Flora of the Southeastern United States
2022 Edition

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Click the number at the start of a key lead to highlight both that lead and its corresponding lead. Click again to show only the two highlighted leads. Click a third time to return to the full key with the selected leads still highlighted.

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Keyed in multiple places:

Key to Santalaceae

1 Leaves alternate; monoecious herb or shrub.
..2 Herb, < 2 (-3) dm tall; leaves 1-4 cm long, glabrous; inflorescence a terminal panicle of cymes; [tribe Comandreae or family COMANDRACEAE]
..2 Shrub, > 4 dm tall; leaves 5-15 cm long, pubescent; inflorescence a terminal raceme; [tribe Pyrularieae or family CERVANTESIACEAE]
1 Leaves opposite; dioecious shrubs.
....3 Aerial (epiphytic) shrubs, parasitic on tree trunks and branches; leaves either coriaceous and brittle when live, or minute and scale-like; [tribe Visceae or family VISCACEAE]
......4 Leaves minute and scale-like; stems to 3 cm long; [south to nw. NJ and ne. PA]
......4 Leaves flat, 2-6 cm long, 1-5 cm wide; stems to 50 cm long; [widespread in our area]
....3 Terrestrial shrubs, parasitic via root connections; leaves herbaceous, flexible when live.
........5 Staminate flowers in terminal umbel-like dichasia; pistillate flowers (and fruits) solitary, terminal; clumped shrub to 4 m tall; [tribe Thesiae or family THESIACEAE]
........5 Staminate flowers in axillary umbels; pistillate flowers (and fruits) solitary, axillary; rhizomatous shrub to 1 m tall; [tribe Santaleae or family SANTALACEAE]

Key J4: shrubs and subshrubs with opposite simple leaves with entire margins

1 Aerial and epiphytic, hemiparasitic shrub
1 Terrestrial, autotrophic or hemiparasitic shrub or subshrub.
..2 Leaves succulent, nearly as thick as wide; [brackish to saline situations]
..2 Leaves herbaceous (succulent in Borrichia), much wider than thick; [various habitats].
....3 Creeping or short subshrubs, the stems primarily prostrate, < 2 dm tall.
......4 Well-developed leaves 4-6 per stem; inflorescence a head subtended by 4 large white bracts
......4 Well-developed leaves many per stem; inflorescence of individual flowers axillary in pairs or clusters or in terminal cymes.
........5 Flowers yellow; leaves with pellucid or dark punctate glands (use at least 10× magnification)
........5 Flowers white, pale pink, or deep pink; leaves lacking sessile, punctate glands.
..........6 Leaves linear; flowers pale to deep pink, 5-merous
..........6 Leaves orbicular or elliptic; flowers white to pale pink, 4-merous or 5-merous
............ 7 Leaves elliptic; flowers 5-merous; fruit a brownish capsule
............ 7 Leaves orbicular; flowers 4-merous; fruit a red berry
....3 Upright or scrambling shrubs, > 3 dm tall
............ ..8 Scrambling shrubs, armed with recurved paired spines at the nodes
............ ..8 Upright shrubs, unarmed.
............ ....9 Inflorescence a terminal head of many flowers.
............ ......10 Head spherical, lacking an involucre of conspicuous bracts or phyllaries
............ ......10 Head flattened, either subtended by 4 large white bracts or by an involucre with >5 green phyllaries.
............ ........11 Head subtended by 4 large white bracts; leaves with prominently parallel-arcing secondary veins; flowers 4-merous
............ ........11 Head subtended by an involucre of >5 green phyllaries; leaves with venation otherwise; flowers 5-merous
............ ....9 Inflorescence otherwise, either of a solitary flower, or one of a wide variety of inflorescences with flowers attached at different points along branched or unbranched axes. {add: [Lagerstroemia] LYTHRACEAE; [Rosmarinus] LAMIACEAE; [Laguncularia] COMBRETACEAE; [Buxus] BUXACEAE; [Exochorda] ROSACEAE; various other [see spreadsheet]}
............ ..........12 Inflorescence flat-topped (broader than long), terminal, a compound cyme or corymb.
............ ............ 13 Flowers bright yellow; stamens many; leaves < 1.5 cm wide; fruit a capsule; leaves with pellucid or dark punctate glands (use at least 10× magnification)
............ ............ 13 Flowers white or creamy; stamens 4-5; leaves > 1.5 cm wide; fruit a drupe; leaves lacking sessile, punctate glands.
............ ............ ..14 Petals 5; foliage with simple hairs
............ ............ ..14 Petals 4; foliage with T-shaped hairs
............ ..........12 Inflorescence either terminal and not flat-topped, or axillary and variously shaped, or terminal and solitary, or leaf-opposed.
............ ............ ....15 Carpels many (> 9), either separate or fused; stamens many; perianth segments either many and undifferentiated into calyx and corolla, maroon, brown, or yellow (in CALYCANTHACEAE), or differentiated into a fleshy and persistent calyx of 5-9 sepals, and a deciduous corolla of 5-9 red (or white) petals (Punica in LYTHRACEAE).
............ ............ ......16 Fruit a wrinkled, 3-7 cm long, brown to black, elliptical aggregate of nearly spherical, large achenes; flowers solitary in axils; perianth segments many and undifferentiated into calyx and corolla, maroon, brown, or yellowish; ovary superior; branches unarmed
............ ............ ......16 Fruit a leathery, 4-15 cm in diameter, reddish, spherical berry with obpyramidal seeds surrounded by a juicy sarcotesta (pomegranate); perianth differentiated, the sepals fleshy and persistent on the fruit, the petals deciduous, 5-9, bright red to white; ovary inferior; branches typically armed with axillary spines
............ ............ ....15 Carpels 1-5, fused; stamens either 1-5 or 8-10; perianth segments 4-5 or 8, variously colored; fruit a simple capsule, drupe, or berry; flowers 2-many, in axillary or terminal inflorescences (pistillate flowers sometimes solitary in SANTALACEAE and THESIACEAE); [Eudicots].
............ ............ ........17 Ovary inferior; corolla absent, radially symmetrical, or bilaterally symmetrical; fruit either a berry or a 1-seeded drupe.
............ ............ ..........18 Flowers unisexual and plants dioecious; corolla absent; pistillate flowers solitary, staminate flowers in pedunculate umbels or cymes, either terminal or axillary; fruit a 1-seeded drupe; leaves acute to acuminate at the apex
............ ............ ............ 19 Inflorescences terminal
............ ............ ............ 19 Inflorescences axillary
............ ............ ..........18 Flowers bisexual and plants hermaphroditic; corolla present; flowers paired, terminal or axillary, or in axillary spikes; fruit a berry; leaves rounded, obtuse, to acute (or acuminate in Lonicera maackii) at the apex
............ ............ ........17 Ovary superior; corolla radially symmetrical (absent in Forestiera in OLEACEAE); fruit either a 1-4-seeded drupe, or a many-seeded berry, or a capsule.
............ ............ ............ ..20 Stamens 8-10, of 2 different lengths in each flower; petals separate, 4-5 (-7), pink purple, 10-15 mm long; stems strongly arching, rooting at the tips; [plants of flooded to saturated wetlands]
............ ............ ............ ..20 Stamens either (1-) 2 (-4), or 4-5, or 10, all of the same length; petals fused (separate in RHAMNACEAE and BUXACEAE, but then < 5 mm long and white or cream), white, bright-yellow, lilac, or pink; stems erect (or at least not arching and rooting at the tips); [plants of various habitats].
............ ............ ............ ....21 Petals separate, 4-5, white or cream; stamens 4-5.
............ ............ ............ ......22 Fruit a loculicidal capsule, dehiscing into 3 valves; branches square in ×-section; leaves < 2 cm long; [alien, cultivated and weakly established, of temperate areas]
............ ............ ............ ......22 Fruit a drupe with 2-4 pyrenes; branches round or nearly so in ×-section; leaves > 2 cm long; [natives, of peninsular FL]
............ ............ ............ ....21 Petals fused, 4-5, white, bright yellow, lilac, or pink; stamens either (1-) 2 (-4) or 10; fruit either a capsule or a 1-seeded drupe.
............ ............ ............ ........23 Perianth 5-merous; corolla pink; stamens 10; fruit a 5-locular capsule
............ ............ ............ ........23 Perianth 4-merous; corolla white, bright yellow, or lilac; stamens (1-) 2 (-4); fruit either a 1-seeded drupe or a 2-locular capsule

Key L: epiphytic angiosperms {Note that epiphytic Pteridophytes are not re-keyed here; seek them in Keys A4 and A6}

1 Stems yellow to bright orange, lacking leaves
1 Stems green or brown, with leaves (scale-like or larger).
..2 Leaves opposite, orbicular or oblanceolate, rounded at the apex; [Eudicots]
..2 Leaves alternate, either orbicular or oblanceolate (rounded at the apex), or scale-like, or elongate and tapering, or lanceolate-elliptic.
....3 Leaves orbicular, rounded at the apex; [Basal Angiosperms]
....3 Leaves either scale-like, or elongate and tapering, or lanceolate-elliptic; [Monocots].
......4 Leaves either scale-like or elongate and tapering; flowers radially symmetrical
......4 Leaves lanceolate-elliptic; flowers bilaterally symmetrical