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Key to Poaceae
2 Plant a small to medium grass, culms < 2 m tall and usually also < 5 mm in diameter at the base. | |
5 Spikelets 2-flowered, often dorsally compressed, falling entire at maturity (the abscission below the glumes), the upper floret usually bisexual, the lower one male or sterile. | |
6 Glumes often as long as or longer than the lemmas and concealing the florets; spikelets usually arranged in obvious pairs or triplets, with 1 spikelet sessile or shortly pedicellate and the other 1 (or 2) spikelets pedicellate (the pedicellate sometimes vestigial or absent) | |
5 Spikelets 1-, 2-, or many-flowered, usually terete or somewhat laterally compressed, either abscising at maturity above the glumes or if 2-flowered then both florets bisexual, or the upper sterile. | |
10 Glumes present; palea various; [habitats various]. | |
Key to Poaceae, Key A: bamboo grasses (woody trees and shrub grasses) (tribe Bambuseae)
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1 Rhizomes pachymorph, having root-bearing internodes thicker than the culm; culms usually in single clumps | |
2 Mid-culm branches consistently 2, unequal, rarely with a smaller central third branch | |
2 Mid-culm branches not consistently 2, initially 1-9. | |
3 Dwarf bamboos, < 1 (-1.5) m tall. | |
4 Leaf blade margins not or only slightly bleached in winter, terminal blade parallel to shoot axis, blades often variegated | |
4 Leaf blade margins more or less bleached in winter, terminal blade often angled from shoot axis, blades usually not variegated | |
5 Mid-culm branches initially 1-3 (-5). | |
Key to Poaceae, Key B: robust herbaceous grasses
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1 Inflorescence an array of spikes, the spikelets closely imbricate in 2 rows along the rachis of the spikes, the spikes alternate along the primary inflorescence axis; [tribe Zoysieae; subtribe Sporobolinae] | |
1 Inflorescence otherwise, either the spikelets embedded or in grooves in an thickened rachis, or the inflorescence a slender or broad panicle. | |
2 Spikelets embedded in the thickened rachis (the inflorescence thus like an ear of corn), or fitting into grooves in the thickened rachis (the inflorescence thus cylindrical and resembling a rat's tail). Spikelets unisexual, with male and female spikelets in separate inflorescences or in different parts of the same inflorescence; [tribe Andropogoneae]. | |
2 Spikelets not embedded or fitting into grooves in the rachis, the inflorescence a slender or broad panicle (the spikelets visibly separate and often pedicelled). | |
4 Spikelets with a single floret, this unisexual (either pistillate or staminate); plants with aerenchymatous culms, [plants of seasonally or tidally flooded wetlands]; [tribe Oryzeae]. | |
5 Pistillate spikelets on the upper branches of the panicle, staminate spikelets on the lower branches; lemmas and paleas clasping along their margins; plants annual | |
5 Pistillate and staminate spikelets intermingled on the same branches of the inflorescence; lemma margins free; plants perennial | |
4 Spikelets with 2 or more florets, at least some of these bisexual; plants without aerenchymatous culms, [plants of uplands or temporarily to seasonally flooded wetlands]. | |
6 Spikelets with 2-8 florets, these bisexual. | |
7 Leaves cauline; plants short to long rhizomatous, usually somewhat clumped (short rhizomatous to cespitose in Neyraudia). | |
9 Lemmas 3-7 veined, the apices not bifid (instead either unawned, entire, or only minutely awned); plants rhizomatous; [tribe Arundineae] | |
10 Lemmas pilose; rachilla glabrous; plants short-rhizomatous (somewhat clumped); culms to 10 m tall; [plants of uplands or saturated or temporarily flooded wetlands] | |
10 Lemmas glabrous; rachilla sericeous; plants long rhizomatous; culms to 4 m tall; [plants of uplands or saturated, tidally flooded, or seasonally flooded wetlands] | |
11 Spikelets falling separately, not attached to rachis segments, stalks, or bristles; spikelets <2× as long as wide; [tribe Paniceae]. | |
11 Spikelets falling with attached rachis segments, stalks, or bristles; spikelets >3× as long as wide; [tribe Andropogoneae]. | |
19 Spikelets unawned or with awns to 5.2 mm long; anthers 3; [exotic, rarely encountered] | |
Key to Poaceae, Key C: bur, bead, or bulblet grasses of various tribes
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1 Fertile spikelets variously spiny or bead-like. | |
2 Pistillate spikelets concealed within a hard, beadlike shell, this white, black, or variously colored; [tribe Andropogoneae] | |
3 Bur formed from accrescent branchlets, with fewer and less regularly arranged straight prickles (these typically retrorsely scabrous); [tribe Paniceae] | |
Key to Poaceae, Key D: rattail or cob grasses
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1 All spikelets unisexual, the pistillate and staminate spikelets either in separate inflorescences, or the pistillate spikelets below the staminate spikelets in the same inflorescence; leaves 9-120 mm wide; [tribe Andropogoneae]. | |
2 Pistillate and staminate spikelets in separate inflorescences, the pistillate inflorescences axillary, staminate inflorescences terminal | |
1 Some spikelets bisexual; leaves 1-25 mm wide. | |
5 Plants obviously and prominently rhizomatous (Eremochloa) or stoloniferous (Stenotaphrum); lower glume with pectinate margins (Eremochloa) or irregularly toothed (Stenotaphrum) | |
6 Plant rhizomatous; lower glume with pectinate margins; [commonly naturalized turf grass and roadside weed]; [tribe Andropogoneae] | |
6 Plant stoloniferous; lower glume irregularly toothed; [naturalized turf grass and allegedly also native]; [tribe Paniceae] | |
5 Plants annual; lower glume with smooth and cartilaginous margins (or absent); [rare waifs]; [tribe Poeae]. | |
8 Lower glumes of the sessile spikelets rough, rugose, pitted, tuberculate or alveolate between the keels | |
Key to Poaceae, Key E: grasses of tribe Andropogoneae (also including grasses also keyed in Keys B, C, and D)
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1 Leaves ovate-lanceolate, 2-10 cm long, 2.5-7× as long as wide; plants weak-stemmed annuals, branching, decumbent, rooting at the lower nodes; [exotic weeds]. | |
1 Leaves lanceolate to linear, either longer or proportionately narrower; plants either perennial or coarse annuals with erect and mostly unbranched culms. | |
3 Spikelets embedded in the thickened rachis (the inflorescence thus like an ear of corn), or fitting into grooves in the thickened rachis (the inflorescence thus cylindrical and resembling a rat's tail), or the pistillate inflorescences enclosed in a hard, bead-like, pearly-white, modified bract. | |
4 Spikelets unisexual, with male and female spikelets in separate inflorescences or in different parts of the same inflorescence. | |
4 Spikelets, or at least one of each pair, bisexual. | |
7 Culms 2-45 cm tall; leaves 1-5 mm wide; plants perennial, obviously and prominently rhizomatous; lower glume with pectinate margins; [commonly naturalized turf grass and roadside weed] | |
7 Culms 30-400 cm tall; leaves 2-25 mm wide; plants annual or perennial, cespitose or short-rhizomatous; lower glume winged or not, but not pectinate. | |
8 Lower glumes of the sessile spikelets rough, rugose, pitted, tuberculate or alveolate between the keels | |
14 Spikelets unawned or with awns to 5.2 mm long; anthers 3; [exotic, rarely encountered] | |
16 Spikelets awned, the awn 10-20 cm long. | |
19 Inflorescence a false panicle with numerous inflorescence units; peduncles with 2 rames in digitate clusters (1 almost sessile, the other rame stalked and both bearing 7-14 heterogamous spikelet pairs), disarticulation in the rames beneath the bisexual spikelets; spikelets a mix of heterogamous and homogamous pairs; awns of the lemma 1-3.5 cm long; [non-natives] | |
19 Inflorescences true panicles (with evident rachises); disarticulation beneath the sessile spikelets (or not occurring at all); spikelets only heterogamous; awns of the lemma (5-) 10-40 mm long (when present); [native and non-natives]. | |
20 Pedicelled spikelet represented by pedicel only; apex of sheath bearing 2 auricles 1-10 mm long; awns of the lemmas once or twice-geniculate; [native] | |
24 Rame internodes and pedicels with a translucent medial groove; pedicelled spikelets smaller than the sessile spikelets (or vestigial/absent) | |
24 Rame internodes and pedicels without a translucent medial groove; pedicelled spikelets ca. the same size as sessile spikelets |
Key to Poaceae, Key F: grasses of tribe Paniceae (also including grasses keyed as well in Keys B, C, D, and H)
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2 Spikelets (at least the terminal) subtended by 1-many stiff, terete bristles. | |
6 Plants not as above (if robust, lacking shiny black florets and not arching or climbing); [collectively widespread, including s. FL]. | |
8 Leaves < 8× as long as wide; ligules present, membranous or of hairs. | |
11 Subterranean (cleistogamous) inflorescences present; aerial inflorescences with elongate rachises; spikelets of the aerial inflorescences often sterile; spikelets glabrous; leaves either with a white cartilaginous margin or prominently ciliate; [of the Coastal Plain] | |
11 Subterranean inflorescences absent; aerial inflorescences either with digitate or subdigitate branches and glabrous spikelets, or with elongate rachises and conspicuously pubescent spikelets; spikelets of the aerial inflorescences fertile; spikelets glabrous, ciliate, or pubescent; leaves various (often not as above); [collectively widespread] | |
13 Spikelets not subtended by a cuplike callus. | |
14 Leaves primarily lacking ligules (at least the upper, and often all, leaves without ligules, if vestigial ligules present, these of hairs) | |
14 Leaves with ligules, these either membranous or of hairs. | |
16 Inflorescence of 1-sided, spikelike primary branches. | |
17 Spikelets with lower lemmas (and lower glumes, if present) adjacent to the branch axes. | |
16 Inflorescence either paniculate with well-developed secondary branchlets or if the primary branches spikelike, then the spikelets not borne in a 1-sided arrangement on the spicate branches. | |
21 Inflorescences open panicles, or if narrowed, all or nearly all the panicle branches readily visible. | |
23 Plant developing a terminal (“spring”) inflorescence usually before mid-summer, followed by lateral (“autumnal”) inflorescences from lower, mid, and/or upper nodes, these often included or hidden among the fascicles of smaller “autumnal” leaves; often developing a rosette of overwintering basal leaves | |
23 Plant developing a terminal inflorescence usually after mid-summer, the lateral inflorescences, when present, from the upper nodes, usually appearing at the same time as the terminal panicle, and not hidden by dense fascicles of smaller leaves; plants lacking a rosette of overwintering basal leaves. | |
24 Spikelets smooth, not tuberculate. | |
25 Panicle < 2 cm wide at maturity. | |
28 Panicles constricted, 0.3-1.6 cm wide; spikelets subsessile to short-pediceled; summit of fertile palea not enclosed by fertile lemma | |
25 Panicle > 2 cm wide at maturity. | |
29 Fertile lemmas not rugose. | |
32 Plants with rhizomes; fertile lemma 1.6-4 mm long. | |
33 Rhizomes about 1 cm thick with pubescent scale-like leaves; lower portion of culm hard, nearly woody | |
35 Culms slightly compressed below; ligules 0.5 mm long or less; spikelet pedicels appressed, the spikelets subsecund, usually some obliquely bent above the first glume; fertile lemma 1.8-2.2 mm long | |
Key to Poaceae, Key G: wheat grasses of tribe Triticeae (and a few unrelated mimics)
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3 Spikelets sessile. | |
4 Spikelets (not including the awns) < 3× as long as the rachis internodes. | |
5 Glumes subulate to narrowly lanceolate or linear, narrowing from below midlength, with 1 vein throughout or at least distally (sometimes 1-3 veins at midlength in Pascopyrum); [tribe Triticeae] | |
6 Awns short, 5 mm or less in length; glumes not keeled; spikelets with 2-12 florets | |
Key to Poaceae, Key H: finger grasses
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1 Spikelets 2-flowered, often dorsally compressed, falling entire at maturity (the abscission below the glumes), the upper floret usually bisexual, the lower one male or sterile; [tribe Paniceae] | |
3 Spikelets with lower lemmas (and lower glumes, if present) adjacent to the branch axes. | |
1 Spikelets 1-, 2-, or many-flowered, usually terete or somewhat laterally compressed, either abscising at maturing above the glumes or if 2-flowered then both florets bisexual, or the upper sterile; [tribes Cynodonteae and Zoysieae] | |
7 Spikes normally 2 or more, alternate, digitate, subdigitate, or verticillate along the main inflorescence axis; second glume lacking a recurved spine; fresh plants not aromatic with a citrus odor. | |
8 Spikes arranged along the central inflorescence axis alternately, solitary at each node or inflorescences consisting of a solitary spikelike raceme (Zoysia). | |
9 Spikelets with 1 bisexual floret, sometimes also accompanied by modified male, sterile, or rudimentary florets above the fertile floret. | |
10 Spikelets with modified male, sterile, or rudimentary florets above the fertile floret; [plants of uplands]; [tribe Cynodonteae; subtribe Boutelouinae] | |
11 Spikelets with 1 solitary fertile floret; inflorescence of solitary spike-like racemes; [waifs or uncommon non-native turf grasses of disturbed areas]; [tribe Zoysieae; subtribe Zoysiinae] | |
9 Spikelets with 2 or more bisexual florets (sometimes also with additional reduced florets); [tribe Cynodonteae; subtribe Eleusininae]. | |
8 Spikes arranged along the central inflorescence axis in a digitate, subdigitate, or verticillate manner, all or most nodes with 2 or more spikes; [tribe Cynodonteae]. | |
15 Spikelets with 1 or more sterile florets. | |
17 Spikes to 7 cm long, terminating in a point (the spikes acuminate); [tribe Cynodonteae; subtribe “incertae sedis”] | |
17 Spikes to 22 cm long, terminating in a functional or rudimentary spikelet (the spikes acute to obtuse); [tribe Cynodonteae; subtribe Eleusininae]. | |
Key to Poaceae, Key I: rice grasses, of tribe Oryzeae (also including grasses keyed as well in Key B)
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1 Lemma margins free; plants perennial. | |
1 Lemmas and paleas clasping along their margins; plants annual or perennial. | |
3 Spikelets either pistillate or staminate, the upper branches of the panicle with pistillate spikelets, the lower branches with staminate spikelets; grains terete | |
3 Spikelets bisexual; grains laterally flattened. | |
Key to Poaceae, Key J: dense spike grasses
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1 Lemma 0.5-6 mm long. | |
3 Glume awns not hairy | |
Key to Poaceae, Key K: grasses with 1 floret and unawned lemmas
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2 Spikelets variously shaped, sometimes rounded but not clearly orbiculate, the glumes keeled or not. | |
3 Florets soft, papery; glumes and lemmas keeled in ×-section; spikelets 0.7-10.8 mm long. | |
4 Florets with a conspicuous tuft of hairs on the callus; [tribe Zoysieae; subtribe Sporobolinae] | |
4 Florets not conspicuously hairy on the callus. | |
5 Lemma 1-veined; ligule of hairs; grain becoming mucilaginous when wet; [tribe Zoysieae; subtribe Sporobolinae] | |
5 Lemma 1-5-veined; ligule a membrane (the summit sometimes ciliolate); grain not becoming mucilaginous when wet. | |
Key to Poaceae, Key L: grasses with 1 floret and awned lemmas
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3 Lemma hardened, distinctly different than the glumes in texture when mature; [tribe Stipeae]. | |
4 Rhizomatous perennials; primary leaves cauline (the basal leaves < 2 cm long or merely represented by sheaths), 8-16 mm wide | |
4 Cespitose perennials; primary leaves basally disposed, 0.2-10 mm wide. | |
3 Lemma neither hardened nor distinctly different than the glumes in texture when mature. | |
15 Inflorescences with disarticulating branches;
Lower glumes 0.9-4 mm long; spikelets disarticulating below the glumes, the spikelet falling as a whole; spikelets appressed to divergent from the raceme axes; sheaths not strongly overlapping; [tribe Cynodonteae; subtribe Eleusininae] | |
15 Inflorescences without disarticulating branches; lower glumes 1.5-3 mm long (Muhlenbergia paniculata) or (2-) 3.5-7 mm long (Gymnopogon); spikelets disarticulating above the glumes (which often remain on the inflorescence); spikelets strongly appressed to the raceme axes; sheaths strongly overlapping or not , if so then at least on the upper culm, therefore hiding the culm; [tribe Cynodonteae; subtribe “incertae sedis”] | |
16 Blades with thick, white margins and well-developed midrib; inflorescence branches strongly divergent; glumes unequal, usually shorter (sometimes only slightly so) than the solitary floret | |
16 Blades lacking both thick, white margins and well-developed midribs; glumes subequal, usually exceeding the 1-2(4) floret(s) | |
14 Spikelets pedicellate and arrayed in a more complex, open (and sometimes loosely contracted) panicle. | |
19 Lemma awns (when present) 0.2-1.5 mm long; culms to 200 cm tall; plants cespitose or rhizomatous perennials | |
18 Spikelets disarticulated above the glumes, the floret falling, leaving the glumes attached to the pedicels. | |
Key to Poaceae, Key M: grasses with 2+ florets, these exceeded and usually concealed by the glumes
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1 Spikelets disarticulating below the glumes, the spikelets falling as a whole or in clusters; [tribe Poeae] | |
2 Spikelets monomorphic. | |
4 Fertile floret basal, with 1-several sterile florets towards the tip of the spikelet; [tribe Cynodonteae; subtribe “incertae sedis”] | |
12 Awns of the lemmas 2-8 mm long, therefore conspicuous and not merely a minute midvein extension, bent or geniculate and often basally-twisted and exceeding lemma apices; lemma apices bifid or bicuspidate. | |
13 Panicle branches densely pubescent, usually glistening yellowish-brown or yellowish to tan; panicles either 1-5 cm (Trisetum aureum) OR 5-20 cm long (Trisetum flavescens); leaves evenly distributed; ligules 0.5-1(2) mm, obtuse, lacerate, sometimes ciliolate; plants sometimes with rhizomes to 7 cm long (T. flavescens); [occurring very sporadically in disturbed sites throughout our area or as a ballast waif in NJ] | |
13 Panicles branches glabrous or sparsely pubescent, usually silvery-shiny; panicles (5)20-30(50) cm long; leaves mostly basal or evenly distributed; ligules 0.5-4 mm, truncate or rounded; plants not rhizomatous; [a native of cool, moist, undisturbed northern sites; rare and disjunct in high elevation areas s. to VA] | |
14 Awns straight or geniculate (sometimes only slightly so), if geniculate then the awn not bearing a distinct ring at the joint and not bearing a clavate distal segment; [collectively widespread]. | |
Key to Poaceae, Key N: grasses with 2+ florets, these readily visible by extending past the glumes
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1 Plants dioecious; plants strongly rhizomatous-clonal; [plants of saline situations, coastal or more rarely inland]; [tribe Cynodonteae; subtribe Monanthochloinae] | |
1 Plants bisexual; plants cespitose or weakly short- rhizomatous; [plants of various habitats, including saline]. | |
5 Mature spikelets not stiff; grains neither protruding, nor shaped with a bottleneck-like beak. | |
6 Inflorescence branches usually strongly ascendant (sometimes weakly spreading); culms often disarticulating at the first node; [tribe Arundineae] | |
6 Inflorescence branches ascending, widely divergent, or nodding; culms not disarticulating at the first node; [collectively widespread, dry to wet sites] | |
7 Florets 3-34 per spikelet; lemmas unawned; [tribe Eragrostideae; subtribe Eragrostidinae] | |
14 Ligule membranous (the membrane apex ciliate); lower 1-4 florets sterile; disarticulation above the glumes and between the florets; [of various, usually moist, habitats, collectively widespread]; [tribe Centotheceae] | |
13 Lowermost florets of the spikelet fertile; [tribe Poeae]. | |
16 Lemmas rounded at the apex, not awned. | |
17 Inflorescence pedicels to 0.5 mm thick (if to 0.8 mm as in Sclerochloa dura then culms usually prostrate and leaves also exceeding inflorescence length); panicle branches flexible; lemmas hardened or membranous but not coriaceous at maturity. | |
18 Inflorescences usually exceeded by the leaves (or sometimes similar in length); culms usually prostrate or procumbent (occasionally ascending); plants often compact and low-growing; mature lemmas hardened | |
18 Inflorescences exceeding the leaves; culms usually erect; plants compact or not; mature lemmas membranous | |