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Dryopteris intermedia (Muhlenberg ex Willdenow) A. Gray. Fancy Fern, Evergreen Woodfern. Phen: Jun-Sep. Hab: Cove forests, other moist, rocky forests, over a variety of substrates. Dist: NL (Newfoundland) west to MN, south to n. GA and s. MO.

Origin/Endemic status: Native

Taxonomy Comments: This species is one of the diploid "parent species" of the e. North American reticulately-evolved Dryopteris complex. Its genome (symbolized II) forms half of the genome of the tetraploids Dryopteris campyloptera and Dryopteris carthusiana and 2/3s of the genome of the triploid hybrid Dryopteris ×triploidea (Dryopteris carthusiana × intermedia, with chromosome symbolized IIS), which is frequent in areas with both parents present.

Synonymy: = C, FNA2, Il, K1, K3, K4, Mo1, NE, NY, Pa, RAB, S13, Sf, Tat, Tn, Va, W, WV, Montgomery & Paulton (1981); = Dryopteris austriaca (Jacq.) Woyn. ex Schinz & Thell. var. intermedia (Muhl. ex Willd.) Morton – G; = Dryopteris spinulosa (O.F.Müll.) Watt var. intermedia (Muhl. ex Willd.) Underw. – F

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image of plant© Alan Cressler: Dryopteris intermedia, Southern Appalachian Mountain Bog, North Carolina 1 by Alan Cressler source
image of plant© Alan Cressler: Dryopteris intermedia, Chestnut Cove Boulder Field, Brasstown Bald, Brasstown Wilderness, Chattahoochee National Forest, Towns County, Georgia 4 by Alan Cressler source
image of plant© Gary P. Fleming
image of plant© Erik Danielson source | Original Image ⭷
image of plant© Alan Cressler: Dryopteris campyloptera (left), Dryopteris intermedia (right) , Big Frog Mountain, Big Frog Wilderness, Cherokee National Forest, Polk County, Tennessee 1 by Alan Cressler source
image of plant© Alan Cressler: Dryopteris carthusiana (right), and Dryopteris intermedia (left), Southern Appalachian Mountain Bog, North Carolina 1 by Alan Cressler source
image of plant© Keith Bradley | Original Image ⭷
image of plant© Alan M. Cressler | Original Image ⭷
image of plant© Alan Cressler: Dryopteris intermedia (top), Dryopteris intermedia X marginalis (middle), Dryopteris marginalis (bottom), Guess Creek, Jackson County, Alabama 1 by Alan Cressler source
image of plant© Erik Danielson source | Original Image ⭷
image of plant© Alan Cressler: Dryopteris celsa (left), Dryopteris Xseparabilis (center), Dryopteris intermedia (right), Big Creek, Great Smoky Mountains National Park, Haywood County, North Carolina 1 by Alan Cressler source