Rubus heterophyllus Willdenow. Pleatleaf Dewberry. Phen: May-Jun. Hab: Typically occurs on poor, well-drained soils, both sands and gravels. Dist: NS to MN, south to w. VA (Roanoke Co.), WV, and IA

Origin/Endemic status: Native

Synonymy: > Rubus botruosus L.H.Bailey; > Rubus coloniatus L.H.Bailey; > Rubus exutus L.H.Bailey; < Rubus pensilvanicus Poir. – FNA9; > Rubus plicatifolius Blanch. – Il, K4, Ladd & Thomas (2015); > Rubus prior L.H.Bailey; > Rubus problematicus L.H.Bailey; < Rubus recurvicaulis Blanch. – C; > Rubus rhodinsulanus L.H.Bailey; > Rubus rosendahlii L.H.Bailey; > Rubus semierectus Blanch.; > Rubus victorinii L.H.Bailey

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Wetland Indicator Status:

• Eastern Mountains and Piedmont: FAC (taxonomic split from wetland indicator species)
• Great Plains: FAC (taxonomic split from wetland indicator species)
• Midwest: UPL (taxonomic split from wetland indicator species)
• Northcentral & Northeast: FACU (taxonomic split from wetland indicator species)