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| 1 Pappus plumose, the barbels along each pappus bristle mostly 0.5-1.0 mm long or longer. |
| ..2 Inner phyllaries with apices prolonged, slightly dilated, and petaloid (white to yellow, pink, or purplish); heads 3-5 mm in diameter, with 4-6 flowers per head. |
| ....3 Heads pedunculate; [east of the Mississippi River in e. LA, s. MS, AL, FL, GA, and SC]. |
| ......4 Phyllary apices lavender to pink or magenta |
| ......4 Phyllary apices white to cream or light yellow |
| ....3 Heads sessile (plants that are top-injured or knocked over and prostrate may develop pedunculate heads); [west of the Mississippi River in AR, OK, LA, and TX]. |
| ........5 Phyllary apices loosely erect, few if any diverging > 30 degrees from the axis of the head; colored portions of the phyllaries lavender to pink or magenta (rarely mainly green with whitish margins); [widespread in w. and s. AR, e. and se. OK, w. LA, and e. TX] |
| ........5 Phyllary apices spreading to recurving, many or most spreading at more than 30 degrees from the axis of the head; colored portions of the phyllaries either white to cream or light yellow or lavender to pink or magenta; [e., se., and s. TX]. |
| ..........6 Corms depressed-globose, flat or rounded at the base; phyllary apices white to cream or light yellow (rarely lavender to pink or magenta); [e. TX, from c. Bastrop County and Travis County northeastwards] |
| ..........6 Corms elongate and tapering downwards (napiform or turnip-like); phyllary apices lavender to pink or magenta; [s. and se. TX, from s. Bastrop County, Fayette County, and Calhoun County southwestwards]. |
| ..2 Inner phyllaries not prominently petaloid; heads 10-20 mm in diameter, with 2-60 flowers per head. |
| ............ 7 Heads in corymbiform or broadly racemiform arrays, at least some of the heads on peduncles 20-70 mm long. |
| ............ ..8 Involucres 8-11 mm in diameter; inflorescences corymbiform; leaves 2-5 mm wide; [e. TX] |
| ............ ..8 involucres 15-20 mm in diameter; inflorescences broadly racemiform (at least when well-developed, with > 5 heads); leaves 1-2 (-2.5) mm wide; [c. peninsular FL scrub] |
| ............ 7 Heads in narrowly racemiform or spiciform arrays (or borne singly), the heads sessile or on short peduncles 1-10 (rarely to 25) mm long. |
| ............ ....9 Leaves with 3-5 nerves; florets 10-60 per head; corolla lobes densely hairy. |
| ............ ......10 Outer phyllaries as long as or (more usually longer than) the inner phyllaries, spreading or reflexed, the spreading portion typically > 2 mm long. |
| ............ ........11 Heads usually single, each subtended by foliaceous bracts longer than the head; ; stems glabrous; [Ouachita Mountains of AR and e. OK] |
| ............ ........11 Heads in racemiform arrays; stems glabrous or hairy; [collectively widespread]. |
| ............ ..........12 Stems glabrous; leaves glabrous; phyllaries glabrous and eciliate |
| ............ ..........12 Stems hairy; leaves sparsely pilose to glabrescent; phyllaries glabrous or pubescent on the faces, ciliate-margined |
| ............ ......10 Outer phyllaries shorter than the inner phyllaries, erect-appressed to spreading or reflexed, the spreading portion 0-2 mm long. |
| ............ ............ 13 Stems and leaves usually glabrous; inner phyllaries usually apically rounded to truncate, apiculate, all essentially erect and appressed, usually with a narrow hyaline border |
| ............ ............ 13 Stems and leaves hirsute to hirsute-pilose; inner phyllaries apically acute-acuminate, all usually spreading to reflexed on the distal 1/3 (outer) to 1/5 (inner), usually without a hyaline border |
| ............ ....9 Leaves with 1 nerve (or sometimes with 3-5 nerves in L. acidota); florets either (2-) 3-6 (-8) or 8-14 (-16) per head. |
| ............ ............ ..14 Heads borne in loose spiciform arrays, the heads spaced widely and the stem therefore not obscured; florets 8-14 (-16) per head; [se. TX] |
| ............ ............ ..14 Heads borne in dense spiciform arrays, the heads overlapping and the stem therefore generally obscured; florets (2-) 3-6 (-8) per head. |
| ............ ............ ....15 Corms elongate and becoming rhizomatous |
| ............ ............ ....15 Corms globose, depressed-globose, or ovate. |
| ............ ............ ......16 Leaves with 3 (-5) nerves; basal leaves fibrous-persistent; cauline leaves abruptly reduced in size; [prairies, Coastal Plain of w. LA and e. TX]. |
| ............ ............ ......16 Leaves with 1 nerve; basal leaves withering completely; cauline leaves nut reduced or only gradually so; [collectively more widespread from MO and KS south to w. LA and e. and c. TX]. |
| ............ ............ ........17 Stems stipitate-glandular; [TX] |
| ............ ............ ........17 Stems glabrous. |
| ............ ............ ..........18 Phyllaries 6-11, in 2-3 series, oblong-lanceolate to lanceolate-triangular, subequal (the inner 11-12 mm long), the apices acute to acute-attenuate (sometimes with narrow tips), lateral veins usually evident beyond middle, continuing nearly to tips); flowering mid Jul–Aug (-Sep) |
| ............ ............ ..........18 Phyllaries 11–18 in (4-) 5-6 series, mostly oblong-obovate, unequal (the inner 7-9 mm long), apices obtuse to truncate-rounded (each tipped by thick, often indurate apicula or mucros), lateral veins usually not evident beyond middle or 2/3s point; flowering mostly mid-late Aug (-Oct) |
| 1 Pappus barbellate, the barbels along each pappus bristle 0.1-0.3 (-0.4) mm long. |
| ............ ............ ............ 19 Heads mostly 1-30 in subcorymbiform or corymbiform arrays; [of either c. peninsular FL or Bibb Co., AL] |
| ............ ............ ............ ..20 Phyllaries in 6-7 series, the apices rounded or obtuse; [Florida scrub, c. peninsular FL] |
| ............ ............ ............ ..20 Phyllaries in 3-4 (-5) series, the apices acuminate; [dolomite glades, Bibb County, AL] |
| ............ ............ ............ 19 Heads usually > 10 in a spiciform or racemiform arrangement; [collectively widespread]. |
| ............ ............ ............ ....21 Leaves 3-5-veined. |
| ............ ............ ............ ......22 Stems glabrous or sparsely sessile-glandular; leaf bases not fibrous-persistent; corolla tubes pilose inside; [c. peninsular FL, west side of the peninsula] |
| ............ ............ ............ ......22 Stems glabrous or puberulent (not glandular); leaf bases fibrous-persistent; corolla tubes glabrous inside; [collectively widespread]. |
| ............ ............ ............ ........23 Phyllary apices obtuse to rounded. |
| ............ ............ ............ ..........24 Basal and lower cauline leaves (2-) 4-8 mm wide, cauline usually abruptly reduced in size at ca. midstem, continuing distally as linear, bract-like leaves; heads in a densely (- to loosely) spiciform arrangement; involucres 7-9 mm, purplish to greenish; florets 5-6 (-8) per head; [mainly of the Coastal Plain] |
| ............ ............ ............ ..........24 Basal and lower cauline leaves 4-10 (-20) mm wide, cauline usually gradually reduced in size distally; heads in a densely to loosely spiciform arrangement; involucres (7-) 8-11 mm, usually greenish; florets (4-) 6-8 (-12) per head; [of the Mountains and Piedmont] |
| ............ ............ ............ ........23 Phyllary apices acuminate or acute. |
| ............ ............ ............ ............ 25 Phyllary apices erect and appressed; florets (2-) 3-4 (-5) per head |
| ............ ............ ............ ............ 25 Phyllary apices reflexed to spreading, somewhat dilated and often slightly petaloid; florets (4-) 5-8 per head |
| ............ ............ ............ ............ ..26 Stems moderately to densely puberulent; leaves moderately to densely puberulent (rarely glabrescent); corms globose to elongate |
| ............ ............ ............ ............ ..26 Stems glabrous or sparsely puberulent only near the inflorescence; leaves glabrous; corms globose |
| ............ ............ ............ ....21 Leaves 1 (-3) veined. |
| ............ ............ ............ ............ ....27 Mid and inner phyllaries with a point of some kind: either apically acuminate or acute or (if rounded-retuse) minutely involute-cuspidate to apiculate. |
| ............ ............ ............ ............ ......28 Phyllaries narrowly triangular, the margins without hyaline borders; phyllary tips divergent from the involucre, usually acuminate to acute; florets 10–12 per head; [w. LA and e. TX] |
| ............ ............ ............ ............ ......28 Phyllaries variously oblong, lanceolate, or ovate, the margins usually apparently hyaline; phyllary tips erect or appressed to the involucre, acuminate, acute, cuspidate, or apiculate; florets 3-10 (-12); [east of Mississippi River]. |
| ............ ............ ............ ............ ........29 Stems hirtellous with spreading to slightly deflexed hairs or variously puberulent to hirsute. |
| ............ ............ ............ ............ ..........30 Involucres 4-6 (-7) mm high; stems hirsute to puberulent to pilose-puberulent or strigose-puberulent |
| ............ ............ ............ ............ ..........30 Involucres 8-15 mm high; stems hirtellous with spreading to slightly deflexed hairs. |
| ............ ............ ............ ............ ............ 31 Corms elongated into rhizomes |
| ............ ............ ............ ............ ............ 31 Corms globose or depressed-globose, or sometimes somewhat elongated. |
| ............ ............ ............ ............ ............ ..32 Heads sessile, relatively crowded in a cylindric arrangement. |
| ............ ............ ............ ............ ............ ....33 Heads rigidly ascending, appressed to the rachis and to each other, densely overlapping; phyllary margins usually not ciliolate; phyllary tips erect; [FL peninsula, Panhandle, and sw. GA and s. AL] |
| ............ ............ ............ ............ ............ ....33 Heads spreading to ascending, not strongly overlapping; phyllary margins ciliolate; phyllary tips slightly spreading; [FL Panhandle (Franklin and Wakulla counties] |
| ............ ............ ............ ............ ............ ..32 Heads sessile to short-pedunculate, in a relatively loose, spiciform, racemose, or paniculate, commonly secund arrangement. |
| ............ ............ ............ ............ ............ ......34 Stems glabrous (rarely sparsely hirtellous); leaves and phyllaries sparsely or not at all gland-dotted; involucres 11-15 mm high; inner phyllaries with acute apices |
| ............ ............ ............ ............ ............ ......34 Stems minutely puberulent-hirtellous; leaves and phyllaries gland-dotted; involucres 7-10 (-14) mm high; inner phyllaries with abruptly acuminate apices, often apiculate or mucronulate |
| ............ ............ ............ ............ ........29 Stems glabrous or sometimes sparsely pilose. |
| ............ ............ ............ ............ ............ ........35 Phyllaries apically usually rounded-retuse and minutely involute-cuspidate to apiculate; corolla tubes glabrous within. |
| ............ ............ ............ ............ ............ ..........36 Stems and basal leaves glabrous; basal leaves mostly arising from congested nodes at very base of plant, (1-) 2-6 (-9) mm wide, abruptly reduced in size distally, surfaces minutely white-dotted by stomates, not glandular-punctate |
| ............ ............ ............ ............ ............ ............ 37 Leaves in a diffuse, few-leaved basal rosette, 2–3 mm wide and 20–30 cm long, mostly somewhat ascending; culms (60–)75–100(–150) cm tall, 2–3 mm wide, the inflorescence averaging 25% of the culm height; flowering mostly from October to November, during the dry season in peninsular Florida |
| ............ ............ ............ ............ ............ ............ 37 Leaves in a well-defined, densely leafy basal rosette, (4–)5–7(–10) mm wide and 15–20 cm long, mostly lax and spreading, not ascending; culms (40–)50–80(–120) cm tall, 4–6 mm wide, the inflorescence averaging half the culm height; flowering mostly from July to September, during the wet season in peninsular Florida |
| ............ ............ ............ ............ ............ ..........36 Stems and basal leaves glabrous to very sparsely pilose, leaves usually with a few spreading cilia near insertion; basal and lower cauline leaves arising from numerous, separated nodes on proximal part of stem, 1-2 (-2.5) mm wide and relatively even-sized, surfaces glandular-punctate |
| ............ ............ ............ ............ ............ ........35 Phyllaries apically acute; corolla tubes pilose within. |
| ............ ............ ............ ............ ............ ............ ..38 Heads often in a secund arrangement; involucres 7-15 mm; phyllaries obovate; florets 3-6. |
| ............ ............ ............ ............ ............ ............ ..38 Heads in a secund arrangement or not; involucres (6-) 7-9 mm; phyllaries ovate-triangular to generally oblong; florets 4-10 (-12). |
| ............ ............ ............ ............ ............ ............ ....39 Heads densely arranged, on internodes 1-2 (-5) mm long, often secund; phyllary apex sharply acuminate-acute, distinctly involute, lamina relatively thin, glands consistently present and superficial at least on proximal portion; florets 4-7 (-9); basal and lower cauline leaves 2-5 mm wide, gradually reduced in length distally |
| ............ ............ ............ ............ ............ ............ ....39 Heads loosely arranged, on internodes 6-15 (-20) mm long, not secund; phyllary apex sharply acute to obtuse-angled with a thickened apiculum, not markedly involute, lamina relatively thick, usually with evidently sunken punctate glands, without superficial glands; florets 7-10 (-12); basal and lower cauline leaves 4-9 (-12) mm wide, quickly reduced in width and length distally |
| ............ ............ ............ ............ ....27 Mid and inner phyllaries lacking a point: apically rounded, obtuse, or truncate. |
| ............ ............ ............ ............ ............ ............ ......40 Stems glabrous (rarely sparsely to moderately pilose in L. pilosa). |
| ............ ............ ............ ............ ............ ............ ........41 Involucres 5-7 (-9) mm; florets 4-5 (-6); corolla tubes glabrous within; pappus bristles usually about half the length of corolla tubes |
| ............ ............ ............ ............ ............ ............ ........41 Involucres 6-10 mm; florets (6-) 7-13 (-17); corolla tubes internally pilose; pappus bristles as long as the corolla tubes (shorter in some populations of L. helleri). |
| ............ ............ ............ ............ ............ ............ ..........42 Stems 15-55 cm; leaves and phyllaries weakly or not at all punctate; pappus bristles 1/3-2/3 to equal the corolla tube length; [of the Mountains and upper Piedmont] |
| ............ ............ ............ ............ ............ ............ ............ 43 Pappus either 0.5 as long as the corolla tube or longer; plants usually < 4 dm tall; [restricted to high elevation areas near Grandfather Mountain NC (and Roan Mountain?)] |
| ............ ............ ............ ............ ............ ............ ............ 43 Pappus about as long as the corolla tube; plants usually > 4 dm tall; [in high elevation areas in w. NC, as well as in mid to lower elevation sites from nw. NC northwards through w. VA and e. WV] |
| ............ ............ ............ ............ ............ ............ ..........42 Stems 40-120 cm; leaves and phyllaries distinctly punctate-glandular to weakly punctate; pappus bristles equal the corolla tube length; [of the Coastal Plain and Piedmont]. |
| ............ ............ ............ ............ ............ ............ ............ ..44 Stems glabrous; heads loosely arranged, on internodes (2-) 5-10 (-14) mm; peduncles 0-2 (-7) mm; involucres 6-8 mm; phyllaries in 3-4 (-5) series |
| ............ ............ ............ ............ ............ ............ ............ ..44 Stems glabrous to sparsely or moderately pilose; heads densely arranged, on internodes (1-) 2-5 (-7) mm; peduncles 0-10 (-17, -80 in proximal part of capitulescence) mm; involucres (7-) 8-10 mm, phyllaries in (3-) 4-5 (-6) series |
| ............ ............ ............ ............ ............ ............ ......40 Stems puberulent to strigose. |
| ............ ............ ............ ............ ............ ............ ............ ....45 Involucres 2.5-7 mm wide; florets 3-12. |
| ............ ............ ............ ............ ............ ............ ............ ......46 Stems and peduncles puberulent to pilose-puberulent or strigose-puberulent; heads usually on ascending peduncles 2-10 (-12) mm; involucres 2.5-4 (-5) mm wide; phyllaries apically rounded or obtuse to acute or acuminate; florets 3-6 (-9). |
| ............ ............ ............ ............ ............ ............ ............ ........47 Lower stem leaves with slender petioles and blades elliptic to lance-elliptic, the blades (8-) 11-22 (-27) mm wide (bracts subtending proximal heads 2+ mm wide, grading gradually from distal cauline leaves); phyllary apices usually acuminate to acute, rarely obtuse |
| ............ ............ ............ ............ ............ ............ ............ ........47 Lower stem leaves usually lanceolate, linear, linear-oblanceolate, or oblanceolate, sometimes narrowly spatulate, 2-8 (-10) mm wide (bracts subtending proximal heads to 2 mm wide, abruptly differentiated from distal cauline leaves); phyllary apices usually rounded or obtuse-truncate, rarely acute |
| ............ ............ ............ ............ ............ ............ ............ ......46 Stems and peduncles stiffly short-strigose with closely ascending hairs; heads on divergent, arcuate-ascending peduncles 10-25 (-30) mm; involucres 5-7 mm wide; phyllaries apically rounded to nearly flat; florets 7-12 |
| ............ ............ ............ ............ ............ ............ ............ ....45 Involucres 13-22 (-25) mm wide or (6-) 8-15 mm wide (L. squarrulosa); florets 11-80. |
| ............ ............ ............ ............ ............ ............ ............ ..........48 Heads usually on peduncles usually 8-50 mm (rarely subsessile); phyllaries erect, not reflexing; florets ca. 30-80 (19-33 in L. scariosa); corolla tubes glabrous or pilose within. |
............ ............ ............ ............ ............ ............ ............ ............ 49 Phyllaries with hyaline borders that are erose to lacerate; corolla tubes glabrous inside; [perhaps not reaching our area, reported from IL and MO] |
| ............ ............ ............ ............ ............ ............ ............ ............ 49 Phyllaries with margins not or very narrowly hyaline, entire or nearly so; corolla tubes glabrous or pilose inside; [collectively widespread in our area]. |
| ............ ............ ............ ............ ............ ............ ............ ............ ..50 Leaves or leafy bracts 8-20 (-25) below the heads, cauline usually abruptly reduced above the basal; florets 19-33 per head; [of the Central and Southern Appalachians] |
| ............ ............ ............ ............ ............ ............ ............ ............ ..50 Leaves or leafy bracts 20-85 below the heads, usually continuing relatively even-sized upward above the basal; florets ca. 30-80 per head; [of WV and PA northward] |
| ............ ............ ............ ............ ............ ............ ............ ............ ....51 Lower cauline leaves mostly 25-50 (-55) mm wide; leaves glandular-dotted and also puberulent |
| ............ ............ ............ ............ ............ ............ ............ ............ ....51 Lower cauline leaves mostly 7-20 (-26) mm wide; leaves glabrous or puberulent, lacking or with few glands; [PA northwards] |
| ............ ............ ............ ............ ............ ............ ............ ..........48 Heads usually sessile, less commonly subsessile on peduncles 1-8 mm (rarely more); at least outer phyllaries usually reflexing; florets 11-26 (-30) per head; corolla tubes pilose within. |
| ............ ............ ............ ............ ............ ............ ............ ............ ......52 Phyllaries glabrous, bullate, with broad, conspicuous, often erose to lacerate or irregular, hyaline border |
| ............ ............ ............ ............ ............ ............ ............ ............ ......52 Phyllaries glabrous to puberulent or puberulent-hirtellous, essentially flat (not bullate), without hyaline border or border narrow and inconspicuous |