A family of a single genus and about 750 species, of nearly cosmopolitan distribution. Murakami et al. (1999) conducted a molecular phylogenetic analysis of the Aspleniaceae, which confirmed that Camptosorus should be included in Asplenium, but suggested that Phyllitis is better separated from Asplenium. Later and more comprehensive studies have shown that Phyllitis and Camptosorus are deeply embedded in Asplenium (Schneider et al. 2004a; Xu et al. 2020), a conclusion followed here.
Ref: Kessler & Smith (2018); Kramer & Viane In Kramer & Green (1990); Murakami et al. (1999); PPG I (2016); Schneider et al. (2004a); Wagner, Moran, & Werth (1993) In Flora of North America Editorial Committee (1993b); Xu et al. (2020). Show full citations.