Cystopteris tennesseensis Shaver. Common name: Tennessee Bladder Fern. Phenology: Apr-Jun. Habitat: Moist to dry outcrops of calcareous rocks, including coquina limestone (‘marl’) in the outer Coastal Plain. Distribution: PA, KY, IL, WI, and IA south to NC, nw. GA, n. AL, AR, and OK.
Origin/Endemic status: Native
Taxonomy Comments: This species is a fertile allotetraploid derived from hybridization between C. bulbifera and C. protrusa. Its genome can be symbolized BBPP. Haufler, Windham, & Ranker (1990) consider this a "successfully fledged and vigorous young species", adapted to a hybrid niche not successfully utilized by either parent.
Synonymy ⓘ: = Ar, C, FNA2, GrPl, Il, K4, Mo1, NS, Pa, RAB, Tn, Va, W; = Cystopteris ×tennesseensis Shaver — WV, Shmakov, Batkin, & Vaganov (2018); = Cystopteris tennesseensis Shaver ssp. tennesseensis — POWO, Windham & Pryer (2022); = n/a — Tat. Basionym: Cystopteris tennesseensis Shaver 1950
Links to other floras: = Cystopteris tennesseensis - FNA2
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Heliophily ⓘ: 2
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native range: eastern & central United States
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