1 Ligule with a dense ring of short
hairs 0.5-1 mm long, and behind it a ring of longer
hairs 1-5 (-8) mm long called the pseudoligule (see introduction for discussion of the so-called double
ligule ).
2 Culm bases hard, thickened,
corm -like; plants producing
basal branches instead of
rosettes , the branches ultimately producing the largest leaves, 4-13 cm long and 4-11 mm wide;
vernal spikelets 2.3-2.7 mm long, 1.2-1.5 mm wide,
autumnal spikelets 2.6-2.9 mm long, 1.0-1.1 mm wide;
spikelets basally
attenuate , the
glumes attached 0.2-0.4 mm below expanding
base of upper
glume and lower
lemma ; [currently known only from sand ridges in peninsular FL]
2 Plants without this combination of characters.
3 Culms > 100 cm long;
node pubescence densely
retrorse ; larger
blades 12-20 cm long;
spikelets acute ;
vernal spikelets 1.9-2.3 mm long, slightly < 2× as long as wide;
autumnal spikelets 2.2-2.5 mm long, > 2× as long as wide
3 Culms < 100 cm long (rarely longer);
node pubescence various; larger
blades 3-12 cm long;
spikelets rounded to blunt, < 2× as long as wide.
4 Plants velvety-
pubescent throughout;
nodes retrorsely bearded ;
internode pubescence retrorse to spreading;
blades usually
ciliate above
base as well as basally; mature
vernal panicles 3-7 cm long, 2-5 cm wide, about 2/3 as wide as long
4 Plants densely
pubescent to
glabrate , but lacking velvety
pubescence ;
nodes with spreading to
ascending pubescence (sometimes
retrorse in
D. villosissimum );
internode pubescence ascending ,
appressed , or spreading;
blades ciliate only at
base (sometimes appearing
ciliate laterally in
D. ovale var.
ovale ); mature
vernal panicles 4-10 cm long, nearly or equally as wide.
5 Lower
internodes with
retrorse or spreading
hairs to 3+ mm long;
blade adaxial surface
pilose with
hairs to 4+ mm long, these
appressed and angled away from the leaf
axis ;
spikelets 2.1-2.5 mm long
6 Spikelets 2.1-2.6 mm long; lower
culm blades usually sparsely
appressed -
pubescent on both surfaces, eciliate or
ciliate at the
base only
6 Spikelets 2.5-3.1 mm long; lower
culm blades usually
glabrous adaxially except for long
hairs at or near the
margin (appearing
ciliate ),
appressed -
pubescent abaxially
1 Ligule a single structure, without a pseudoligule.
7 Culm bases hard, thickened,
corm -like; plants producing
basal branches instead of
rosettes , the branches ultimately producing the largest leaves, 4-13 cm long and 4-11 mm wide;
vernal spikelets 2.3-2.7 mm long, 1.2-1.5 mm wide,
autumnal spikelets 2.6-2.9 mm long, 1.0-1.1 mm wide;
spikelets basally
attenuate , the
glumes attached 0.2-0.4 mm below expanding
base of upper
glume and lower
lemma ; [currently known only from dry sand ridges in peninsular FL]
7 Plants without this combination of characters.
8 Ligule 2-5 mm long,
ciliate
9 Ligule a stramineous to light brown membrane, with or without terminal
cilia ;
peduncle antrorsely scabrous but not hairy.
10 Panicle rachis smooth,
pellucid -
punctate ; first
glume 0.3-0.6 (-0.8 ) mm long, as broad as or broader than long,
truncate to
obtuse ; larger leaves 10-25 cm long, 8-15 mm wide;
ligule 0.5-1.3 mm long; lowest elongate
culm internode > 2 mm in diameter; lowest
nodes usually
glabrous or
pubescent
10 Panicle rachis scabrous or smooth, not
pellucid -
punctate ; first
glume 0.5-1.1 mm long, longer than wide, rounded to
acute ; larger leaves 3.5-12 cm long, 3-9 mm wide;
ligule 0.1-0.6 mm long; lowest elongate
culm internode < 2 mm in diameter; lowest
nodes retrorsely bearded or
glabrous .
11 Lowest
nodes usually
retrorsely bearded ;
ligules (0.1-) 0.3-0.6 mm long; largest
vernal blades 7-12 cm long, (4.5-) 6-9 mm wide;
panicle peduncle scabrous ;
spikelets ovate -
lanceolate ,
acute , 2.0-2.4 mm long; first
glume lanceolate , blunt to
acute ; fertile
lemma smooth
11 Lowest
nodes usually
glabrous ;
ligules 0.1-0.2 (-0.3) mm long; largest
vernal blades 3.5-7 cm long, 3-6 mm wide;
panicle peduncle smooth;
spikelets elliptic , blunt to
acute , 1.6-2.2 mm long; first
glume ovate to
rotund , rounded to
acute ; fertile
lemma papillose
9 Ligule entirely of white
hairs ;
peduncle variously hairy or
glabrous , but not
antrorsely scabrous .
12 Culms without a
viscid band below the
retrorsely , spreading, or
antrorsely bearded nodes ;
internodes and
sheaths not velvety-
pubescent ,
blades velvety-
pubescent in some.
13 Basal as well as
cauline blades linear , 5-15 cm long,; 1-4 mm wide, 30× or more as long as wide;
spikelets 2.8-3.8 (-4.5) mm long
13 Blades < 15× as long as wide (if > 15×, then
spikelets 1.9-2.5 mm long and
sheaths retrorsely pilose in
D. laxiflorum ).
14 Blades (15-) 20× or more as long as wide, increasing in length upwards on
culm ;
blade adaxial surface (weakly-) strongly
scabrous ;
sheaths glabrous ; second
glume and
sterile lemma exceeding fertile
lemma
14 Blades < 15× as long as wide, similar or more often decreasing in size upwards on
culm (if increasing, then
sheaths strongly
retrorsely pilose in
D. laxiflorum );
blade adaxial surface smooth to moderately
scabrous ; second
glume and
sterile lemma shorter than to exceeding fertile
lemma .
15 Nodes retrorsely bearded ; larger
vernal leaves 7-15 mm wide.
16 Sheaths retrorsely pilose with
hairs 2-3 mm long;
basal leaves usually numerous,
ascending , similar in size and shape to the
culm leaves;
culms branching only at the
base in
autumnal phase
16 Sheaths glabrous or
pilose (if
pilose , then
hairs not both
retrorse and 2-3 mm long);
basal leaves
rosette -forming, usually much shorter than
culm leaves;
culms branching above
base in
autumnal phase.
17 Plants rarely > 1 m in length; lower
internodes glabrous to sparsely hairy;
vernal cauline leaves 3-7;
spikelets 1.8-2.8 mm long,
elliptic to lance-
ovate , blunt to
acute ,
glabrous or
pubescent
17 Plants often > 1 m in length; lowest
internodes moderately to densely
villous ;
vernal cauline leaves 6-15;
spikelets 1.8-2.2 mm long,
obovoid to broadly
ellipsoid , rounded to blunt,
pubescent
15 Nodes spreading to
antrorsely bearded ; larger
vernal leaves 3-10 mm wide.
18 Culm internodes , at least the lower,
strigose ,
pilose , or
villous ;
culm nodes bearded with
ascending or spreading
hairs ;
blade surfaces
glabrous or variously hairy.
19 Lower
nodes bearded with erect-
ascending , soft, and long
hairs ; mid-
culm blades usually 20× or more as long as wide.
20 Spikelets (3.0-) 3.2-3.8 mm long,
fusiform , pointed at summit,
attenuate at
base , with both
glumes attached 0.3-0.5 mm below
sterile lemma , the
autumnal spikelets 3.5-3.8 mm long; larger
vernal blades 3-6 mm wide, the lower and mid-
culm blades of similar width;
autumnal blades involute
20 Spikelets 1.8-3.1 mm long,
obovate , blunt, and the
base not
attenuate except in
D. arenicoloides with
autumnal spikelets 2.3-3.1 mm long; larger
vernal blades 2-8 mm wide, the lower usually wider and often shorter than mid-
culm blades ;
autumnal blades involute or flat.
21 Longest
vernal blades to 16 cm, widest
vernal blades 4-8 mm, usually longitudinally
wrinkled ;
vernal and
autumnal spikelets 2.3-3.1 mm long;
autumnal blades flat, the larger to 9 cm × 2-4 mm
21 Longest
vernal blades to 6 (
D. aciculare ) or 12 (
D. arenicoloides ) cm long, 2-5 mm wide, not noticeably
wrinkled ;
vernal spikelets 1.5-2.8 mm long,
autumnal spikelets either 1.5-2.3 (
D. aciculare ) or 2.3-3.1 (
D. arenicoloides );
autumnal blades involute , the larger to 6 cm × 1.5 mm.
22 Longest
vernal blades to 12 cm, widest
vernal blades 3-4 (-5) mm;
vernal spikelets 2.1-2.8 mm long;
autumnal spikelets 2.3-3.1 mm long, pointed,
attenuate , the
glumes attached 0.3-0.5 mm below
sterile lemma ; first
glumes 0.7-1.5 mm long; larger
autumnal blades to 6 cm by 1.5 mm
19 Lower and often mid-
culm nodes bearded with spreading, stiffish, and short-to-long
hairs ; mid-
culm blades usually 15× or less as long as wide.
23 Blades stiff, often longitudinally
ribbed , at least the lower
villous or strongly
pilose on the
abaxial surface, and
pilose to
hirsute on the
adaxial surface, at least proximally.
24 Spikelets 2.3-3.0 mm long;
ligule < 1 mm long; mid-
culm blades 4-8 mm wide
24 Spikelets 1.7-2.0 (-2.3) mm long;
ligule 1-2.5 mm long; mid-
culm leaves 2-5 mm wide
23 Blades not noticeably stiff nor longitudinally
ribbed ,
pubescent or
strigose underneath,
glabrous above or with a few long
hairs near the
base .
25 Spikelet bases strongly
attenuate ,
apices blunt to beaked; first
glume (2/5-) ½-2/3 (-4/5) as long as
spikelet , attached 0.3-0.5 (-0.7) mm belong expansion of second
glume and
sterile lemma ; larger
vernal blades 3-6 (-7) mm wide
25 Spikelet bases not noticeably
attenuate ,
apices rounded to
obtuse ; first
glume < ½ as long as
spikelet , attached < 0.3 mm below expansion of second
glume and
sterile lemma ; larger
vernal blades 5-12 mm wide.
26 Culms 50-140 cm long; lowest
internodes usually purplish; leaf
sheaths often
viscid -spotted; leaf
blades glabrous abaxially,
glabrous and
scabrous adaxially;
spikelets 1.8-2.2 mm long
26 Culms 15-60 cm long; lowest
internodes usually olive- or bluish-green; leaf
sheaths not
viscid -spotted; leaf
blades appressed -
pubescent abaxially,
glabrous and smooth to
appressed pubescent adaxially;
spikelets 2.1-3.1 mm long.
27 Spikelets 2.1-2.6 mm long; lower
culm blades usually sparsely
appressed -
pubescent on both surfaces, eciliate or
ciliate at the
base only
27 Spikelets 2.5-3.1 mm long; lower
culm blades usually
glabrous adaxially except for long
hairs at or near the
margin (appearing
ciliate ),
appressed -
pubescent abaxially
2 Leaf
blades , at least the lower,
cordate or subcordate at the
base , 4-25 mm wide.
3 Culm blades 4-12 mm wide;
spikelets obpyriform when viewed dorsally, strongly plano-
convex when viewed laterally, usually markedly reddish-purple basally; fertile
lemma papillose
3 Culm blades 5-25 mm wide;
spikelets elliptic to
elliptic -
obovoid when viewed dorsally or laterally, greenish to faintly purple-tinged basally; fertile
lemma not
papillose .
4 Ligule ciliate , 0.2-0.8 mm long;
culm blades 5-14 mm wide;
spikelets 1.8-2.6 mm long; first
glumes 0.5-1 mm long
4 Ligule an eciliate membrane 0-0.3 mm long;
culm blades 5-25 cm wide;
spikelets 2.0-3.7 mm long; first
glumes 0.5-2.6 mm long.
5 Internodes crisp-
puberulent ; larger
culm blades 4-8 (-11) cm long, 5-10 (-12) mm wide, broadest near the
base ;
spikelets 2.1-2.7 mm long; first
glumes 0.7-0.9 mm long
5 Internodes glabrous to sparsely
pubescent , larger
culm blades 5-18 cm long, 5-25 mm wide, broadest near the middle or broadly
linear ;
spikelets 2.0-3.7 mm long; first
glumes 0.5-2.6 mm long.
6 Cauline blades 5-18 cm long, 6-14 mm wide, averaging about 10 × as long as wide, the
margins nearly parallel;
spikelets 2.9-3.7 mm long; first
glumes 11-26 mm long, 2/5 or more as long as the
spikelets
6 Cauline blades 7-12 cm long, (6-) 12-25 mm wide,
ovate -
lanceolate , averaging about 6 × as long as wide, the
margins curved most of their length;
spikelets 2.0-3.2 mm long; first
glumes 5-10 mm long, about 1/4 as long as
spikelets .
7 Culms erect;
blades more nearly symmetrical;
spikelets 2.0-3.2 mm long;
sterile lemma apex usually blunt to rounded
7 Culms basally
decumbent ;
blades strongly asymmetrical-
falcate ;
spikelets 3.0-3.2 mm long;
sterile lemma apex usually pointed
2 Leaf
blades tapering to the
base , 2-12 mm wide.
8 Ligule a stramineous to light brown membrane, with or without terminal
cilia .
9 Ligule a
ciliate membrane;
spikelets (2.6) 2.8-3.4 (3.6) mm long; first
glume (0.9) 1.1-1.6 mm long
9 Ligule an eciliate membrane;
spikelets 1.6-2.8 (3.0) mm long; first
glume 0.3-1.1 mm long.
10 Panicle rachis smooth,
pellucid -
punctate ; first
glume 0.3-0.6 (-0.8 ) mm long, as broad as or broader than long,
truncate to
obtuse ; larger leaves 10-25 cm long, 8-15 mm wide;
ligule 0.5-1.3 mm long; lowest elongate
culm internode > 2 mm in diameter; lowest
nodes usually
glabrous or
pubescent
10 Panicle rachis scabrous or smooth, not
pellucid -
punctate ; first
glume 0.5-1.1 mm long, longer than wide, rounded to
acute ; larger leaves 3.5-12 cm long, 3-9 mm wide;
ligule 0.1-0.6 mm long; lowest elongate
culm internode < 2 mm in diameter; lowest
nodes retrorsely bearded or
glabrous .
11 Lowest
nodes usually
retrorsely bearded ;
ligules (0.1-) 0.3-0.6 mm long; largest
vernal blades 7-12 cm long, (4.5-) 6-9 mm wide;
panicle peduncle scabrous ;
spikelets ovate -
lanceolate ,
acute , 2.0-2.4 mm long; first
glume lanceolate , blunt to
acute ; fertile
lemma smooth
11 Lowest
nodes usually
glabrous ;
ligules 0.1-0.2 (-0.3) mm long; largest
vernal blades 3.5-7 cm long, 3-6 mm wide;
panicle peduncle smooth;
spikelets elliptic , blunt to
acute , 1.6-2.2 mm long; first
glume ovate to
rotund , rounded to
acute ; fertile
lemma papillose
8 Ligule of short white
hairs or absent.
12 Leaves basally disposed, usually matted or cushion-forming, larger than the mid and upper
culm leaves;
blade margins uniformly
papillose -
ciliate ;
culms branching only at the
base , 0.5-3.5 dm tall;
internodes glabrous or sparsely
pubescent
12 Basal leaves usually
rosette -forming, not matted or cushion-forming, usually much smaller than
culm leaves;
blade margins glabrous , or
ciliate only below the middle (or sometimes
papillose -
ciliate throughout in
D. portoricense ssp.
patulum , which has densely
puberulent internodes );
culms branching at the
nodes in age, 1.5-7.5 dm tall.
13 Blades progressively longer upwards on
culm , the larger 9-20 cm long and 3-8 mm wide, (15-) 20 × or more as long as wide, firm but not stiff, not longitudinally
grooved or
wrinkled ;
blade adaxial surface (weakly-) strongly
scabrous ;
sheaths glabrous ; second
glume and
sterile lemma exceeding fertile
lemma
13 Blades similar or more often decreasing in size upwards on
culm , the larger 3-16 cm long and 2-25 mm wide; if 15 × or more as long as wide, then
blades stiff and longitudinally
grooved or
wrinkled (sect.
Angustifolia );
blade adaxial surface smooth to moderately
scabrous ;
sheaths glabrous or
pilose ; second
glume and
sterile lemma shorter than to exceeding fertile
lemma .
14 Blades of mid-
culm leaves typically long and stiff,
acuminate ,
linear or narrowly
lanceolate , usually > 10× as long as wide, only 2-6 mm wide when < 8 cm long.
15 Vegetative parts
glabrous (
spikelets pubescent , lowest
internodes and
sheaths sometimes sparsely
pubescent ,
blades and
sheaths sometimes
ciliate ); mature
panicles less than ¼ as wide as long, the branches erect-
ascending , the
spikelets often subsecund;
autumnal blades 4-10 cm long,
involute , < 2 mm wide;
spikelets 1.8-2.2 mm long.
16 Larger
vernal blades 5-8 cm long and 3-5 mm wide;
vernal panicles usually less than ¼ as wide as long,
spikelets often subsecund along the strongly
ascending to erect branches;
spikelets 1.8-2.2 mm long, second
glume and
sterile lemma pubescent , summits blunt, about equaling fertile
lemma ; [of coastal wetlands and Piedmont
prairies from NC and AR south to the Caribbean and Central America]
16 Larger
vernal blades 5-9 cm long and 2-3 mm wide;
vernal panicles usually ¼ to ½ as wide as long, the branches
ascending ;
spikelets 2.3-3.0 mm long, second
glume and
sterile lemma sparsely
pubescent to
glabrous , summits pointed to blunt, exceeding fertile
lemma ; [of c. and s. FL]
15 Vegetative parts
pubescent , at least in the lower portion of plant; mature
panicles usually more than half as wide as long, the branches spreading-
ascending , the
spikelets not noticeably subsecund;
autumnal blades 1-6 cm long,
involute and < 2 mm wide in
D. aciculare and
D. arenicoloides , or to 9 cm long, flat and 2-4 mm wide in
D. angustifolium (
autumnal blade dimensions not yet known for
D. wilcoxianum );
spikelets 1.5-3.2 mm long.
17 Culms 15-35 cm long; all
culm leaf
blades long-
hirsute on both surfaces, their
sheaths densely
papillose -
hirsute ; leaf
blades of the primary branches often exceeding the immature
vernal panicles
17 Culms 15-75 cm long;
culm leaf
blades glabrous to sparsely
pubescent , or the lower softly
villous abaxially in
D. arenicoloides ; lower
sheaths usually
villous , mid-
culm and upper
sheaths glabrous (though often
ciliate basally); leaf
blades of primary branches not exceeding immature
vernal panicles .
18 Spikelets (3.0-) 3.2-3.8 mm long,
fusiform , pointed at summit,
attenuate at
base , with both
glumes attached 0.3-0.5 mm below
sterile lemma , the
autumnal spikelets 3.5-3.8 mm long; larger
vernal leaf
blades 3-6 mm wide, the lower and mid-
culm leaf
blades of similar width;
autumnal leaf
blades involute
18 Spikelets 1.8-3.1 mm long,
obovate , blunt, and the
base not
attenuate (except in
D. arenicoloides with
autumnal spikelets 2.3-3.1 mm long); larger
vernal leaf
blades 2-8 mm wide, the lower usually wider and often shorter than mid-
culm leaf
blades ;
autumnal leaf
blades involute or flat.
19 Longest
vernal blades to 16 cm, widest
vernal leaf
blades 4-8 mm, usually longitudinally
wrinkled ;
vernal and
autumnal spikelets 2.3-3.1 mm long;
autumnal leaf
blades flat, the larger to 9 cm by 2-4 mm
19 Longest
vernal leaf
blades to 6 cm (
D. aciculare ) or 12 cm (
D. arenicoloides ) long, 2-5 mm wide, not noticeably
wrinkled ;
vernal spikelets 1.5-2.8 mm long,
autumnal spikelets either 1.5-2.3 (
D. aciculare ) or 2.3-3.1 (
D. arenicoloides );
autumnal leaf
blades involute , the larger to 6 cm by 1.5 mm.
20 Longest
vernal leaf
blades to 12 cm, widest
vernal leaf
blades 3-4 (-5) mm;
vernal spikelets 2.1-2.8 mm long;
autumnal spikelets 2.3-3.1 mm long, pointed,
attenuate , the
glumes attached 0.3-0.5 mm below
sterile lemma ; first
glumes 0.7-1.5 mm long; larger
autumnal leaf
blades to 6 cm by 1.5 mm
14 Blades of mid-
culm leaves
lanceolate , thin or firm but not stiff, usually < 10× as long as wide, usually 7 mm or more wide when as much as 8 cm long.
21 Spikelet bases strongly
attenuate ,
apices blunt to beaked; first
glume (2/5-) ½-2/3 (-4/5) as long as
spikelet , attached 0.3-0.5 (-0.7) mm below expansion of second
glume and
sterile lemma
21 Spikelet bases not noticeably
attenuate ,
apices rounded to
acute ; first
glume < ½ as long as
spikelet , attached < 0.3 mm below expansion of second
glume and
sterile lemma .
22 Spikelets 2.9-3.8 mm long, broadly
elliptic , rounded at the summit, with broad and thick
nerves
22 Spikelets 2.1-2.9 mm long,
elliptic or
obovate , rounded or pointed at the summit, the
nerves often raised, but not broad and thick.
24 Culms 50-150 cm long; at least lowest
sheaths papillose -
pilose .
25 Lowest
internodes usually purple-
villous ;
nodes retrorsely (-spreading)
bearded ;
spikelets 1.8-2.2 mm long, moderately densely
pubescent ; first
glume broadly
ovate , subacute, 0.4-0.7 mm long, usually < 1.5 × as long as wide; second
glume about as long as
sterile and fertile
lemmas ; fertile
lemma 1.7-1.8 mm long
25 Lowest
internodes glabrous or
pubescent or spreading
villous , usually not purplish;
nodes glabrous , sparsely
pubescent to sparsely
bearded ;
spikelets 2.2-2.8 (-3.0) mm long, sparsely
pubescent ; first
glume 0.7-1.1 mm long, usually > 1.5 × as long as wide; second
glume shorter than
sterile and fertile
lemmas ; fertile
lemma 2.1-2.2 mm long
24 Culms 18-100 cm long;
sheaths glabrous or the lowest softly
pubescent but not
papillose -
pilose .
26 Spikelets strongly plano-
convex when viewed laterally,
obpyriform when viewed dorsally, (2.1-) 2.2-2.6 mm long; fertile
lemma and
palea papillose or smooth; first
glume usually broadened above
base , rounded or broadly tapered distally;
glume bases often reddish-purple.
27 Culm leaves 2-5 mm wide; fertile
lemma and
palea smooth, shiny
27 Culm leaves 4-12 mm wide; fertile
lemma and
palea papillose -roughened
26 Spikelets biconvex to
elliptic when viewed laterally, variously shaped but not
obpyriform when viewed dorsally; fertile
lemma and
palea smooth or
reticulate (or
papillose in D. lucidum with weak and
sprawling culms and
spikelets 1.7-2.3 mm long); first
glume usually tapered from near
base to
apex ;
glume bases various
23 Culm internodes crisp-
puberulent (sparsely so in
D. webberianum) ;
sheaths puberulent or
glabrous .
28 Spikelets elliptic , sub-
acute to pointed, greenish or faintly purple-tinged basally
28 Spikelets strongly plano-
convex when viewed laterally,
obpyriform when viewed dorsally, broadly rounded, usually markedly reddish-purple basally.
29 Fertile
lemma and
palea papillose ;
spikelets 2.2-2.6 mm long; lower
culm blades 6-12 mm wide,
glabrous
29 Fertile
lemma and
palea smooth (minutely
reticulate but not
papillose );
spikelets (1.8) 1.9-2.2 (-2.5) mm long; lower
culm blades 2-8 mm wide,
glabrous ,
glabrate , or
puberulent (especially abaxially)
1 Longer
hairs of
ligule 2-5 mm long.
2 Ligule with a distinct ring of short
hairs in front of the long
hairs .
3 Peduncle ,
panicle axis , and
sheaths puberulent with
hairs 0.1 mm long; larger
blades 3-6 cm long, 3-5 mm wide;
spikelets 1.3-1.7 mm long
3 Peduncle ,
panicle axis , and
sheaths pubescent with
hairs more than 0.1 mm long; larger
blades 12-20 cm long, 7-9 mm wide;
spikelets 1.9-2.3 mm long
2 Ligule without a distinct ring of short
hairs in front of the long
hairs .
4 Culms > 100 cm long;
node pubescence densely
retrorse ; larger
blades 12-20 cm long;
vernal spikelets 1.9-2.3 mm long, nearly 2× as long as wide;
autumnal spikelets 2.2-2.5 mm long, > 2× as long as wide
4 Culms < 100 cm long (-rarely longer);
node pubescence various; larger
blades 3-12 cm long;
spikelets (1.0-) 1.1-2.0 mm long, less than 2 × as long as wide, rounded to blunt.
5 Blades of mid-
culm leaves
lanceolate , thin or firm but not stiff, length and width various, less than 15× as long as wide
5 Blades of mid-
culm leaves
linear or narrowly
lanceolate , stiff,
acuminate , often
involute , 4-10 cm long, 2-5 mm wide, about 15-20× as long as wide
1 Longer
hairs of
ligule < 2 mm long.
6 Culm leaves basally
crowded ,
ascending , usually matted or cushion-forming, larger than the mid and upper
culm blades .
7 Sheaths conspicuously
retrorsely long-
pilose ; longer
blades 6-18 cm long and 7-12 mm wide;
spikelets (1.4-) 1.9-2.3 (-2.5) mm long
7 Sheaths variously
pubescent or
glabrous , but not conspicuously
retrorsely long-
pilose ; longer
blades 2-6 cm long and 1-8 mm wide;
spikelets 0.9-2.1 mm long.
8 Leaf
blades 1-4 mm wide,
glabrous , the
margins eciliate or basally
ciliate ;
spikelets 0.9-1.5 mm long,
glabrous ;
autumnal form branched from lower and mid
nodes as well as from
basal nodes .
9 Leaf
blades 1.5-4 (-5) cm long;
spikelets 0.9-1.2 (-1.4) mm long
9 Leaf
blades 4-12 (-20) cm long, some at least 7 cm long;
spikelets 1.2-1.5 mm long
8 Leaf
blades 2-10 mm wide,
pubescent or
glabrous , the
margins coarsely
papillose -
ciliate throughout;
spikelets 1.1-2.1 mm long,
glabrous or
pubescent ;
autumnal form branched from
basal nodes only.
10 Spikelets pubescent , 1.5-2.1 mm long;
blade surfaces
glabrous
11 Leaf
blades glabrous , or sparsely
pilose only near the
adaxial base ;
spikelets 1.4-2.1 mm long
11 Leaf
blades pilose , at least abaxially;
spikelets 1.1-1.6 mm long
6 Culm leaves not basally
crowded , the lowest leaves spreading and
rosette -forming, usually smaller than the
culm leaves.
12 Blades progressively longer upwards on
culm , the larger 9-20 cm long and 3-8 mm wide, (15-) 20 × or more as long as wide, firm but not stiff, not longitudinally
grooved or
wrinkled ;
blade adaxial surface (weakly-) strongly
scabrous ;
sheaths glabrous ; second
glume and
sterile lemma exceeding fertile
lemma
12 Blades similar or more often decreasing in size upwards on
culm ; if 15 × or more as long as wide, then larger
blades 4-10 cm long and 2-5 mm wide, stiff and longitudinally
grooved or
wrinkled (sect.
Angustifolia );
blade adaxial surface smooth to moderately
scabrous ;
sheaths glabrous or
pilose ; second
glume and
sterile lemma shorter than to equaling fertile
lemma .
13 Blades of mid-
culm leaves
linear or narrowly
lanceolate , stiff,
acuminate , often
involute , 4-10 cm long, 2-5 mm wide, about 15-20× as long as wide.
14 Ligule (1-) 1.5-2 (-2.5) mm long;
nodes bearded with spreading-
ascending hairs ;
blades moderately
hirsute ;
autumnal blades flat
13 Blades of mid-
culm leaves
lanceolate to lance-
elliptic , thin or firm but not stiff, length and width various, < 15× as long as wide.
16 Ligule 0.7-1.5 mm long; first
glume acute ;
spikelets elliptic when viewed dorsally,
biconvex or
elliptic when viewed laterally, not strongly nerved
16 Ligule < 0.5 mm long; first
glume obtuse to
truncate ;
spikelets obovate when viewed dorsally, plano-
convex when viewed laterally, strongly nerved.
17 Spikelets (1.8-) 1.9-2.2 (-2.5) mm long; first
glume 0.8-1.2 mm long; lower
culm blades 2-8 mm wide
17 Spikelets 1.5-1.8 mm long; first
glume 0.5-0.8 mm long; lower
culm blades 2-5 mm wide
15 Internodes variously hairy but not crisp-
puberulent .
18 Culms (55-) 70-140 cm long;
nodes retrorsely bearded ;
vernal cauline leaves 6-15;
blades 6-15 cm long, 8-13 mm wide,
glabrous
18 Culms < 60 cm long;
nodes not
retrorsely bearded ;
vernal cauline leaves 2-7;
blades to 14 cm long and 17 mm wide,
glabrous or
pubescent .
19 Internodes (sparsely-) moderately to densely
pubescent to
pilose ;
ligule 1-5 mm long;
blade margins either weakly
ciliate ,
papillose -
ciliate basally only, or eciliate, lacking a white-beige
cartilaginous edge 0.2 mm wide.
20 Larger mid-
culm blades 4-7 cm long, 4-7 mm wide,
glabrous to sparsely
pubescent adaxially;
ligule 1-5 mm long;
spikelets 1.1-1.5 mm long
20 Larger mid-
culm blades 3-6 cm long, 3-5 mm wide, long-
pilose adaxially;
ligule often with a ring of
hairs < 1 mm and scattered longer
hairs to 4 mm;
spikelets 1.3-1.7 mm long
19 Internodes sparsely
pilose ;
ligule < 1 mm long;
blade margins various.
21 Blades 5-14 cm by 5-17 mm, basally
truncate to
cordate ,
margins only
ciliate basally if at all, without a white-beige
margin up to 0.2 mm wide
21 Blades 1.5-6 (8) cm by 2-8 mm, basally narrowed to rounded,
blade margins either coarsely
papillose -
ciliate throughout or
glabrous with a white-beige
cartilaginous edge about 0.2 mm wide.
22 Blades with white-beige
cartilaginous margins 0.2 mm wide;
spikelets 1.4-1.7 mm long;
autumnal form branching from middle and upper
nodes
22 Blade margins coarsely
papillose -
ciliate throughout;
spikelets 1.1-2.1 mm long;
autumnal form branching from the
base .
23 Spikelets pubescent , 1.5-2.1 mm long;
blade surfaces
glabrous
24 Blades glabrous , or sparsely
pilose only near the
adaxial base ;
spikelets 1.4-1.8 mm long
24 Blades pilose ;
spikelets 1.1-1.6 mm long