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4Rayflorets mostly in 2-3 series, blades 1.3-4.2 mm, often not exceeding mature pappus (S. tenuifolium and S. bracei in 1 series to 8.5 mm); plants appearing entirely glabrous (leaf axils or margins sometimes inconspicuously strigillose); plants either taprooted annuals or perennials with succulent, linear, entire leaves; [coastal wetlands, often saline, or disturbed areas inland]; [subgenus Astropolium]
(c) Wright, Janet - CC-BY-NC, permission granted to NCBG
5 Leaves not succulent, usually broader or toothed; raylaminae 1.3-4.2 mm, usually in 2-3 series (laminae 3.7-7 mm and in 1 series in S. divaricatum); plants annual, often copiously branched, heads (10-) 30-100 (-150+).
9 Free tips of phyllaries flat, tipped with a white spine (otherwise acute); stem leaves crowded (internodes mostly <1 cm), linear or oblong, 1-4 mm wide, usually upcurved; [subgenus Virgulus, section Ericoidei]
8Phyllary tips acute to acuminate, sometimes mucronate or apiculate (e.g., S. puniceum), flat (sometimes spreading at tips when foliaceous); flowers pink, purple, or lavender (rarely white in S. firmum).
2Phyllarieslinear, in 4-7 series; disc flowers 4.0-5.5 mm long; [dry, rocky soils of open woodlands and prairies; IL and n. MO south through MO and se. KS to s. AR and se. OK]
2Phyllaries broadly lanceolate, foliaceous, in 3-5 series; disc flowers 5.8-6.8 mm long; [moist forests of the higher-elevation Southern Appalachians; w. NC and e. TN south to nw. SC and n. GA]
4 Heads at first at ends of long, bracteate branches, then produced and maturing as axillary and nearly sessile or on very short lateral branches, commonly on one side of the main stem and appearing secund to subsecund, in paniculiform arrangements; ray flowers in 2-3 series, corollas mostly 2-3.5 (-4) mm long, the blades 0.2-0.4 mm wide (dry), blue to purple, coiling back in 2-3 (-4) coils; disc flowers 11-23; [e. GA s. to s. FL, w. to LA]
4 Heads often at ends of long, bracteate branches, axillary heads usually maturing on elongate lateral branches, the whole arrangement often diffusely paniculiform to pyramidal-paniculiform, or heads more distally disposed and the arrangement corymbiform to thyrsiform; ray flowers in 1 series, corollas mostly 4-7 mm long, the blades 0.4-0.8 mm wide (dry), blue to white, coiling back in 3-5 coils; disc flowers (20-) 33-45 (-50); [sc. U.S. east to AL and scattered eastward as an introduction]
1 Mid and upper stem leaves > 8× as long as wide; phyllaries tipped with a small, white spine; rays white (to pale pink); involucre 2.5-4.5 (-5) mm high; disc flowers 6-12 (-20) per head; [section Ericoidi]
1 Mid and upper stem leaves 2-7× as long as wide; phyllaries not spine-tipped; rays purple, lavender, rose, blue (rarely nearly white); involucre > 5 mm high (except sometimes as short as 4 mm high in S. adnatum of s. GA and FL west to LA); disc flowers (6-) 11-110 per head.
2 Stem leaves mostly ≥1.5 cm long, spreading to erect-ascending; [collectively widespread].
4 Midstem leaves <3cm long, bases rounded to cuneate (sometimes subclasping); leaves and bracts silvery-sericeous (glabrous in S. concolor var. devestitum); leaves held erect-ascending, lower often twisted by 90 degrees to bring edge in line with stem; plants not stipitate-glandular; [section Virgulus].
6Phyllaries with lower (outer) surfaces glabrous to glabrate (rarely sparsely to densely sericeous), the marginsciliate with thick-based, spreading hairs 0.5-1.0 mm long (much longer than those of the surfaces); largest phyllaries with terminal, green, pubescent portion 2/3-3/4 as long as the total phyllary length (longer than the proximal, indurated, glabrate portion); middle phyllaries mostly > 8 mm long and > 2.5 mm wide, averaging 9 (-14) mm long × 3 (-5) mm wide; involucre 8-15 mm tall; [mostly of the West Gulf Coastal Plain, disjunct e. to VA and s. GA]
6Phyllaries densely sericeous, the vestiture of the margins similar to that of the surfaces; largest phyllaries with terminal, green, pubescent portion 1/4-1/3 (-1/2) as long as the total phyllary length (shorter than the proximal, indurated, glabrate portion); middle phyllaries mostly < 7 mm long and < 2 mm wide, averaging 6 mm long × 2 mm wide; involucre 7-11 mm tall; [mostly of the Great Plains, s. to ne. TX, nw. AR, and s. IL]
15Involucres 8-10 (-12) mm high; disc flowers 8-10 mm long, white with purplish lobes; heads 4-5 (-6) cm across (ray tip to ray tip), the rays 14-24 mm long; plants strongly rhizomatous, forming clonal colonies with the stems mostly scattered along the rhizome (new stems typically arising at least several cm from the old ones); achenes 2.5-4.0 mm long, pale gray-brown, the trichomes about 0.4 mm long and distributed on and between the ribs; anthers purplish; pollen white
15Involucres 5.5-7.5 (-8.5) mm high (or to 12 mm high in S. patens var patentissimum, barely entering our area in w. KY and w. MS); disc flowers 5.5-8 mm long, either white with purplish lobes or bright yellow; heads 3-4 (-4.5) cm across (ray tip to ray tip), the rays 10-18 (-20) mm long; plants cespitose, generally with 1 or more stems arising from caudices (the new stems arising near the old); achenes 2.0-4.0 mm long, tan, gray, brown, dark-brown, or black, the trichomes various (see below); anthers purplish or yellow; pollen white or yellow.
16Disc flowers white with purplish lobes; stem leaves 7.5-12.5 (-14) cm long, thin in texture, soft-pubescent, the venation apparent, rugose-veiny and wrinkled; anthers purplish; pollen white; achenes 2.5-4.0 mm long, the trichomes concentrated on the ribs, < 0.4 mm long, appressed; [primarily of the Mountains, less commonly the Piedmont, mostly in moist, shady to semi-sunny situations]
16Disc flowers bright yellow; stem leaves (2-) 3-7 (-9) cm long, thick in texture, scabrous, the venation inconspicuous; anthers yellow; pollen yellow; achenes 2.0-3.5 mm long, the trichomes distributed on and between the ribs, mostly > 0.4 mm long, spreading; [collectively widespread in our area, mostly in dry, semi-sunny to sunny situations]
18 Mid-stem leaves mostly 5-7 cm long, separated by internodes of (1-) 1.5-3 (-4) cm at their densest, usually spreading; plants usually 0.8-1.6 m tall; heads mostly 9-12 mm wide
18 Mid-stem leaves mostly 3-5 cm long, separated by internodes of (0.5-) 1-2 (-3) cm at their densest, often adnate-ascending; plants usually 0.4-0.8 m tall; heads mostly 7-11 mm wide.
19 Stems and leaves generally eglandular (except sometimes for scattered glands on distal branches), but with dense ascendingeglandularhairs; plants usually dull greyish-green, not much darkening when dried; leaves usually with gradual or irregular reduction from base of stem to summit; mid-stem leaves with length/width (2-) 2.5-4 (-4.5), not forming a distinct overlapping cluster; bracts on proximal thirds of peduncles mostly 2-3 (-5) mm wide
19 Stems and leaves with dense to sparse stipitate-glands, with or without eglandularhairs; plants somewhat bluish-waxy, often becoming blackish when dried; leaves abruptly reduced in size with each branching order; mid-stem leaves [directly below inflorescence] with length/width (1.5-) 2-2.5(-3), clustered and overlapping along 10-20 cm; bracts on proximal thirds of peduncles mostly 1-2 (-3) mm wide
1 Mid-stem leaves narrowed above the claspingbase to an elongate petiolar portion (1/2-1/8 of the blade length) which flares abruptly into an ovate to lanceolate upper portion 10-55 mm wide; stem slightly to strongly zigzag in the upper portion; [series Punicei]
4 Lower stem leaves attenuate, sessile to winged-petiolate; leaves thick-textured, often slightly fleshy; stems erect; [marshes, pine savannahs; Atlantic Coastal Plain of GA northwards, disjunct in w. NC]
5 Leaves linear, > 10× as long as wide; leaf bases only subclasping
8 Upper and lower leaf surfaces distinctly different in color (the lower pale green, the upper dark green); upper stem leaves (on branches leading to heads) equal or nearly so in size to mid-stem leaves
1Ray flowerblades (3-) 3.8-7.5 (-11) mm long, white (rarely slightly blue or violet); involucres (2.5-) 3.1-5.1 (-6.5) mm high; [various habitats, including glades].
2Disc flowers (13-) 17-39 (-67); ray flowers (10-) 16-28 (-38); involucres (2.5-) 3.5-5.1 (-6.5) mm high.
4 Stems sparsely to densely hairy with short pubescence; plants 3-10 dm tall; upper stem leaves with axillaryfascicle shoots; [prairies, woodlands, barrens, and glades of n. IL and n. IA south to n. AR and ne. OK]
7Rays 9-17 (-19); leaves (3-) 4-9 cm long; array leaves and bracts thick, firm, linear-oblong; [MD, PA, OH, KY, s. IL, se. MO, AR, and se. OK and southwards]
8Inflorescencepaniculiform, ±crowded, branches mostly erect-ascending and restricted to upper ½ of plant (broadly pyramidal with spreading-arching branches in S. racemosum var. subdumosum); branch leaves and pedunclebracts decreasing gradually in size, often broader; [collectively widespread].
10Peduncles about equal to or shorter than subtending array leaves; upper stems glabrate or slightly scabrous; leaves firm, slightly fleshy; [outer Coastal Plain of NC and SC]
11 Upper stems, array branches, and both faces of leaves strigoso-scabrous or hirtellous (short, stiff, spreading hairs), usually densely so; plants usually stipitate-glandular distally (on upper stems, array branches, and leaves); leaves and bracts firm, stiffly spreading, sessile.
17 Leaves thin, margins planar or (S. racemosum, S. eulae) recurved; involucres 3-5.6 (-6) mm (-7.2 mm in S. hesperium); rays usually white, sometimes pinkish-tinged (commonly purplish-blue in S. hesperium); [collectively widespread].
19Disccorolla lobes < 1/2 the length of the corollalimb, erect to spreading; raylaminae (4.2-) 5-10 (-14)mm; rays 16-45 (occasionally as few as 12 in S. racemosum and S. tradescantii).
21 Plants short-rhizomatous and cespitose; leaf blades mostly < 1 cm wide, usually held erect-ascending; stems 1.5-6 dm tall, 1-3 mm wide at base; [rocky shores and river shoals, n. NJ northwards].
21 Plants long-rhizomatous and colonial; leaf blades to 3 cm wide, lax or spreading; stems (3-) 6-15 dm tall, 3-6+ mm wide at base; [collectively widespread].
23Disccorolla lobes mostly widely spreading, about 1/2 length of limbs, usually yellow (fading brownish); rayblades mostly 3.5-5.5 mm; leaves pubescent across face abaxially
28 Heads not congested or congested only distally on lateral branches of arrays; involucres 3.6-5.6 (-6) mm high, and mostly > 4 mm high; [collectively widespread].
1 Leaves strongly basally disposed, much reduced on the mid and upper stem and the basal and lower stem leaves sometimes withering by anthesis; leaf bladeslinear or linear-lanceolate (lower) to linear-subulate (upper stem), > 10× as long as wide; leaf margins strongly scabrous; [West Gulf Coastal Plain]