 67 Inflorescence a raceme, the flowers fewer (< 20) and distinctly pedicelled; capsule 5-locular |
1 Leaves either 3-foliolate or palmately 4-11-foliolate (all the leaflets attached at a common point, or the leaflets slightly pedate in Helleborus foetidus in RANUNCULACEAE). |
..2 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela |
..2 Inflorescence, flower, and fruit structure various, but not with the combination of features as above. {add under 2b [ Cynosciadium] APIACEAE} |
....3 Leaflets obviously and sharply serrate; pistils 1 or 5-many; fruit an achene, or an aggregate of achenes, drupelets, or follicles. |
......4 Leaflets 7-11, slightly pedate in their arrangement, evergreen |
......4 Leaflets 3-9, palmate, deciduous |
........5 Pistil 1; fruit an achene |
........5 Pistils 5-many; fruit an aggregate of achenes, drupelets, or follicles |
....3 Leaflets entire, finely denticulate, or very obscurely toothed (or irregularly serrate or lobed in Cardamine in BRASSICACEAE); pistil 1; fruit simple, a legume, capsule, silique, or berry. |
..........6 Corolla bilaterally symmetrical; petals 5; fruit a legume; carpel 1 |
..........6 Corolla radially symmetrical; petals 4 or 5; fruit either an elongate capsule or a berry; carpels 1, 2, or 5. |
............ 7 Leaflets radially arranged at the summit of the petiole, not differentiated in size or placement into a terminal leaflet and 2 lateral leaflets; leaflets prominently notched at the apex; petals 5, yellow; inflorescence axillary, cymose or umbelliform; carpels 5 |
............ 7 Leaflets differentiated in size and placement into a terminal leaflet and 2 or more lateral leaflets; leaflets not regularly notched at the apex (a few may be slightly retuse); petals 4, white, pink, or yellow; inflorescence terminal and racemose; carpels 1 or 2. |
............ ..8 Carpels 2; fruit a red berry; petals connate, purplish-blue |
............ ..8 Carpel 1; fruit a capsule; petals separate, white, pink, or yellow. |
............ ....9 Stem leaves 1-3, alternate [or whorled or opposite]; leaflets 3, irregularly serrate, lacerate, or additionally divided or lobed; fruit a silique; carpels 2 |
............ ....9 Stem leaves >3, alternate; leaflets (1-) 3-7, each entire or obscurely toothed; fruit a capsule; carpel 1 |
1 Leaves either 1-pinnately compound (all leaflets attached to a central rachis) or more complexly compound (with several orders of branching, some leaflets at least attached to second-order branches from the rachis). |
............ ......10 Inflorescence an involucrate head subtended by phyllaries, heads solitary or many, variously arrayed in secondary inflorescences; fruit a cypsela; ovary inferior |
............ ......10 Inflorescence various, but not as above; fruit various, not as above; ovary superior. |
............ ........11 Leaves 1-pinnately compound (all leaflets attached to a central rachis). |
............ ..........12 Flowers bilaterally symmetrical, papilionaceous; fruit a legume; leaves even-pinnately or odd-pinnately compound, the terminal leaflet sometimes replaced by a tendril; leaflets entire or at most minutely denticulate |
............ ..........12 Flowers radially symmetrical (or barely bilaterally symmetrical in Erodium in GERANIACEAE); fruit a capsule, capsular but of 5 mericarps, or an aggregate of achenes, nutlets, or follicles (in some cases the # of pistils from many down to 2 or even 1); leaves odd-pinnately compound, never with tendrils; leaflets serrate (or entire to shallowly lobed in Polemonium in POLEMONIACEAE, Cardamine in BRASSICACEAE, and Floerkea in LIMNANTHACEAE). |
............ ............ 13 Pistils many (only 1-2 in Agrimonia, Poteridium, Poterium, and Sanguisorba); fruit an aggregate of achenes, nutlets, or follicles; hypanthium present; stamens 5-many (only 4 in Poteridium and Sanguisorba) |
............ ............ 13 Pistil 1 (or deeply 2-3-lobed in Floerkea in LIMNANTHACEAE); fruit a silique, capsule, schizocarp of 2-3 mericarps, or a capsular schizocarp of 5 mericarps ( Erodium in GERANIACEAE); hypanthium absent; stamens 3-6. |
............ ............ ..14 Petals 3 |
............ ............ ..14 Petals 4-5. |
............ ............ ....15 Petals 4, distinct; stamens 6; inflorescence a terminal raceme; fruit a silique/silicle |
............ ............ ....15 Petals 5, fused (distinct in Erodium in GERANIACEAE); stamens 5; inflorescence axillary or terminal, cymose, consisting of subcapitate, umbel-like, or helicoid cymes; fruit either a capsule, or a capsular schizocarp of 5 mericarps ( Erodium in GERANIACEAE). |
............ ............ ......16 Flowers slightly bilaterally symmetrical (2 of the petals of different size than the other 3); fruit a capsular schizocarp of 5 mericarps; carpels 5 |
............ ............ ......16 Flowers radially symmetrical; fruit either a loculicidal capsule or a berry; carpels 2 or 3. |
............ ............ ........17 Fruit a berry; fresh plant rankly fragrant |
............ ............ ........17 Fruit a capsule; fresh plant not aromatic. |
............ ............ ..........18 Capsule 2-valvate; carpels 2; leaflets prominently serrate or with some tooth-like sublobes |
............ ............ ..........18 Capsule 3-valvate; carpels 3; leaflets with entire margins |
............ ........11 Leaves more complexly compound (with 2 or more orders of branching, some leaflets at least attached to second-order branches from the rachis). |
............ ............ ............ 19 Leaves 2× even-pinnate; flowers in spikes or spherical heads; fruit a legume or loment. |
............ ............ ............ 19 Leaves either 2× odd-pinnate or more complexly 2-4× ternately or ternately-pinnately compound; flowers in various inflorescences; fruits various (not legumes or loments). |
............ ............ ............ ..20 Leaf segments linear, less than 2 mm wide. |
............ ............ ............ ....21 Inflorescence an umbel; ovary inferior, of 2 fused carpels; fruit a schizocarp of 2 mericarps |
............ ............ ............ ....21 Inflorescence either a terminal solitary flower or terminal raceme or panicle; ovary superior, either of 2 fused carpels or of 1-5or many distinct 1-carpellate pistils; fruit either a capsule or an aggregate of follicles or achenes. |
............ ............ ............ ......22 Ovary of 2 fused carpels; fruit a capsule (1-seeded and indehiscent in Fumaria) |
............ ............ ............ ......22 Ovary of 1-5or many distinct 1-carpellate pistils; fruit an aggregate of follicles or achenes |
............ ............ ............ ..20 Leaf segments broader, lanceolate, ovate, or elliptic, > 5 mm wide. |
............ ............ ............ ........23 Herbaceous vine climbing by axillary tendrils; stamens 8 |
............ ............ ............ ........23 Erect or sprawling herb; stamens 5-6 or >15. |
............ ............ ............ ..........24 Leaflets sharply serrate, with usually many teeth on each leaflet side, the total number of “points” per leaflet > 10. |
............ ............ ............ ............ 25 Inflorescence an umbel; ovary inferior, of 2 fused carpels; fruit a schizocarp of 2 mericarps; inflorescence an umbel |
............ ............ ............ ............ 25 Inflorescence a panicle or raceme; ovary superior, of 1-8 carpels; fruit an aggregate of follicles, a single follicle, or an indehiscent berry-like fruit. |
............ ............ ............ ............ ..26 Flowers bisexual; carpels 1-8; fruit an aggregate of follicles, a single follicle, or an indehiscent berry-like fruit |
............ ............ ............ ............ ..26 Flowers unisexual; carpels (in pistillate flowers) of 3 (-5) carpels; fruit an aggregate of follicles |
............ ............ ............ ..........24 Leaflets entire, or with 1-several, broad, obtuse, rounded, or broadly acute “sublobes”, especially towards the tip of the leaflet, the total number of “points” per leaflet < 10. |
............ ............ ............ ............ ....27 Inflorescence an umbel; ovary inferior, of 2 fused carpels; fruit a schizocarp of 2 mericarps |
............ ............ ............ ............ ....27 Inflorescence a raceme, panicle, or cyme; ovary superior, of either 1-2 fused carpels or of many separate 1-carpellate pistils. |
............ ............ ............ ............ ......28 Perianth bilaterally symmetrical, the corolla of 4 fused petals; plant a scandent vine or erect or sprawling herb |
............ ............ ............ ............ ......28 Perianth radially symmetrical, of 1-5 whorls of separate perianth parts; plant an erect herb. |
............ ............ ............ ............ ........29 Perianth of 4-5 whorls of 3 parts each (some of the whorls modified into nectaries); pistil 1, 1-carpellate; fruit a drupelike, blue, naked seed; largest leaflets > 6 cm long, obviously longer than wide |
............ ............ ............ ............ ........29 Perianth of 1 whorl; of 4-5 parts; pistils many, each 1-carpellate; fruit an aggregate of achenes or an aggregate of follicles; largest leaflets either < 6 cm long, or if longer than 6 cm, also about as wide as long |
1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela |
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.). |
..2 Plant a vine, climbing by tendrils or twining. |
....3 Vine climbing by twining. |
......4 Leaf margins entire; flowers bisexual; plants hermaphroditic; petals connate, large and showy |
......4 Leaf margins serrate; flowers unisexual; plants dioecious; petals absent |
....3 Vine climbing by tendrils. |
........5 Ovary inferior; petals connate; flowers unisexual |
........5 Ovary superior; petals distinct; flowers bisexual |
..2 Plant an herb, sometimes sprawling, reclining (e.g. Cymbalaria in PLANTAGINACEAE, Aconitum in RANUNCULACEAE), but lacking climbing adaptations such as tendrils or twining stems. |
..........6 Ovary inferior; inflorescence an umbel; fruit a schizocarp of 2 mericarps |
............ 7 Involucre well-developed and obvious |
............ 7 Involucre absent or minute |
..........6 Ovary superior; inflorescence various, not an umbel; fruit various, a capsule, an aggregate of achenes or follicles, a ring of (>2) mericarps. |
............ ..8 Perianth uniseriate, the corolla absent (the calyx petaloid and white in Cnidoscolus); flowers unisexual; plants either with stinging hairs or not |
............ ..8 Perianth biseriate (uniseriate in Aphanes in ROSACEAE and in Trautvetteria in RANUNCULACEAE); flowers bisexual; plants lacking stinging hairs. |
............ ....9 Pistils many (or 2-3 in Aphanes in ROSACEAE), each with 1 carpel, arranged spirally or in a ring (if in a ring, of 2-5); fruit an aggregate of achenes, follicles, or utricles. |
............ ......10 Perianth bilaterally symmetrical, either hooded or spurred; fruit an aggregate of follicles |
............ ......10 Perianth radially symmetrical, not hooded or spurred; fruit an aggregate of utricles or achenes (plumose achenes in Geum) |
............ ........11 Stamens showy, bright white, dilated towards the tip; pistils ca. 15; fruit an aggregate of utricles |
............ ........11 Stamens not showy, white, or dilated towards the tip; pistils many (> 25); fruit an aggregate of achenes. |
............ ..........12 Flowers lacking a hypanthium; achenes short-beaked |
............ ..........12 Flowers with a prominent hypanthium; achenes with an elongate, plumose beak |
............ ....9 Pistil 1, with 1-to many carpels (in many MALVACEAE, the carpels loosely united in a ring of more than 5 around the style); fruit a capsule, an achene, a follicle, or a ring of 3 or 5-many 1-seeded mericarps. |
............ ............ 13 Perianth uniseriate, the corolla absent |
............ ............ 13 Perianth biseriate, with well-developed and differentiated calyx and corolla |
............ ............ ..14 Corolla bilaterally symmetrical, the petals connate (except distinct in Delphinium in RANUNCULACEAE); fruit a capsule, a follicle, or a schizocarp of 3 1-seeded mericarps. |
............ ............ ....15 Corolla not spurred; fruit an elongate (10-20 cm) capsule with 2 curved beaks |
............ ............ ....15 Corolla with a nectar spur; fruit < 3 cm long. |
............ ............ ......16 Petals distinct; fruit a follicle |
............ ............ ......16 Petals connate; fruit a capsule or a schizocarp of 3 1-seeded mericarps. |
............ ............ ........17 Carpels 2; fruit a capsule; stamens 4 |
............ ............ ........17 Carpels 3; fruit a schizocarp of 3 1-seeded mericarps; stamens 8 |
............ ............ ..14 Corolla radially symmetrical, the petals distinct (fused and tubular in Ipomoea); fruit a capsule or a schizocarp consisting of a ring of 5-many 1-seeded mericarps. |
............ ............ ..........18 Stem trailing; petals fused and tubular |
............ ............ ..........18 Stem erect; petals separate. |
............ ............ ............ 19 Stamens many, connate into a stamen tube; carpels 5-many, completely or only loosely fused; fruit a capsule or a schizocarp of 5-many mericarps borne in a ring; calyx often subtended by an epicalyx (an additional calyx-like, green, foliaceous whorl of bracts) |
............ ............ ............ 19 Stamens 5 or 10, distinct; carpels 2 or 5, fused; fruit a capsule or a schizocarp of 5 1-seeded mericarps. |
............ ............ ............ ..20 Fruit a schizocarp of 5 1-seeded mericarps; carpels 5; stamens 10 |
............ ............ ............ ..20 Fruit a capsule with 2 locules, loculicidal; carpels 2; stamens 5 |