1 Flowers (all, or at least many) solitary on scapes
5 Petals 6-30 mm long, clawed
5 Petals 2-4 mm long, not clawed
6 Stems angular distally; blooming Apr to early Jun
6 Stems not angular distally; blooming Apr to Oct
3 Leaves not
clasping at
base .
7 Petals with contrasting dark yellow, brown, maroon, or purple veins; [tribe
Brassiceae ]
8 Petals yellow;
basal leaves with 3-10 lobes per
margin ; stems usually sparsely to densely
hispid basally;
cauline leaf blades usually
lobed (but fewer than
basal leaves)
8 Petals yellow or creamy white;
basal leaves with 1-4 lobes per
margin ; stems sparsely to densely
pubescent ;
cauline leaf blades often undivided
7 Petals mostly uniform in color.
9 Ovaries and young fruits 2-segmented; petals 5-30 mm long; [tribe
Brassiceae ]
10 Stem leaves
pinnatifid
10 Stem leaves
entire ,
serrate , or
sinuate
11 Valves of the fruit 1-veined; sepals erect or
ascending (rarely spreading); terminal
segment of fruit
conic , usually seedless
11 Valves of the fruit 3-7-veined (1-veined in
Rhamphospermum nigrum ); sepals spreading to
reflexed ; terminal
segment of fruit
either linear or ensiform (flattened and 2-
edged ).
12 Terminal
segment of fruit
linear 0.1-1.6 cm long, seedless or 1-seeded.
12 Terminal
segment of fruit
ensiform (flattened and 2-
edged ), (1-) 1.5-2.5 (-3) cm long, seedless
9 Ovaries and young fruits unsegmented; petals 1-8 mm long.
13 Inflorescence leafy
bracteate (most of the pedicels subtended by bracts)
13 Inflorescence not
bracteate (occasionally with some lower pedicels bearing bracts).
14 Stigmas distinctly 2-
lobed ; [tribe
Sisymbrieae ]
14 Stigmas
entire , rarely indistinctly 2-
lobed ; [tribe
Cardamineae ]
2 Flowers white, cream, pinkish, purple, lavender, or blue.
15 Leaves strictly
basal ,
or both
basal and
cauline and also
auriculate .
16 Plants with only
basal leaves or
cauline leaves much reduced.
17 Plants annual; flowers
zygomorphic ; petals 0.5-2.5 mm long; [tribe
Iberideae ]
17 Plants annual or perennial; flowers
actinomorphic ; petals absent,
rudimentary , or to 16 mm long; [tribe
Cardamineae ]
16 Plants with some well-developed
cauline leaves.
18 Ovaries and young fruits
ovate ,
orbicular or
cordate .
19 Ovules 1 per
ovary ; [tribe
Calepineae ]
20 Ovules 2 (rarely 4) per
ovary ; [tribe
Lepidieae ]
20 Ovules 6-16 per
ovary .
21 Leaves mostly
entire ; plants not
fetid ; [tribe
Noccaeeae ]
21 Leaves often
toothed ; plants
fetid ; [tribe
Thlaspideae ]
18 Ovaries and young fruits
linear .
22 Plants aquatic or
semi- aquatic; rooting at nodes; [tribe
Cardamineae ]
22 Plants terrestrial, though sometimes growing partially submerged; not rooting at nodes.
23 Flowers blue, rarely pink or white; petals 12-32 mm long; [tribe
Brassiceae ]
23 Flowers white, sometimes pink or purple; petals <16 mm long.
24 Petals larger, conspicuous, 7-16 mm long.
25 Cauline leaves simple or
pinnately to
palmately compound , typically not
auriculate ; if simple
and auriculate , then plants usually with a mix of compound and simple
cauline leaves and the
simple leaves ovate or
oblong and never deeply
pinnatifid (as in
C. clematitis )
25 Cauline leaves conspicuously or slightly
auriculate , always simple; blades
elliptic ,
lanceolate ,
ovate , or
oblong , deeply
pinnatifid to
entire
26 Basal leaves
oblanceolate ,
unlobed ; petals white to cream
26 Basal leaves
incised or
lyrate -
lobed ; petals white to pinkish-purple
24 Petals smaller, inconspicuous, < 7 mm long.
27 Cauline leaves
sessile ; [tribe Boechereae]
27 Cauline leaves usually
petiolate ; [tribe Cardamineae]
15 Leaves
basal and
cauline but not
auriculate (
basal leaves sometimes withering upon flowering, thus only appearing alternate).
28 Plants aquatic, rooting at nodes; [tribe Cardamineae].
29 Submersed leaves
dissected into
filiform segments
29 Leaves not
dissected into
filiform segments
28 Plants terrestrial, though sometimes growing partially
submersed ; not rooting at nodes.
30 Stamens 2 or 4 (per individual flower).
31 Ovaries and young fruits
linear ; [tribe Cardamineae]
31 Ovaries and young fruits
ovate ,
orbicular or
cordate .
32 Flowers
zygomorphic ; filaments appendaged; [tribe Iberideae]
32 Flowers
actinomorphic ; filaments not appendaged; [tribe Lepidieae]
30 Stamens 6 (per individual flower).
33 Petals 15-30 mm long; fruit conspicuously coin-like,
oblong to suborbicular and latiseptate (flattened parallel to
septum ); [tribe Lunarieae]
33 Petals rarely to 20 mm long; fruit
ovate to
linear (if rounded not strongly flattened like a coin, or if flattened the fruit long and slender as in
Warea ).
34 Inflorescence corymbose ; claws (of the petals) strongly differentiated from blades; fruit latiseptate and
stipitate ,
linear in shape; [natives of sandhills and coastal plain, NC to s. MS]
34 Inflorescence elongate or
corymbose (if so, not also collectively as above); claws differentiated or not from blades; fruit
terete , latiseptate, or augustiseptate (flattened perpindicular to
septum ), stipate or not,
linear to
ovate in shape; [widespread natives and non-natives]
35 Ovaries and young fruits 2-segmented; [tribe Brassiceae]
35 Ovaries and young fruits unsegmented.
36 Ovaries and young fruits
ovate ,
orbicular or
cordate ; [tribe Lepidieae]
37 Ovules 2 (-4) per
ovary
36 Ovaries and young fruits
linear .
38 Petals with contrasting dark veins; [tribe Brassiceae]
38 Petals mostly uniform in color.
39 Leaves smelling of garlic when crushed,
reniform or
cordate ,
dentate ; [tribe Thlaspideae]
39 Leaves not smelling of garlic, rarely
reniform or
cordate , with margins various.
40 Cauline leaves
sessile ; [tribe Boechereae]
40 Cauline leaves usually
petiolate ; [tribe
Cardamineae ].
41 Base of plant usually
glabrous ; seeds not
winged
41 Base of plant
pubescent ; seeds
winged
1 Fruits silicles (< 3× as long as wide).
3 Plants
succulent , leaves usually fleshy
3 Plants
herbaceous , not
succulent , leaves not fleshy
4 Fruits 2 cm or more wide
5 Fruits with 20 or more seeds
5 Fruits with 16 or fewer seeds.
6 Fruits with 2-16 seeds.
8 Fruits with more than 2 seeds.
10 Stems 5-12 (-20) dm tall; leaf blades of
basal leaves (3-) 5-12 (-17) cm long
10 Stems 0.5-1.5 (-2) dm tall; leaf blades of
basal leaves 0.3-1.5 (-2) cm long
11 Plants not
fetid ; seeds usually smooth, yellowish to medium brown
11 Plants
fetid ; seeds
striate or
alveolate , dark gray to dark brown or black
1 Fruits siliques (> 3× as long as wide).
12 Fruits
indehiscent or breaking into 1-seeded segments.
13 Fruits not segmented, 1-seeded
13 Fruits segmented, usually more than 1-seeded.
14 Styles
obsolete ; plants
glabrous ; [coastal]
14 Styles 1-5 mm; plants
pubescent ; [inland]
12 Fruits
dehiscent , more than 1-seeded.
16 Cauline leaf
blade bases
auriculate (except
B. juncea ); terminal
silique segment seedless (rarely 1-2-seeded),
conic
16 Cauline leaf
blade bases not
auriculate ; terminal
silique segment 1-5-seeded (sometimes 0).
17 Fruit valves 3-7-veined.
18 Valves of the fruit 3 (-5) veined
18 Valves of the fruit 3-5 (-7) veined.
19 Terminal
segment of fruit
linear 0.1-1.6 cm long, seedless or 1-seeded.
19 Terminal
segment of fruit
ensiform (flattened and 2-
edged ), (1-) 1.5-2.5 (-3) cm long, seedless
20 Fruits (all, or at least some) solitary, borne on scapes
20 Fruits in racemes (or
corymbose clusters).
21 Fruits latiseptate (flattened parallel to the
septum ).
22 Replums strongly flattened; fruit valves dehiscing elastically, coiled
22 Replums
terete or rounded; fruit valves not dehiscing elastically or coiled.
24 Fruit
stipitate ;
cauline leaves
entire , not
auriculate ; flowers pink to purple (or white in
W. carteri ), [natives of sandhills and coastal plain, NC to s. MS]
24 Fruit
sessile or short-
stipitate ;
cauline leaves usually
lobed or at least
dentate , the bases
auriculate or somewhat
clasping ; flowers yellow; [non-natives, widespread]
23 Seeds
winged at least distally.
21 Fruits
terete or 4-angled.
26 Plants aquatic or
semi- aquatic; rooting at nodes
26 Plants terrestrial, though rarely growing partially
submersed ; not rooting at nodes.
27 Fresh leaves smelling of garlic when crushed,
reniform or
cordate ,
dentate
27 Fresh leaves not smelling of garlic when crush, rarely
reniform or
cordate ,
dentate or not.
29 Stigma lobes obviously
decurrent
29 Stigma lobes sometimes distinct but not
decurrent .
30 Lower
cauline leaves
entire or
dentate .
31 Cauline leaves spaced, not concealing stem, spreading to slightly
ascending ; petals 7-12 mm long
32 Basal leaves
obovate to
oblanceolate ,
unlobed ;
cauline leaves
entire ; siliques 8-13 cm long
32 Basal leaves
incised or
lyrate -
lobed ;
cauline leaves with
dentate to
serrate margins; siliques 2-4 cm long
31 Cauline leaves usually ascendant and concealing stems (at least basally); petals 2-5 mm long (except for 5-10 mm long in
B. missouriensis )
33 Terminal
leaflet lobes nearly
entire to
crenate or
undulate
33 Terminal
leaflet lobes sharply
toothed or cut
34 Basal leaves usually withered by flowering;
cauline leaf lobes
entire to lyrately
lobed ; usually petals white to pink; [native of
riparian habitats]
34 Basal leaves usually present at flowering or after;
cauline leaves with
linear ,
filiform ,
lyrate , or
runcinate lobes (occasionally
unlobed , but if so then leaves usually
linear or otherwise thin); petals yellow; [non-natives of various habitats]