*Perilla frutescens (Linnaeus) Britton. Perilla, Beefsteak-plant, Shiso. Phen: Aug-Oct; Oct-Dec. Hab: Moist disturbed areas. Dist: Native of India.

Origin/Endemic status: E. and se. Asia

Taxonomy Comments: Var. crispa (Bentham) Deane (leaves purple above and below; leaf margins laciniate-dentate and also crisped) and var. frutescens (leaves purple below; leaf margins dentate, not crisped) are sometimes recognized; these appear to be recently-derived cultivars rather than taxonomically distinct entities.

Synonymy: = Ar, C, Fl6, G, GrPl, K4, Mi, Pa, RAB, S, Tn, Tx, W, WH3; = Perilla fructescens – NcTx, orthographic variant; > Perilla frutescens (L.) Britton var. crispa (Benth.) W.Deane – F, Il, K1, K3, NE, NY, Tat, WV; > Perilla frutescens (L.) Britton var. frutescens – F, Il, K1, K3, NE, NY, WV

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Wetland Indicator Status:

• Atlantic and Gulf Coastal Plain: FACU
• Eastern Mountains and Piedmont: FACU
• Great Plains: FAC
• Midwest: FAC
• Northcentral & Northeast: FAC

Heliophily: 5