Cystopteris protrusa (Weatherby) Blasdell. Common name: Lowland Bladder Fern. Phenology: Apr-Jun. Habitat: Rich woods or on moss- and soil-covered talus in boulderfields, loess bluffs, occasionally on ledges of rock outcrops. Distribution: NY and ON west to MN, south to GA, Panhandle FL (Washington and Gadsden counties) (Wunderlin & Hansen 2006; L. Byrd, pers.comm., 2020), AL, MS, LA, e. TX, and OK.
Origin/Endemic status: Native
Taxonomy Comments: This species is a diploid involved in the reticulate evolution of Cystopteris in e. North America. It is one parent of C. tennesseensis and of C. tenuis. Its genome can be symbolized PP.
Synonymy ⓘ: = Ar, C, Can, ETx1, Fl1, FNA2, GrPl, Il, K4, Mo1, NE, NS, NY, Pa, POWO, RAB, Tn, TxFerns, Va, W, WH3, WV, Shmakov, Batkin, & Vaganov (2018); = Cystopteris fragilis (L.) Bernh. var. protrusa Weath. — F, G, Sf, Tat; < Cystopteris fragilis (L.) Bernh. — Tx; < Filix fragilis (L.) Underw. — S13. Basionym: Cystopteris fragilis (L.) Bernh. var. protrusa Weath. 1935
Links to other floras: = Cystopteris protrusa - FNA2
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Wetland Indicator Status:
- Atlantic and Gulf Coastal Plain: FAC
- Eastern Mountains and Piedmont: FAC
- Great Plains: FACW
- Midwest: FACW
- Northcentral & Northeast: FACU
Heliophily ⓘ: 2
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native range: eastern North America
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