Asplenium heteroresiliens W.H. Wagner. Marl Spleenwort, Carolina Spleenwort, Wagner's Spleenwort, Morzenti's Spleenwort. Phen: Apr-Oct. Hab: Fairly moist outcrops of calcareous sedimentary rocks, such as coquina limestone (‘marl’), along small blackwater streams or larger rivers, at low elevations, and rarely also on old ruins made of tabby (a cement made from lime, sand, and oyster shells). Dist: Rare and scattered from se. NC to se. GA, sw. GA, and n. FL, on the Coastal Plain

Origin/Endemic status: Endemic

Other Comments: This species is an apogamous (producing viable spores asexually) allopentaploid derived from hybridization of the sexual tetraploid H. heterochroum Kunze and the apogamous triploid A. resiliens. Its chromosome complement can be symbolized EEEHH.

Synonymy: = K4, RAB; = Asplenium ×heteroresiliens – Fl1, FNA2, K1, K3, WH3; < Asplenium heterochroum Kunze – Sf

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