Flora of the Southeastern United States
2022 Edition


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Cystopteris protrusa (Weatherby) Blasdell. Lowland Bladder Fern. Phen: Apr-Jun. Hab: Rich woods or on moss- and soil-covered talus in boulderfields, occasionally on ledges of rock outcrops. Dist: NY and ON west to MN, south to GA, Panhandle FL (Washington and Gadsden counties) (Wunderlin & Hansen 2006; L. Byrd, pers.comm., 2020), AL, MS, LA, e. TX, and OK.

Origin/Endemic status: Native

Syn: = Ar, C, ETx1, Fl1, FNA2, GrPl, Il, K1, K3, K4, Mo1, NE, NY, Pa, RAB, Tn, TxFerns, Va, W, WH3, WV, Shmakov, Batkin, & Vaganov (2018); = Cystopteris fragilis var. protrusa – F, G, Sf; < Cystopteris fragilis – Tx

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image of plant© Bruce A. Sorrie
image of plant© Bruce A. Sorrie