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Barker, M.S., and W.D. Hauk. 2003. An evaluation of Sceptridium dissectum (Ophioglossaceae) with ISSR markers: implications for Sceptridium systematics. Amer. Fern J. 93(1): 1-19.

Cao, D., and W.D. Hauk. 2022. Assessing morphological species and interspecific relationships in North American grapeferns (Sceptridium; Ophioglossaceae) using ISSR markers. Syst. Bot. 47(3) 748-761.

Hauk, W.D. 1996. Phylogenetics of Ophioglossaceae: the evolutionary consequences of morphological reduction. Ph.D. dissertation, University of North Carolina at Chapel Hill, Biology Dept.

Hauk, W.D., C.R. Parks, and M.W. Chase. 2003. Phylogenetic studies of Ophioglossaceae: evidence from rbcL and trnL-F plastid DNA sequences and morphology. Molec. Phylogen. Evol. 28: 131-151.

PPG I. 2016. A community-derived classification for extant lycophytes and ferns. J. Syst. Evol.

Wagner, W.H., Jr., and F.S. Wagner. 1993. Ophioglossaceae. In Flora of North America Editorial Committee. 1993b. Flora of North America north of Mexico. Volume 2, pteridophytes and gymnosperms. Oxford Univ. Press, New York, NY. 475 pp.

Zhang, L., and L.-B. Zhang. 2022. Phylogeny, character evolution, and systematics of the fern family Ophioglossaceae based on Sanger sequence data, plastomes, and morphology. Molec. Syst. Evol. 173: 107512.

Zhang, L., X.‐P. Fan, S. Petchsri, L. Zhou, R. Pollawatn, X. Zhang, X.‐M. Zhou, N.T. Lu, R. Knapp, S. Chantanaorrapint, P. Limpanasittichai, H. Sun, X.‐F. Gao, and L.‐B. Zhang. 2020. Evolutionary relationships of the ancient fern lineage the adder's tongues (Ophioglossaceae) with description of Sahashia gen. nov. Cladistics 36: 380-393.