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Cystopteris tennesseensis Shaver. Tennessee Bladder Fern. Phen: Apr-Jun. Hab: Moist to dry outcrops of calcareous rocks, including coquina limestone (‘marl’) in the outer Coastal Plain. Dist: PA, KY, IL, WI, and IA south to NC, nw. GA, n. AL, AR, and OK.

Origin/Endemic status: Native

Taxonomy Comments: This species is a fertile allotetraploid derived from hybridization between C. bulbifera and C. protrusa. Its genome can be symbolized BBPP. Haufler, Windham, & Ranker (1990) consider this a "successfully fledged and vigorous young species", adapted to a hybrid niche not successfully utilized by either parent.

Synonymy: = Ar, C, FNA2, GrPl, Il, K1, K3, K4, Mo1, Pa, RAB, Tn, Va, W; = Cystopteris ×tennesseensis – WV, Shmakov, Batkin, & Vaganov (2018); = Cystopteris tennesseensis Shaver ssp. tennesseensis – Windham & Pryer (2022); = n/a – Tat

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Heliophily ?: 2

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image of plant© Alan Cressler: Cystopteris tennesseensis, Madison County, Alabama 1 by Alan Cressler source | Original Image ⭷
image of plant© Alan Cressler: Cystopteris tennesseensis, Atwood Gulf, Pigeon Mountain, Crockford-Pigeon Mountain Wildlife Management Area, Walker County, Georgia 2 by Alan Cressler source | Original Image ⭷
image of plant© Alan Cressler: Cystopteris tennesseensis, Atwood Gulf, Pigeon Mountain, Crockford-Pigeon Mountain Wildlife Management Area, Walker County, Georgia 4 by Alan Cressler source | Original Image ⭷

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