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1 Leaves either 2-3-foliolate or palmately 4-11-foliolate (all the leaflets attached at a common point). | |
2 Leaves either 3-foliolate or palmately or pedately 4-11-foliolate. | |
3 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
3 Inflorescence, flower, and fruit structure various, but not with the combination of features as above. | |
7 Flowers radially symmetrical; fruit a 2-valved or 5-valved capsule; [plant of uplands or wetlands] | |
8 Leaflets notched at the tip; flowers pink, white, or yellow; [plants of uplands or temporarily flooded wetlands] | |
9 Petals 5 or more; stamens 10 or more; fruit either a legume or an aggregate of achenes or follicles | |
10 Stamens many, fused into a staminal tube; carpels 10-20, in a ring; pubescence stellate (sometimes mixed with simple hairs) | |
10 Stamens 10-many, separate, or fused but not all into a staminal tube; carpel either 1 (FABACEAE), or 3-7 in a ring (RANUNCULACEAE), or many and spirally arranged on a conical receptacle (RANUNCULACEAE or ROSACEAE) | |
12 Fruit an aggregate of achenes (borne on a fleshy, expanded receptacle in Fragaria and some Potentilla) | |
1 Leaves 1-pinnately compound (all leaflets attached to a central rachis) or more complexly compound (with several orders of branching, some leaflets at least attached to second-order branches from the rachis). | |
15 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
15 Inflorescence, flower, and fruit structure various, but not with the combination of features as above. | |
16 Flowers radially symmetrical; fruit a silique/silicle, or a schizocarp of mericarps, or an achene. | |
17 Petals 0 or 5 (if 0, the sepals petaloid); stamens 2, 4, 5, or many. | |
14 Leaves more complexly compound (with 2 or more orders of branching, some leaflets at least attached to second-order branches from the rachis). | |
19 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
19 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head subtended by bracts, but then with other features differing, such as stamens 4, or green calyx present, or petals separate, or fruit a schizocarp of mericarps, etc.). | |
21 Inflorescence various, but not an umbel; ovary superior; fruit an aggregate of follicles or an elongate capsule. | |
23 Inflorescence various, but not an umbel; fruit an aggregate of follicles or achenes, an elongate capsule, or a naked seed resembling a drupe. | |
26 Corolla bilaterally symmetrical; fruit an elongate capsule; [cultivated exotic, rarely persistent near gardens] | |
27 Leaflets 1-6 cm long, about as long as broad if > 4 cm long; pistils 4-many; fruit an aggregate of follicles or achenes; flowers mainly 4-5-merous | |
28 Pubescence of the stem and lower leaf surface glandular; flowers unisexual, on the same plant (monoecious); stamens 10; pistils 2, partly fused; fruit an aggregate of follicles | |
28 Pubescence of the stem and lower leaf surface non-glandular (or absent); flowers either bisexual (the plants hermaphroditic), or unisexual and the male and female flowers on separate plants (the plants dioecious); stamens 15 or more; pistils 1-8, separate; fruit an aggregate of follicles, a follicle, or a red or white berry. | |
29 Flowers bisexual (plants hermaphroditic); carpels 1-8 per flower; inflorescence a raceme, or a panicle of racemes with just a few branches; fruit an aggregate of follicles, a follicle, or a red or white berry | |
29 Flowers unisexual (plants dioecious); carpels 3-4 per pistillate flower; inflorescence a panicle of racemes, with numerous branches; fruit an aggregate of follicles |
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1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head subtended by bracts, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or petals separate, or fruit a schizocarp of mericarps, etc.). | |
2 Basal leaves 2-lobed, pinnately lobed, or palmately lobed (not considering cordate, hastate, or auriculate leaf bases as “lobed”). | |
4 Leaf lobing pinnate. | |
8 Corolla 2-lipped, bilaterally symmetrical or asymmetrical; stamens 2 or 4. | |
4 Leaf lobing palmate. | |
13 Petals 5; stamens 5; fruit a schizocarp of 2 mericarps. | |
16 Perianth of 5 green sepals and 5 colored petals. | |
18 Leaves 2, the single flower terminal and associated with the upper leaf; fruit an aggregate of berries | |
19 Hypanthium present, partially fused or not fused to the pistil; ovary partially inferior to superior | |
23 Leaf margins entire. | |
24 Inflorescence a terminal and/or axillary raceme, panicle, or cyme of many small flowers; fruit an achene; perianth uniseriate, of 0, 4-5, or 6 tepals. | |
24 Inflorescence either a terminal spike, or a 1-7-flowered terminal cyme, or of a solitary axillary or terminal flower; fruit various; perianth biseriate (of differentiated sepals and petals (except uniseriate, of 3 fused sepals in ARISTOLOCHIACEAE). | |
26 Flowers bilaterally symmetrical; inflorescence a terminal spike (with > 20 flowers); petals 4, usually scarious, transparent; sepals 4, green; stamens 4 | |
26 Flowers radially symmetrical; inflorescence either of a solitary flower or of a 1-7-flowered terminal cyme; petals 5, 8-12, or 0; sepals 5 (green), 3 (brown), or 5-9 (yellow); stamens 5, 12, or many. | |
27 Gynoecium either of a single pistil with 6 carpels or of a single pistil with 4 carpels or of 2 nearly separate carpels; fruit a simple capsule (or deeply 2-lobed); flowers white, brown, or greenish, either of 5 fused or distinct white petals and 5 fused or distinct green sepals, or of 3 fused brown or greenish petaloid sepals. | |
28 Flowers brown or green, of 3 fused brown or greenish petaloid sepals (and 0 petals); carpels 6; stamens 12; leaves 4-10 cm wide | |
29 Petals fused; sepals fused; carpels 2; plant pubescent. | |
32 Carpels many, separate, spiral; petals yellow or white. | |
33 Flowers lacking a hypanthium; fruit an aggregate of achenes or aggregate of follicles | |
34 Flowers radially symmetrical; inflorescence an umbel (or composite of umbelliform units, or a terminal panicle. | |
36 Inflorescence a terminal raceme; perianth of 4 green sepals and 4 white petals; fruit a silique/silicle; fresh foliage in spring and summer with a strong garlic odor; larger leaves < 10 cm in diameter | |
35 Ovary inferior; inflorescence an umbel (or a composite of umbellate units); fruit a schizocarp of mericarps. | |
39 Leaves tubular, with a sutured ventral flange, erect or reclining, adapted as a pitfall for insects (flat, phyllodial leaves sometimes present as well, common in the winter in some species, such as S. oreophila) | |
39 Leaves flat, not sutured into a tubular shape. | |
43 Corolla white, lavender, or blue, 2-lipped and bilaterally symmetrical. | |
40 Stem leaves alternate; perianth radially symmetrical (less commonly bilaterally symmetrical); stamens 5, 6-8, 9, 10 (rarely 4). | |
49 Corolla bilaterally symmetrical (barely so in Limosella in SCROPHULARIACEAE); stamens 2, 4, 6, 8, or 10. | |
49 Corolla radially symmetrical; stamens 5, 10, 4-6, or 9. | |
56 Perianth of green sepals and more brightly colored petals; stamens 5 or 10; carpels 2, 3, 4, or 5. | |
57 Leaves lacking sticky gland-tipped hairs. | |
59 Inflorescence of a solitary, terminal flower; carpels 2-3 (-4). | |
60 Corolla of separate petals or united only for a short length (< ¼ its length). | |
59 Inflorescence of several to many flowers; carpels 5 (3 in Galax in DIAPENSIACEAE). | |
64 Inflorescence a somewhat to very diffuse panicle, with 3 or more orders of branching, not giving at all the impression that the overall inflorescence is made of racemose units. | |
66 Inflorescence of 1-several terminal and axillary racemes, the plant typically well-branched, especially from the base; stamens 5 | |
66 Inflorescence of a single, terminal raceme, the plant unbranched; stamens 10 (or 5, with 5 staminodes) | |
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2 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
2 Inflorescence, flower, and fruit structure various, but not with the combination of features as above. {add under 2b [Cynosciadium] APIACEAE} | |
7 Leaflets radially arranged at the summit of the petiole, not differentiated in size or placement into a terminal leaflet and 2 lateral leaflets; leaflets prominently notched at the apex; petals 5, yellow; inflorescence axillary, cymose or umbelliform; carpels 5 | |
1 Leaves either 1-pinnately compound (all leaflets attached to a central rachis) or more complexly compound (with several orders of branching, some leaflets at least attached to second-order branches from the rachis). | |
10 Inflorescence an involucrate head subtended by phyllaries, heads solitary or many, variously arrayed in secondary inflorescences; fruit a cypsela; ovary inferior | |
12 Flowers bilaterally symmetrical, papilionaceous; fruit a legume; leaves even-pinnately or odd-pinnately compound, the terminal leaflet sometimes replaced by a tendril; leaflets entire or at most minutely denticulate | |
12 Flowers radially symmetrical (or barely bilaterally symmetrical in Erodium in GERANIACEAE); fruit a capsule, capsular but of 5 mericarps, or an aggregate of achenes, nutlets, or follicles (in some cases the # of pistils from many down to 2 or even 1); leaves odd-pinnately compound, never with tendrils; leaflets serrate (or entire to shallowly lobed in Polemonium in POLEMONIACEAE, Cardamine in BRASSICACEAE, and Floerkea in LIMNANTHACEAE). | |
13 Pistils many (only 1-2 in Agrimonia, Poteridium, Poterium, and Sanguisorba); fruit an aggregate of achenes, nutlets, or follicles; hypanthium present; stamens 5-many (only 4 in Poteridium and Sanguisorba) | |
13 Pistil 1 (or deeply 2-3-lobed in Floerkea in LIMNANTHACEAE); fruit a silique, capsule, schizocarp of 2-3 mericarps, or a capsular schizocarp of 5 mericarps (Erodium in GERANIACEAE); hypanthium absent; stamens 3-6. | |
15 Petals 5, fused (distinct in Erodium in GERANIACEAE); stamens 5; inflorescence axillary or terminal, cymose, consisting of subcapitate, umbel-like, or helicoid cymes; fruit either a capsule, or a capsular schizocarp of 5 mericarps (Erodium in GERANIACEAE). | |
16 Flowers slightly bilaterally symmetrical (2 of the petals of different size than the other 3); fruit a capsular schizocarp of 5 mericarps; carpels 5 | |
17 Fruit a capsule; fresh plant not aromatic. | |
11 Leaves more complexly compound (with 2 or more orders of branching, some leaflets at least attached to second-order branches from the rachis). | |
19 Leaves either 2× odd-pinnate or more complexly 2-4× ternately or ternately-pinnately compound; flowers in various inflorescences; fruits various (not legumes or loments). | |
20 Leaf segments linear, less than 2 mm wide. | |
21 Inflorescence either a terminal solitary flower or terminal raceme or panicle; ovary superior, either of 2 fused carpels or of 1-5or many distinct 1-carpellate pistils; fruit either a capsule or an aggregate of follicles or achenes. | |
23 Erect or sprawling herb; stamens 5-6 or >15. | |
25 Inflorescence an umbel; ovary inferior, of 2 fused carpels; fruit a schizocarp of 2 mericarps; inflorescence an umbel | |
25 Inflorescence a panicle or raceme; ovary superior, of 1-8 carpels; fruit an aggregate of follicles, a single follicle, or an indehiscent berry-like fruit. | |
26 Flowers bisexual; carpels 1-8; fruit an aggregate of follicles, a single follicle, or an indehiscent berry-like fruit | |
26 Flowers unisexual; carpels (in pistillate flowers) of 3 (-5) carpels; fruit an aggregate of follicles | |
27 Inflorescence a raceme, panicle, or cyme; ovary superior, of either 1-2 fused carpels or of many separate 1-carpellate pistils. | |
29 Perianth of 4-5 whorls of 3 parts each (some of the whorls modified into nectaries); pistil 1, 1-carpellate; fruit a drupelike, blue, naked seed; largest leaflets > 6 cm long, obviously longer than wide | |
29 Perianth of 1 whorl; of 4-5 parts; pistils many, each 1-carpellate; fruit an aggregate of achenes or an aggregate of follicles; largest leaflets either < 6 cm long, or if longer than 6 cm, also about as wide as long |
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1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.). | |
2 Perianth uniseriate (represented only by undifferentiated tepals or sepals) or completely absent; flowers usually unisexual, less commonly bisexual). | |
3 Inflorescence a cyathium, consisting of a single pistillate flower (reduced to a single 3-carpellate pistil) and 2 or more staminate flowers (each reduced to 1 stamen), borne in a cup-like involucre, the involucre bearing pointed or rounded glands, these sometimes brightly colored and petaloid, mimicking an individual flower (the cyathia then secondarily arranged in terminal cymes, or solitary and axillary, etc.); fresh plants with milky juice; fruit a 3-lobed, 3-locular capsule | |
3 Inflorescence not a cyathium (and staminate or bisexual flowers with > 1 stamen; fresh plants lacking milky juice (except Stillingia in EUPHORBIACEAE); fruit various, not as above. | |
7 Leaf bases cordate to rounded. | |
8 Styles 3; fruit a 3-lobed, 3-carpellate capsule (1 carpel sometimes aborting); inflorescence either a terminal or leaf opposed raceme, or a dense axillary condensed cyme with conspicuous toothed bracts subtending the flowers | |
4 Leaf margins entire. | |
11 Leaf base cuneate, rounded, or truncate; calyx of 3-4-5 distinct sepals, radially symmetrical, white or yellow; fruit a dry, nutlike drupe or an achene. | |
14 Inflorescence a leaf-opposed (sometimes apparently terminal) spike or raceme; flowers visually white from white petaloid sepals, white bracts, or white stamens. | |
15 Sepals absent; carpels 3-4; stamens 2-6 (-8); fruit a capsule, a 1-seeded drupe, or a schizocarp of 3-4 mericarps; leaf bases cordate or subcordate; [Basal Angiosperms]. | |
14 Inflorescence not leaf-opposed, either simpler (single axillary or glomerules of flowers) or more complexly branched (axillary or terminal panicles or complex cymes); flowers white, reddish, scarious, or greenish. | |
24 Petals connate. | |
25 Inflorescence various, either a terminal head, or axillary and solitary, or variously axillary or terminal and more diffuse. | |
28 Petals 4-7; stamens 1× or 2× as many as the petals, 4-7, 8, 10, 12, or14; leaves herbaceous in texture | |
30 Petals connate (at least basally), 4, 5, 6, 7, or 8; carpels 1, 2, 4, 5, or 6 (rarely 3 in Reseda in RESEDACEAE); fruit a legume or 1-, 2-, or 5-loculed capsule (except a 1-seeded indehiscent pod in Krameria in KRAMERIACEAE). | |
31 Stamens 6-10 (-25), more than the number (4 or 5) of petals and the number (4 or 5) of the sepals; fruit a legume or a 1-6-carpellate capsule. | |
35 Pistil 5-carpellate; capsule 5-locular, explosively dehiscent; inflorescence axillary, small clusters of flowers | |
35 Pistil 2-carpellate; capsule 2 locular, opening gradually or not at all; inflorescence a terminal spike, raceme or panicle (or solitary, axillary flowers in Chaenorrhinum in PLANTAGINACEAE and Krameria in KRAMERIACEAE). | |
36 Fruit 1-seeded, indehiscent; sepals petaloid, red-purple; petals dimorphic, the upper 3 long-clawed, the lower 2 small, thickened, and glandlike | |
37 Stamens 5; corolla not spurred; capsule septicidal; pubescence of the stem and leaves either gland-tipped or dendritically branched | |
37 Stamens 4; corolla with a distinct spur or sac at the base between the the 2 lower calyx lobes (except not spurred in Digitalis and Schwalbea); capsule loculicidal (only at the summit in Antirrhinum and Chaenorrhinum, and septicidal in Schwalbea); pubescence of the stem and leaves neither gland-tipped (except in Antirrhinum and Chaenorrhinum) nor dendritically branched. | |
40 Pistils 4-10 (each 1-carpellate) in a ring, these sometimes fused basally, each with its own style/stigma; fruit either an aggregate of achenes or follicles or a 5 (-7) locular capsule. | |
42 Fruit an aggregate of follicles; leaves fleshy in texture; inflorescence; leaves entire of sparsely and coarsely serrate, with < 12 points per leaf; [plants primarily of dry habitats] | |
44 Petals 5 (rarely 4 or 6); sepals 5 (rarely 4 or 6); stamens 5 (or multiples of 5), 6, or 12; fruit a capsule or a ring of mericarps. | |
47 Flowers 6-merous (the petals and sepals 6, the stamens 6 or 12); corolla pink or purplish (rarely white); fruit a septicidal capsule | |
47 Flowers 5-merous (the petals and sepals 5, stamens 5 or various multiples of 5); corolla yellow, reddish, or blue; fruit a loculicidal or septicidal capsule. | |
48 Stamens 5; corolla yellow or blue; capsule either 10-locular and septicidal or 1-locular (with 3 carpels) and loculicidal. | |
48 Stamens (5-) 10, 15, 20, 30 (-many); corolla white, pink, yellow, or reddish; capsule 3-, 5- (-10)-locular, loculicidal. | |
50 Stamens (5-) 10, 15, 20, 30 (-many); corolla yellow or reddish; capsule 3 (-10)-locular, loculicidal | |
51 Pistils 2, united only by the style and stigma; fruit a schizocarp of 2 follicles (often single by abortion); plant with milky juice when fresh; leaves entire; inflorescence an umbel | |
55 Plant an herb, erect or sprawling; leaves > 1.5 cm long. | |
61 Flowers sessile or very-short pedicelled, solitary in the leaf axils. | |
61 Flowers either solitary and obviously pedicelled, or several in an axillary or lateral inflorescence. | |
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1 Inflorescence an involucrate head subtended by phyllaries, the heads solitary or many and variously arrayed in secondary inflorescences, the ovary inferior, the corolla connate and tubular at least basally, the calyx absent, the stamens 5, the fruit a cypsela | |
1 Inflorescence, flower, and fruit structure various, but not with the combination of features as above (sometimes the flowers in a head, e.g. Eryngium in APIACEAE, but then with other features differing, such as stamens 4, or green calyx present, or fruit a schizocarp of mericarps, etc.). | |
2 Perianth uniseriate, with only undifferentiated tepals; flowers many and small, greenish or brownish, inconspicuous individually; inflorescence of glomerules that are usually further aggregated into racemes or panicles; fruit an achene or utricle | |
2 Perianth biseriate, both sepals and petals present and differentiated (except uniseriate and of 2 white to cream-colored sepals 5-10 mm long in Macleaya in PAPAVERACEAE); flowers larger, usually with the petals prominently colored; inflorescence various, but not as above; fruit a capsule, silique/silicle, or a schizocarp of 2 mericarps. | |
3 Corolla radially symmetrical and either connate into a tube or distinct (except Reseda in RESEDACEAE, with bilateral symmetry but separate petals); stamens 5 or more; fruit a silique/silicle, a schizocarp of 4 mericarps, or a 1-, 3-, or 4-locular capsule (2-locular in Ipomoea in CONVOLVULACEAE and Glaucium in PAPAVERACEAE), opening variously. | |
7 Corolla radially symmetrical; stamens 5 or 6. | |
9 Plant an erect or sprawling herb. | |