Pityopsis aspera (Shuttleworth ex Small) Small var. adenolepis (Fernald) Semple & F.D. Bowers. Phen: Late Jun-Oct. Hab: Longleaf pine sandhills, dry woodlands, forests, and disturbed places, apparently in the NC Mountains only in the Escarpment. Dist: E. MD (R. Simmons, pers. comm., 2016) and e. and c. VA south to n. FL and west to s. MS.

Origin/Endemic status: Endemic

Taxonomy Comments: P. aspera var. adenolepis includes 2 chromosome numbers (2n = 18 and 36), which "account, in part, for the range of variation in involucre, floret, and fruit size" (Semple & Bowers 1985). Nesom (2019a) subsumes this variety under a more broadly defined P. aspera, since he did not find a clear morphological or geographic separation between these varieties. We are tentatively maintaining the varieties, pending further field study of this variation.

Synonymy: = FNA20, K1, K3, Va, Semple & Bowers (1985); = Chrysopsis graminifolia (Michx.) Elliott – F, misapplied; = Heterotheca adenolepis (Fernald) H.E.Ahles – Bowers (1972); = Pityopsis adenolepis (Fernald) Semple; < Chrysopsis graminifolia (Michx.) Elliott var. aspera (Shuttlew. ex Small) A.Gray – C, G, SE1, W; > Heterotheca adenolepis (Fernald) H.E.Ahles – RAB; < Heterotheca aspera (Shuttlew. ex Small) Shinners; > Heterotheca graminifolia (Michx.) Shinners – RAB, misapplied; < Pityopsis aspera (Shuttlew. ex Small) Small – Fl7, K4, S, WH3, Nesom (2019a)

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