Dryopteris campyloptera (Kunze) Clarkson. Phen: Jul-Sep. Hab: Spruce-fir forests, northern hardwood forests. Dist: NL (Newfoundland) and n. QC south to extreme n. PA, and from extreme s. PA south through e. WV and w. VA to e. TN and w. NC
Origin/Endemic status: Native
Taxonomy Comments: This species is a fertile allotetraploid derived from hybridization of D. intermedia and the northern and western D. expansa (C. Presl) Fraser-Jenkins & Jermy, which does not (now) reach our area. The chromosome complement is symbolized EEII.
Synonymy: = C, FNA2, K1, K3, K4, NE, NY, Pa, RAB, Sf, Tn, Va, W, WV, Montgomery & Paulton (1981); = Dryopteris austriaca (Jacq.) Woyn. ex Schinz & Thell. var. austriaca – G; = Dryopteris spinulosa (O.F.Müll.) Watt var. americana (Fisch. ex Kunze) Fernald – F; >< Dryopteris dilatata (Hoffm.) A.Gray – S13, misapplied