Nyssa biflora Walter. Swamp Tupelo, Water Gum, Swamp Black Gum. Phen: Apr-Jun; Aug-Oct. Hab: Blackwater river swamps, depressions in pinelands, pocosins, either where inundated for substantial periods of time or in more-or-less permanently saturated organic peaty soils. Dist: NJ south to s. FL, west to e. TX, primarily on the Coastal Plain, but scattered inland to c. NC, w. SC, c. TN, w. KY (Clark et al. 2005), se. MO, and c. AR.

Origin/Endemic status: Endemic

Synonymy: = FNA12, G, Il, K1, K3, K4, Mo2, S, S13, Va, Burckhalter (1992); = Nyssa biflora Walter ssp. biflora – Zhou et al (2018); = Nyssa biflora Walter var. biflora – Ward (2008b); = Nyssa sylvatica Marshall var. biflora (Walter) Sarg. – C, F, RAB, Tat, Tn, Tx, Eyde (1966), Wen & Stuessy (1993); < Nyssa sylvatica Marshall var. biflora (Walter) Sarg. – Fl5, GW2, WH3

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Wetland Indicator Status:

• Atlantic and Gulf Coastal Plain: OBL
• Eastern Mountains and Piedmont: FACW
• Great Plains: OBL
• Northcentral & Northeast: OBL

Heliophily: 5