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Key to Asteraceae, Key A: woody composites (shrubs and lianas)

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1 Leaves opposite, at least on the lower stem nodes (the leaves higher on the stem sometimes alternate); [tribe Heliantheae or tribe Eupatorieae].
..2 Heads radiate, the rays yellow to orange; disc flowers yellow to orange.
....3 Leaves linear; receptacle epaleate, but densely setose; [extreme s. TX]
....3 Leaves lanceolate or broader, 1-5× as long as wide; receptacle paleate; [collectively widespread].
......4 Leaves ovate, rhombic, or pentagonal, unlobed or 3-5-(-7) lobed; leaf blade margins toothed; leaves in part alternate upwards; [of disturbed, suburban areas of FL peninsula]; [tribe Heliantheae; subtribe Helianthinae]
........5 Ray laminae 5-16 mm long; leaf blades 7-33 cm long, the larger deeply 3-7-lobed; petiole bases dilated at base, fused to form a disc of leaf tissue at the stem
........5 Ray laminae 45-70 mm long; leaf blades 2-8 cm long, unlobed or 3-lobed; petiole bases not dilated at base
......4 Leaves lanceolate to oblanceolate, unlobed or obscurely 3-lobed; leaf blade margins toothed or entire; leaves strictly opposite; [tribe Heliantheae; subtribe Ecliptinae].
..........6 Leaves obovate to oblanceolate, unlobed, entire; heads (5-) 8-13 (-18+) mm in diameter; cypselas 3-4-smgled, not winged; [of tidally inundated salt and brackish marshes of the outer Coastal Plain, MD to s. FL, west to s. TX]
..........6 Leaves lanceolate, sometimes 3-lobed, usually serrate but sometimes nearly entire; heads 4-8 mm in diameter; cypselas flattened and winged; [TX only]
..2 Heads discoid or disciform (ray flowers lacking); disc flowers pink, purple, whitish, yellow, or orange.
............ 7 Disc flowers yellow to orange
............ 7 Disc flowers pink, purple, or white.
............ ..8 Heads small (involucres 2-8 mm high), solitary, axillary in the axils of leaves or leafy bracts (similar to the leaves but smaller) or interpretable as arrayed in bracteate racemes; heads nodding, the involucre 2-7 mm high; [collectively widespread in coastal portions of our area, of maritime situations]; [tribe Heliantheae; subtribe Ambrosiinae]
 Iva
............ ..8 Heads small to larger (involucres 3-15+ mm high), many, terminal on the branches of corymbiform or paniculiform arrays; heads erect, the involucre 4-12 mm high; [of Coastal Plain of FL, s. GA, and TX].
............ ....9 Pappus of 4-10 scales, 0.3-7 mm long; [tribe Heliantheae; subtribe Chaenactidinae]
............ ....9 Pappus of 5-70 barbellate bristles, 3-12 mm long; [tribe Eupatoriae]
............ ......10 Leaf blades spatulate to obovate, cuneate to attenuate at the base, rounded or shallowly notched at the apex; leaf margins entire; leaves opposite below, alternate higher on the stems; [native of FL]
............ ......10 Leaf blades triangular or lanceolate, truncate, cordate, or cuneate at the base, acute to acuminate at the apex; leaf margins serrate; leaves opposite throughout.
............ ........11 Phyllaries broad, 2-3× as long as wide, the inner phyllaries translucent-scarious white to pinkish at their broadly rounded apices, falling as the head goes to fruit
............ ........11 Phyllaries narrow, 4-7× as long as wide, the inner phyllaries green at their acute to acuminate apices, persistent on the head in age.
............ ..........12 Involucres 2-3 mm in diameter; phyllaries 7-16 in 1-2 series; disc flowers 3-13 per head; [s. FL only (in our region)]
............ ..........12 Involucres 3-7 mm in diameter; phyllaries 25-35 in 2-3 series; disc flowers 10-60 per head; [TX only (in our region)].
............ ............ 13 Disc flowers white (to light pink); [c. TX]
............ ............ 13 Disc flowers blue to lavender (rarely white); [extreme s. TX]
1 Leaves strictly alternate.
............ ............ ..14 Heads radiate (with ray and disc flowers) or pseudoradiate (with pseudoradiate flowers only.
............ ............ ....15 Rays white, pink, or rose.
............ ............ ......16 Rays pink or rose, 30-50 (-70); sprawling viny shrub to 40 dm tall; leaves ovate, averaging ca. 3× as long as wide, with clasping base, with a midvein and lateral veins and tertiary reticulation
............ ............ ......16 Rays white, 5-34+; shrubs 2-30 dm tall; leaves either obovate to oblanceolate, averaging ca. 4× as long as wide, dentate, or linear, ca. 10× as long as wide, entire, cuneate.
............ ............ ........17 Rays white, 5-11 (-15); leaves linear, ca. 10× as long as wide, entire; [native, s. and se. TX]
............ ............ ........17 Rays white, 21-34+; leaves obovate to oblanceolate, averaging ca. 4× as long as wide, dentate; [non-native, known in our region only from NY]
............ ............ ....15 Rays or pseudorays yellow, orange-yellow, or light creamy yellow.
............ ............ ..........18 Heads pseudoradiate, with 11-25 pseudoray flowers and 0 disc flowers; leaves 8-70 mm wide
............ ............ ..........18 Heads radiate, with (0-) 1-11 (-15) ray flowers and 1-15 (-20) disc flowers; leaves (or their segments) 0.5-10 mm wide.
............ ............ ............ 19 Phyllaries in 1-2 series and equal or subequal in length to one another, the phyllaries often coherent/fused (the involucre thus appearing as a cylinder with ribs extending from base to top; calyculus (of bracts subtending the phyllaries and differing from them in texture, color, or orientation) present; [tribe Senecioneae]
............ ............ ............ 19 Phyllaries in 3-17 series and unequal in length (imbricated); calyculus absent.
............ ............ ............ ..20 Rays light creamy yellow, 5-11 (-15); involucre 3-5 (-7) mm in diameter
............ ............ ............ ..20 Rays bright yellow, (0-) 1-9; involucre 1-2.5 (-3) mm in diameter.
............ ............ ............ ....21 Pappus of 40-60 barbellate bristles; ray blades 4-6 mm long; leaf surfaces minutely pebbled; intricately branched shrubs to 1 m tall, with persistent sterile shoots with evergreen leaves and annual flowering shoots (the entire shoot and leaves deciduous); [Coastal Plain, se. NC south to Panhandle FL and s. MS]
............ ............ ............ ....21 Pappus absent, a low coroniform structure, or of scales; ray blades 2-3.5 mm long; leaf surfaces planar/revolute; shrubs 1-15 dm tall, not differentiated into evergreen, sterile shoots bearing deciduous, fertile shoots; [OK and TX westwards, very rarely as a waif eastwards].
............ ............ ............ ......22 Pappus absent or a low (< 0.1 mm high) crown
............ ............ ..14 Heads discoid (with disc flowers only).
............ ............ ............ ........23 Pappus absent; leaves 1-2× pinnately or 1× palmately lobed.
............ ............ ............ ..........24 Leaves either palmately 3-lobed, or pinnately lobed and the few (<11) lobes > 2 mm long
............ ............ ............ ..........24 Leaves 1-pinnately lobed, the many (>21) lobes mostly < 1.5 mm long and wide
............ ............ ............ ........23 Pappus of bristles; leaves unlobed (the margins entire or serrate).
............ ............ ............ ............ 25 Plants dioecious, either male or female; heads either staminate, with 10-50 flowers, or pistillate, with 20-150 flowers
............ ............ ............ ............ 25 Plants hermaphroditic; heads bisexual, with 5-25 flowers, either all perfect (bisexual), or a mixture of perfect (bisexual) and functionally staminate flowers.
............ ............ ............ ............ ..26 Leaves strongly basally disposed, crowded in the lowermost, woody or semiwoody portion of the stem
............ ............ ............ ............ ....27 Leaves lanceolate to ovate (broadest below the midpoint); leaf margins entire to coarsely toothed.
............ ............ ............ ............ ......28 Heads in leafy, paniculate arrays; leaf blades coarsely dentate, 0.5-4 cm long; shrubs to 12 dm tall; [c. and e. TX]
............ ............ ............ ............ ......28 Heads borne in leafless, corymbiform arrays; leaf blades entire to denticulate, 5-16 (-20) cm long; shrubs to 40 dm tall; [FL and s. TX]
............ ............ ............ ............ ....27 Leaves obovate, spatulate, or orbiculate (with rounded apex) or elliptic (with acute to acuminate apex), broadest at or beyond the midpoint; leaf margins entire.
............ ............ ............ ............ ........29 Leaf blades obovate, spatulate, or orbiculate, rounded or slightly notched at the apex; leaf blade surfaces gray-green on both surfaces, granular when young; disc flowers 5 per head; [FL]
............ ............ ............ ............ ........29 Leaf blades elliptic, acute to acuminate at the apex; leaf blade strongly bicolored above and below, the upper surface dark green and glabrate, the lower surface densely and finely tomentose and strongly whitened, the upper surface dark green and glabrate or glabrous; disc flowers 4-25 per head; [s. TX]

Key to Asteraceae, Key B: herbaceous composites with opposite or whorled leaves
and discoid or disciform heads (lacking ray florets)

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1 Pappus present, of 5-60 barbellate bristles; receptacle naked (without paleae or well-developed bristles); [tribe Heliantheae; subtribe Eupatoriae].
..2 Plant a twining herb, phyllaries and disk florets 4 per head
..2 Plant stiffly erect to sprawling but never twining, phyllaries and disk florets usually > 4 per head.
....3 Leaves in whorls of 3-7, > 2 cm wide
....3 Leaves opposite, rarely alternate or whorled, if whorled, < 2 cm wide.
......4 Cypselae rostrate (with long beaks); the achene portion not bearing ribs
......4 Cypselae not beaked, the achene variously ribbed.
........5 Achenes (and ovaries) 8-10-ribbed; outer phyllaries longitudinally striate
........5 Achenes (and ovaries) (3-) 4-5-ribbed; outer phyllaries not noticeably longitudinally striate.
..........6 Heads pink to blue.
............ 7 Phyllaries in 4-6+ series, deciduous
............ 7 Phyllaries in 2-4 series, persistent (or deciduous in Praxelis).
............ ..8 Heads pink (rarely bluish); receptacles flat; florets 18-25 per head; leaves sometimes gland-dotted
............ ..8 Heads blue; receptacles conic; florets 25-70 per head; leaves gland-dotted.
............ ....9 Bristles ca. 30; florets 35-70+; phyllaries ca. 25, subequal, persistent
............ ....9 Bristles ca. 40; florets 25-30; phyllaries 15-25, unequal, deciduous (absence apparent in fruit)
..........6 Heads white to cream or, rarely, pale lilac.
............ ......10 Florets 3-7 per head
............ ......10 Florets at least 9 per head.
............ ........11 Phyllaries not strongly imbricate, with the principal ones subequal and sub-biseriate; petioles 0.5-10 cm
............ ........11 Phyllaries clearly imbricate, in 3+ series, the margins usually glandular; some species epetiolate
1 Pappus either absent, or of scales, setae, or awns; receptacle naked, paleate, or bearing bristles.
............ ..........12 Leaves whorled, linear, < 2 mm wide; head solitary; [aquatic herb growing in shallow stagnant water]
............ ..........12 Leaves opposite (or alternate in part), broader in shape and > 5 mm wide; heads typically not solitary; [terrestrial or wetland plants].
............ ............ 13 Receptacle naked (epaleate).
............ ............ ..14 Pappus of 2-12 scales; heads white, pink, or blue (yellow in Schkuhria).
............ ............ ....15 Heads with 20-125 florets; leaves serrate or ± dentate (at least distally); [rarely naturalized exotics or s. TX/FL natives]; [tribe Eupatorieae].
............ ............ ......16 Leaves petiolate; corolla tube glabrous, typically blue or lavender colored (sometimes white tinged); heads with 20-125 florets; [naturalized exotics or s. FL native]
............ ............ ......16 Leaves sessile; corolla tube glandular-pubescent, typically white colored (sometimes pink-purple tinged); heads with 75-125 florets; [s. TX native]
............ ............ ....15 Heads with 10-30 florets (or fewer in Schkuhria); leaves entire; [natives, of FL, s. GA, MS, and TX in our area]; [tribe Heliantheae; subtribe Chaenactidinae].
............ ............ ........17 Leaves 3-5 lobed (or foliolate); cypselae with curled hairs (Florestina tripteris) or villous on angles (Schkuhria pinnata); [s. TX in our area or ne. waif].
............ ............ ..........18 Heads discoid, corolla white; cypselae with curled hairs; [s. TX in our area]
............ ............ ..........18 Heads discoid, sometimes with 1 (-2) diminuitive rays, corolla (of discs) yellow; cypselae villous, at least at bases or on angles; [waif of ne US, native in sw. US]
............ ............ ........17 Leaves simple or unlobed; cypselae with straight hairs; [variously distributed in s. GA and FL w. to MO and s. TX]
............ ............ ..14 Pappus none, or of a few bristles or irregular coroniform lobes; heads green to yellow.
............ ............ ............ 19 Heads bright yellow, closely aggregated into corymbiform arrays of flat-topped to dome-shaped glomerules
............ ............ ............ 19 Heads greenish or greenish yellow, borne in spiciform, racemiform, or paniculiform arrays.
............ ............ ............ ..20 Heads in paniculiform arrays, all or most of the heads not subtended by bracts
............ ............ ............ ..20 Heads in spiciform or racemiform arrays, each 1 (-2) heads subtended by a bract
 Iva
............ ............ 13 Receptacle with paleae or well-developed bristles.
............ ............ ............ ....21 Heads small, less than 1 cm in diameter at anthesis (the female heads enlarging in Xanthium); disc florets dull white or suffused with green or purple; florets mainly unisexual (either in the same heads and then males central and females peripheral, or in separate female and male heads); female florets 0-8 per head; [tribe Heliantheae; subtribe Ambrosiinae]
............ ............ ............ ......22 Heads unisexual; cypselas shed within an indurated bur or “nut” with hooked or straight spines developed from the phyllaries and/or paleae.
............ ............ ............ ........23 Involucre of the female heads with tubercles or straight spines developing from the phyllaries; burs 1-8 mm long
............ ............ ............ ........23 Involucre of the female heads with hooked spines developing from the phyllaries/paleae; burs 10-35 mm long
............ ............ ............ ......22 Heads bisexual, with functionally male and female flowers in the same head; cypselas shed individually, not enclosed.
............ ............ ............ ..........24 Heads in paniculiform arrays, all or most of the heads not subtended by bracts
............ ............ ............ ..........24 Heads in spiciform or racemiform arrays, each 1 (-2) heads subtended by a bract
 Iva
............ ............ ............ ....21 Heads larger, mostly > 1 cm in diameter at anthesis; disc florets conspicuously white, yellow, pale yellow, or purple; florets mainly bisexual; female florets > 12 per head (except 2-8 in Polymnia and 8-15 in Verbesina occidentalis).
............ ............ ............ ............ 25 Leaves strongly basally disposed, the basal-most leaves sitting almost flat on the ground and overlapping; disc flowers maroon-purple
............ ............ ............ ............ 25 Leaves mainly or strictly cauline (not basally disposed); disc flowers white, yellow, or reddish-brown.
............ ............ ............ ............ ..26 Disc flowers bright yellow or reddish-brown.
............ ............ ............ ............ ....27 Involucre of phyllaries subtended by a calyculus of bracts obviously different in color, texture, and shape than the phyllaries.
............ ............ ............ ............ ......28 Phyllaries usually distinct, rarely connate proximally; disk florets bright yellow
............ ............ ............ ............ ......28 Phyllaries connate (both proximally-distally connate); disk florets yellow to reddish-brown
............ ............ ............ ............ ....27 Involucre of phyllaries not subtended by a calyculus (phyllaries sometimes in 2+ series, but without obvious calyculi below).
............ ............ ............ ............ ........29 Plants erect; heads without ray florets; [wet pine savannas and bogs; FL panhandle]
............ ............ ............ ............ ........29 Plants primarily prostrate or trailing, sometimes erect; heads with or without ray florets; [habitats various, widespread natives and non-natives]
............ ............ ............ ............ ..26 Disc flowers bright white or pale yellow.
............ ............ ............ ............ ..........30 Stems not winged; plants erect or spreading.
............ ............ ............ ............ ............ 31 Cypselae 3-4 angled; pappus falling (of 2-12 awns or bristles); leaf margins irregularly crenate to serrate, the surfaces sometimes roughened but lacking glands; [e. of TX]
............ ............ ............ ............ ............ 31 Cypselae 5-ribbed; pappus absent; leaf margins entire or serrulate, the surfaces gland-dotted; [s. TX]
............ ............ ............ ............ ............ ..32 Disc flowers bright white; leaves unlobed or with 2 basal lobes
............ ............ ............ ............ ............ ..32 Disc flowers creamy or pale yellow; leaves prominently pinnately lobed, with 5-many lobes

Key to Asteraceae, Key C: herbaceous composites with opposite leaves and radiate heads

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1 Ray florets white, pink, or purple (rarely pale yellow or lavender).
..2 Ray floret (the lamina) persistent on the achene and becoming papery and bleached
..2 Ray floret (the lamina) articulate from the achene and falling, thus the mature heads not appearing papery.
....3 Pappus of a minute crown (coroniform); disc florets white or whitish.
....3 Pappus absent, of 1-many scales, or of retrorsely barbed awns (or plumose bristles in Tridax procumbens); disk florets yellow, red-brown, or pinkish.
......4 Heads with an involucre not subtended by a calyculus.
........5 Rays pink, the laminae 5-14 mm long; [OK and TX westward]
........5 Rays white or whitish-yellow; the laminae typically < 5 mm long (occasionally longer in Polymnia); [collectively widespread]
..........6 Plants larger, 50-150+ cm tall; perennials; leaves > 10 cm long, sometimes with winged petioles or clasping basal appendages; disc florets functionally staminate; [natives of higher-quality, limestone or novaculite habitats]
..........6 Plants smaller, 2-40 cm tall or long (if stems > 40 cm then plants usually procumbent); annuals or perennials; leaves usually < 10 cm long, lacking winged petioles or clasping basal appendages; disc florets bisexual and fertile; [non-natives of disturbed habitats].
............ 7 Plants with leaves simple, unlobed; pappus either absent or of fimbriate to aristate scales; plants annual, erect
............ 7 Plants usually with a mix of simple and 3-lobed leaves; pappus plumose; plants perennial, usually at least somewhat procumbent
......4 Heads with an involucre subtended by a calyculus of bracts (these often but not always reflexed); the phyllaries often appearing somewhat translucent or of a distinctly different color, shape, or texture from the leafy colored bracts below; [tribe Heliantheae; subtribe Coreopsidinae].
............ ..8 Phyllaries connate for at least ¼ their length; heads with or without ray florets; [MS westwards in our area]
............ ..8 Phyllaries distinct; heads with ray florets; [collectively widespread in our area].
............ ....9 Cypselas beaked, not strongly flattened, 7-30 mm long, with 1 groove per face; leaves highly dissected
............ ....9 Cypselas beakless, more or less strongly flattened and also often winged, 1.2-16 mm long, with 0 or 2 grooves per face; leaves simple to highly dissected.
............ ......10 Cypselas 2.5-16 mm long, usually not winged; pappus awns (if present) usually retrorsely barbed; rays white
............ ......10 Cypselas 1.2-8 mm long, usually winged; pappus awns (if present) barbless or antrorsely barbed; rays pink, purple, or white
1 Rays predominantly yellow, orange, or red (sometimes with some brown, maroon, or purple coloration as well).
............ ........11 Receptacle naked, epaleate.
............ ..........12 Leaves and phyllaries with large, scattered, embedded oil glands, making the plants strongly aromatic (the glands translucent in living plants, usually golden-brown or blackish in herbarium specimens); plants annual, decumbent and much branched from the base (except Tagetes, annual and generally erect and sparingly branched); [tribe Heliantheae; subtribe Pectidinae].
............ ............ 13 Leaves unlobed, entire or serrate.
............ ............ ..14 Leaves coarsely toothed, mostly opposite (the distal alternate)
............ ............ ..14 Leaves entire (though with marginal setae), opposite
............ ............ 13 Leaves pinnately lobed or pinnatisect (pinnately divided nearly to the midrib but the leaflets not separate), the margins of terminal segments usually serrate (sometimes entire).
............ ............ ....15 Pinnate leaf segments lanceolate, 3-10 mm wide; calyculus absent
............ ............ ....15 Pinnate leaf segments linear, 1-3 mm wide; calyculus present, of linear bracts and subtending the involucre.
............ ............ ......16 Phyllaries distinct to their bases, or nearly so; ray florets 5-8; heads borne singly or in pairs/triplets; leaf surfaces puberulent
............ ............ ......16 Phyllaries connate > ½ their length; ray florets 5-21; heads borne singly; leaf surfaces glabrous to variously pubescent/floccose
............ ..........12 Leaves and phyllaries lacking embedded oil glands, though smaller punctate glands sometimes present; perennial or annual plants, upright and little or moderately branched below the inflorescence.
............ ............ ........17 Leaf surfaces densely scabrous and canescent; [s. TX westward]
............ ............ ........17 Leaf surfaces glabrous to hirsute but never canescent; [collectively widespread]
............ ............ ..........18 Leaves mainly basal or basally disposed (a few pairs low on the stem), the blades elliptic to ovate, the surfaces hirsute and stipate-glandular
............ ............ ..........18 Leaves mainly cauline, the blades primarily linear, lanceolate oblanceolate, or occasionally oblong (if ovate, then leaves mostly cauline), the surfaces glabrous to strigose but not stipitate-glandular
............ ............ ............ 19 Pappus absent or of 1-2 antrorsely-barbed or subplumose bristles; leaf blades usually lobed; [c. TX westward]
............ ............ ............ 19 Pappus absent, of scales, or coroniform (if coroniform then with 6-8 barbellulate bristles as in Jamesianthus); leaf blades unlobed; collectively widespread, including c. TX]
............ ............ ............ ..20 Heads smaller and many (10-300+), arranged into dense, flat-topped corymbs; disc florets 1-15; ray florets 0-2, the laminae inconspicuous; phyllaries 6 (-9), in 1 series; [collectively more widespread but absent from n. AL and wc. GA]
............ ............ ............ ..20 Heads larger and fewer (< 9 per inflorescence), arranged singly or in loose corymbs; ray florets 6-14, the laminae conspicuous; phyllaries 12-18, broadly ovate, squarrose and in several imbricate series; [on calcareous substrates in n. AL and wc. GA]
............ ........11 Receptacle paleate, with paleae, bristles, or scales.
image of plant
Show caption*© Scott Ward
............ ............ ............ ....21 Disc florets functionally staminate (“sterile”, not producing cypselae), the style undivided, their ovaries much smaller than ovaries of the ray florets (which are functionally pistillate).
image of plant
Show caption*© Swarochi Tathagath, some rights reserved (CC BY), uploaded by Swarochi Tathagath
............ ............ ............ ......22 Inner phyllaries prickly with straight or uncinate prickles, and each enveloping a cypsela and swelling into a bur-like structure
............ ............ ............ ......22 Inner phyllaries unarmed, not becoming bur-like (though those of Melampodium do invest the fruit).
............ ............ ............ ........23 Plants with tack-glands or pit-glands on stems, leaves, and/or phyllaries; [waif, e. TX; native further westward]
............ ............ ............ ........23 Plants without tack-glands or pit glands.
............ ............ ............ ..........24 Outer phyllaries orbicular, connate 1/4 - 1/3 of their lengths; [exotic, rare]
............ ............ ............ ..........24 Outer phyllaries orbicular to lance-linear, not connate; [native, collectively widespread in our area].
image of plant
Show caption*© Gary P. Fleming
............ ............ ............ ............ 25 Phyllaries in 2 series; pappus coroniform (Chrysogonum) or absent (Lindheimera)
............ ............ ............ ............ ..26 Plants perennial, often trailing; pappus persistent, forming a minute, half-cup-shaped crown; [e. LA eastward, e. of MS river]
............ ............ ............ ............ ..26 Plants annual, erect; pappus absent (cypselae with 2 pappus-like processes originating from cypsela wings); [w. LA westward, w. of MS river]
image of plant
Show caption*© Alan M. Cressler
............ ............ ............ ............ 25 Taller, robust plants, the stems usually 5-40 cm long at maturity, erect; pappus absent or of 2 awns.
image of plant
Show caption*© Scott Ward
............ ............ ............ ............ ....27 Cypselae strongly flattened, borne in 2-3 series from the 2-3 series of ray florets
image of plant
Show caption*© Scott Ward
............ ............ ............ ............ ....27 Cypselae thick, not flattened, borne in 1 series from the 1 series of ray florets
............ ............ ............ ....21 Disc florets functionally bisexual (“fertile”, producing cypselae), the style divided, their ovaries as large as, or larger than, ovaries of the ray florets (which may be either functionally pistillate or completely neuter).
............ ............ ............ ............ ......28 Ray corolla (the lamina) persistent on the achene and becoming papery and bleached.
............ ............ ............ ............ ........29 Outer phyllaries with a stiff, maroon-colored subulate apex
............ ............ ............ ............ ........29 Outer phyllaries lacking a stiff subulate apex.
............ ............ ............ ............ ..........30 Leaf margins serrate or toothed; plant a perennial; cypselae subterete, 4-5 mm long; [natives]
............ ............ ............ ............ ..........30 Leaf margins entire; plant an annual; cypselae 3-angled or flattened, 6-10 mm long; [non-native waifs]
............ ............ ............ ............ ......28 Ray corolla articulate from the achene and falling after flowering.
............ ............ ............ ............ ............ 31 Pappus absent or of scales or awns (if with awnlike bristles as in Acmella, then these never plumose).
............ ............ ............ ............ ............ ..32 Plants with tack-glands or pit-glands on stems, leaves, and/or phyllaries; [waif, VA and NC northward; subtribe Madiinae]
............ ............ ............ ............ ............ ..32 Plants without tack-glands or pit-glands on stems, leaves, and/or phyllaries; [natives and non-natives, collectively widespread]
............ ............ ............ ............ ............ ....33 Paleae not notably clasping the cypsela; cypselae notably flattened (or weakly compressed/angled in Guizotia); heads small, the receptacle 3-8 mm in diameter (ca. 10-15 mm in Guizotia).
............ ............ ............ ............ ............ ......34 Pappus persistent, of 2-3(-5) scales or awns; ray laminae shorter, 2-5 mm long
............ ............ ............ ............ ............ ........35 Cypselae dimorphic, those of the ray florets with lacerate wings
............ ............ ............ ............ ............ ....33 Paleae either entirely enveloping and falling with each cypsela or conduplicate (V-shaped in cross section), the 2 sides of the V partially clasping the cypsela; cypselae flattened, terete, or angled; heads mostly larger.
............ ............ ............ ............ ............ ..........36 Phyllaries apparently 4, the outer 4 foliaceous and forming a fused quadrangle which conceals the much smaller and narrower inner phyllaries (each inner phyllary subtending a ray floret); cypselae finely 32-40 ribbed
............ ............ ............ ............ ............ ..........36 Phyllaries not as above, instead 5 or more and not forming a conspicuously fused quandrangle; cypselae angled or smooth (sometimes angled, but lacking many fine ribs)
............ ............ ............ ............ ............ ............ 37 Paleae completely surrounding the cypselae of the disc florets, forming a perigynium or sac-like appendage
............ ............ ............ ............ ............ ............ 37 Paleae conduplicate or partially enclosing the cypselae, but not forming a sac-like appendage.
............ ............ ............ ............ ............ ............ ..38 Disc florets with hairy staminal filaments; pappus of 4-6 aristate to lacerate scales; [TX westward]
............ ............ ............ ............ ............ ............ ..38 Disc florets without hairy staminal filaments; pappus absent or of 2-3 scales or awns (sometimes accompanied by up to 8-12 additional shorter scales in Helianthus and Simsia, but these readily falling); [collectively widespread].
............ ............ ............ ............ ............ ............ ....39 Rays primarily yellow; [natives and non-natives, collectively widespread]
............ ............ ............ ............ ............ ............ ......40 Cypselae (of at least the disc florets) strongly flattened and generally also winged (thin-margined in Simsia).
............ ............ ............ ............ ............ ............ ........41 Cypselae 1-2.5 mm (Acmella) or 3.5-6 mm long (Simsia); plants erect, decumbent, or prostrate; stems never winged.
............ ............ ............ ............ ............ ............ ..........42 Phyllaries 8-15; cypselae 1-2.5 mm long; involucre ca. 3-6 mm in diameter
............ ............ ............ ............ ............ ............ ..........42 Phyllaries 13-43; cypselae 3.5-6 mm long; involucre 5-16+ mm in diameter
............ ............ ............ ............ ............ ............ ......40 Cypselae subterete, quadrangular, variously angled, or diamond-shaped in ×-section, not winged.
............ ............ ............ ............ ............ ............ ............ ..44 Creeping herb; leaves serrate and also mostly 3-lobed; peduncles 3-10 cm long
............ ............ ............ ............ ............ ............ ............ ....45 {XXXX}; Leaf surfaces and ray laminae abaxially gland-dotted; [SC waif; native to sw. US]
............ ............ ............ ............ ............ ............ ............ ......46 Leaves linear, lanceolate, or ovate, almost always some leaves on a plant > 7 mm wide; plants from crowns, some species with thickened vertical storage roots (only H. tuberosus producing horizontal tubers); [collectively widespread in our area]
............ ............ ............ ............ ............ ............ ............ ......46 Leaves linear, 0.5-7 mm wide; plants from slender horizontal tubers; [c. FL north to s. AL]
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