Uvularia puberula Michaux. Section: Oakesiella. Carolina Bellwort, Appalachian Bellwort, Coastal Bellwort. Phen: Late Mar-Early May; Aug-Oct. Hab: Dry to moist upland, acidic forests, up to at least 1500m. Dist: S. PA to GA in the Mountains and (more rarely) Piedmont, and from Long Island NY south to GA in the Coastal Plain and Sandhills.

Origin/Endemic status: Endemic

Other Comments: Sometimes interpreted as having two varieties, the montane and Piedmont plants as var. puberula (leaves broader, rounded to slightly clasping at the base, firm in texture, obviously reticulate on the lower surface, the stem puberulent in lines on the ridges) and var. nitida (Britton) Fernald (leaves narrower, more cuneate, thin in texture, the reticulation of cross veins less evident, the stems glabrous); see Wilbur (1963b) and Uttal (1991)While Wilbur (1963b) chose not to recognize varieties, Uttal (1991) supported varietal recognition. The differences seem minor and variable, and poorly correlated with geography.

Synonymy ⓘ: = C, FNA26, K1, K3, K4, NY, Tn, Va, W, Wilbur (1963b); = Oakesiella puberula (Michx.) Small – S, S13; = Uvularia caroliniana (J.F.Gmel.) Wilbur; = Uvularia pudica (Walter) Fernald – G, Pa, RAB, WV, nomen dubium; > Uvularia puberula Michx. var. nitida – Uttal (1991); > Uvularia puberula Michx. var. puberula – Uttal (1991); > Uvularia pudica (Walter) Fernald var. nitida (Britton) Fernald – F; > Uvularia pudica (Walter) Fernald var. pudica – F

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Wetland Indicator Status:

• Atlantic and Gulf Coastal Plain: FAC
• Eastern Mountains and Piedmont: FACU
• Northcentral & Northeast: FACU

Heliophily ⓘ: 4