Dryopteris clintoniana (D.C. Eaton) Dowell. Clinton's Woodfern, Broad Swamp Fern. Phen: Jun-Sep. Hab: Acid seepages, swampy forests, red maple swamps. Dist: NB, QC, and ON, south to DE, DC, MD (Somerset Co.), n. VA (Arlington and Fairfax counties), PA, OH, IN, and IL. Reports from GA are based on the misapplication of the name to Dryopteris goldieana by McVaugh & Pyron (1951).

Origin/Endemic status: Native

Taxonomy Comments: This species is a fertile allohexaploid derived from hybridization of Dryopteris cristata and Dryopteris goldiana; its chromosome complement is symbolized GGLLSS.

Synonymy: = AqW, C, FNA2, G, K1, K3, K4, NE, NY, Pa, S13, Tat, Va, Montgomery & Paulton (1981); = Dryopteris ×clintoniana – Il; = Dryopteris cristata (L.) A.Gray var. clintoniana (D.C.Eaton) Underw. – F

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Wetland Indicator Status:

• Atlantic and Gulf Coastal Plain: FACW
• Eastern Mountains and Piedmont: FACW
• Midwest: FACW
• Northcentral & Northeast: FACW

Heliophily: 4